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<document id="CECAC969461AB60A16FF7A98C703820E" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFC3FFE2FCBEFF325A9D39AA" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) guanajuatoensis Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="20" updateTime="1700168056660" updateUser="ExternalLinkService">
<mods:mods id="D39FB49F2D345E9A10C7640A3CF2597A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0D4896374C55BCDC0A52D83D4585A775">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="634DE267730CEC9ADFEA581D225A00BB">Perkins, Philip D.</mods:namePart>
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<mods:title id="1F4EC50AABAC25B4FE9C89CC3E191067">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BE98477B50DAED8BD435C13856A1596E">
<mods:date id="FBDFF6AE8057565AB8869E806EEF5147">2023</mods:date>
<mods:detail id="B55EC0C741026CF073EFA161CC8FC690" type="pubDate">
<mods:number id="7D7EE2B3B95EB4FED57FAEA56C2CC714">2023-11-07</mods:number>
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<mods:number id="F82612435EFB788EC705B8BFF3E87786">5367</mods:number>
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<mods:number id="65EAFE92DE9222EC48C329F0CEACB027">1</mods:number>
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</mods:location>
<mods:classification id="13E733F19BF3C1CF90C752EDD2F97646">journal article</mods:classification>
<mods:identifier id="CC003C8E8F648711801264EE81A9773B" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
<mods:identifier id="8279376440079E939CD1A8EE47C808DA" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="677FC176756D80300E981277CA99ED1C" type="Zenodo-Dep">10145106</mods:identifier>
<mods:identifier id="31A08C25D491D58146BB340368C3BBF8" type="ZooBank">5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723</mods:identifier>
</mods:mods>
<treatment id="03FD878BFFC3FFE2FCBEFF325A9D39AA" LSID="urn:lsid:plazi:treatment:03FD878BFFC3FFE2FCBEFF325A9D39AA" httpUri="http://treatment.plazi.org/id/03FD878BFFC3FFE2FCBEFF325A9D39AA" lastPageNumber="20" pageId="19" pageNumber="20">
<subSubSection id="C34E6516FFC3FFE2FCBEFF325A513C86" box="[151,830,151,178]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BEB369DFFC3FFE2FCBEFF325A513C86" blockId="19.[151,830,151,213]" box="[151,830,151,178]" pageId="19" pageNumber="20">
<heading id="D0A381F1FFC3FFE2FCBEFF325A513C86" bold="true" box="[151,830,151,178]" fontSize="11" level="2" pageId="19" pageNumber="20" reason="4">
<emphasis id="B920EA8FFFC3FFE2FCBEFF325A513C86" bold="true" box="[151,830,151,178]" pageId="19" pageNumber="20">
<taxonomicName id="4C544D1EFFC3FFE2FCBEFF325BF03C86" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,671,151,178]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="guanajuatoensis" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B920EA8FFFC3FFE2FCBEFF325BF03C86" bold="true" box="[151,671,151,178]" italics="true" pageId="19" pageNumber="20">Hydraena (Hydraenopsis) guanajuatoensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357F4FFC3FFE2FE87FF325A513C86" box="[686,830,151,177]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCBEFF1E5A653CE2" box="[151,778,187,213]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BEB369DFFC3FFE2FCBEFF1E5A653CE2" blockId="19.[151,830,151,213]" box="[151,778,187,213]" pageId="19" pageNumber="20">
<figureCitation id="136F2A18FFC3FFE2FCBEFF1E598C3CE2" box="[151,227,187,213]" captionStart="FIGURE 3" captionStartId="7.[151,250,1639,1663]" captionTargetBox="[206,1381,181,1615]" captionTargetId="figure-18@7.[206,1381,181,1615]" captionTargetPageId="7" captionText="FIGURE 3. Dorsal habitus of Hydraena species, H. leechi Group." figureDoi="http://doi.org/10.5281/zenodo.10145112" httpUri="https://zenodo.org/record/10145112/files/figure.png" pageId="19" pageNumber="20">Figs. 3</figureCitation>
(habitus comparison), 10 (habitus and aedeagus)
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCBEFE825C3A3DBE" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BEB369DFFC3FFE2FCBEFE825C3A3DBE" blockId="19.[151,1437,295,1438]" pageId="19" pageNumber="20">
<emphasis id="B920EA8FFFC3FFE2FCBEFE82583F3D76" bold="true" box="[151,336,295,321]" pageId="19" pageNumber="20">Type Material.</emphasis>
<materialsCitation id="3B3C3CC0FFC3FFE2FD49FE825A213D52" collectingDate="1999-08-11" collectionCode="IEXA" collectorName="R. Arce" country="Mexico" elevation="1950" latitude="21.416388" location="Pto de Palmas" longLatPrecision="21" longitude="-100.083336" municipality="Victoria" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Guanajuato" typeStatus="holotype">
<emphasis id="B920EA8FFFC3FFE2FD49FE8258A23D76" bold="true" box="[352,461,295,321]" pageId="19" pageNumber="20">
<typeStatus id="54EF883FFFC3FFE2FD49FE8258A23D76" box="[352,461,295,321]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
(male): “
<collectingCountry id="F343760DFFC3FFE2FE6FFE825BD43D76" box="[582,699,295,321]" name="Mexico" pageId="19" pageNumber="20">MEXICO</collectingCountry>
:
<collectingRegion id="4990F87FFFC3FFE2FEE7FE825A3D3D76" box="[718,850,295,321]" country="Mexico" name="Guanajuato" pageId="19" pageNumber="20">Guanajuato</collectingRegion>
,
<collectingMunicipality id="6B8FACE7FFC3FFE2FF4CFE825AAD3D76" box="[869,962,295,321]" pageId="19" pageNumber="20">Victoria</collectingMunicipality>
,
<location id="8E8B6046FFC3FFE2FFFFFE8D5DE73D76" LSID="urn:lsid:plazi:treatment:03FD878BFFC3FFE2FCBEFF325A9D39AA:8E8B6046FFC3FFE2FFFFFE8D5DE73D76" box="[982,1160,295,322]" country="Mexico" latitude="21.416388" longLatPrecision="21" longitude="-100.083336" municipality="Victoria" name="Pto de Palmas" pageId="19" pageNumber="20" stateProvince="Guanajuato">Pto de Palmas</location>
,
<quantity id="4CAC9B78FFC3FFE2F8B2FE825D943D75" box="[1179,1275,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="19" pageNumber="20" unit="m" value="1950.0">
<elevation id="0079D1AEFFC3FFE2F8B2FE825D943D75" box="[1179,1275,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="19" pageNumber="20" unit="m" value="1950.0">1950 m</elevation>
</quantity>
,
<date id="FFEA105DFFC3FFE2F926FE825CF83D76" box="[1295,1431,295,321]" pageId="19" pageNumber="20" value="1999-08-11">
<collectingDate id="EFAEE9B5FFC3FFE2F926FE825CF83D76" box="[1295,1431,295,321]" pageId="19" pageNumber="20" value="1999-08-11">11.viii.1999</collectingDate>
</date>
,
<geoCoordinate id="EE60505AFFC3FFE2FCBEFEEE58483D51" box="[151,295,331,358]" degrees="21" direction="north" minutes="24" orientation="latitude" pageId="19" pageNumber="20" precision="15" seconds="59" value="21.416388">21°24’59’’N</geoCoordinate>
,
<geoCoordinate id="EE60505AFFC3FFE2FD1AFEEE58BD3D51" box="[307,466,331,358]" degrees="100" direction="west" minutes="04" orientation="longitude" pageId="19" pageNumber="20" precision="15" seconds="60" value="-100.083336">100°04’60’’W</geoCoordinate>
,
<quantity id="4CAC9B78FFC3FFE2FDC9FEEE5B573D51" box="[480,568,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.462" pageId="19" pageNumber="20" unit="m" value="1462.0">
<elevation id="0079D1AEFFC3FFE2FDC9FEEE5B573D51" box="[480,568,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.462" pageId="19" pageNumber="20" unit="m" value="1462.0">1462 m</elevation>
</quantity>
,
<collectorName id="26A1534BFFC3FFE2FE6DFEE95BF53D52" box="[580,666,331,358]" pageId="19" pageNumber="20">R. Arce</collectorName>
col. 1” (
<collectionCode id="ED45AE58FFC3FFE2FED5FEEE5A2A3D52" box="[764,837,331,357]" pageId="19" pageNumber="20">IEXA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CC0FFC3FFE2FF71FEEE5CF73D52" box="[856,1432,331,358]" collectingDate="1999-08-11" collectionCode="IEXA" collectorName="R. Arce" country="Mexico" elevation="1950" latitude="21.416388" location="Pto de Palmas" longLatPrecision="21" longitude="-100.083336" municipality="Victoria" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Guanajuato" typeStatus="paratype">
<emphasis id="B920EA8FFFC3FFE2FF71FEEE5D7F3D52" bold="true" box="[856,1040,331,357]" pageId="19" pageNumber="20">
<typeStatus id="54EF883FFFC3FFE2FF71FEEE5ABF3D52" box="[856,976,331,357]" pageId="19" pageNumber="20" type="paratype">Paratypes</typeStatus>
(22):
</emphasis>
Same data as holotype (15
<collectionCode id="ED45AE58FFC3FFE2F961FEEE5CFF3D52" box="[1352,1424,331,357]" pageId="19" pageNumber="20">IEXA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CC0FFC3FFE2FCBEFECA5C3E3DBE" box="[151,1361,367,394]" collectingDate="1999-08-12" collectionCode="IEXA" collectorName="R. Arce Col." country="Mexico" elevation="2400" latitude="21.390833" location="Joya Fria" municipality="Victoria" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Guanajuato" typeStatus="paratype">
<collectingRegion id="4990F87FFFC3FFE2FCBEFECA58743DBE" box="[151,283,367,393]" country="Mexico" name="Guanajuato" pageId="19" pageNumber="20">Guanajuato</collectingRegion>
,
<collectingMunicipality id="6B8FACE7FFC3FFE2FD0FFECA58EC3DBE" box="[294,387,367,393]" pageId="19" pageNumber="20">Victoria</collectingMunicipality>
,
<location id="8E8B6046FFC3FFE2FDA7FED558973DBE" LSID="urn:lsid:plazi:treatment:03FD878BFFC3FFE2FCBEFF325A9D39AA:8E8B6046FFC3FFE2FDA7FED558973DBE" box="[398,504,367,394]" country="Mexico" latitude="21.390833" municipality="Victoria" name="Joya Fria" pageId="19" pageNumber="20" stateProvince="Guanajuato">Joya Fría</location>
,
<date id="FFEA105DFFC3FFE2FE2AFECA5BE13DBE" box="[515,654,367,393]" pageId="19" pageNumber="20" value="1999-08-12">
<collectingDate id="EFAEE9B5FFC3FFE2FE2AFECA5BE13DBE" box="[515,654,367,393]" pageId="19" pageNumber="20" value="1999-08-12">12.viii.1999</collectingDate>
</date>
.
<emphasis id="B920EA8FFFC3FFE2FEB0FECA5D263DBE" box="[665,1097,367,394]" italics="true" pageId="19" pageNumber="20">
<geoCoordinate id="EE60505AFFC3FFE2FEB0FECA5A423DBE" box="[665,813,367,393]" degrees="21.390833" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="1" value="21.390833">21.390833 N</geoCoordinate>
, -100.144444 O,
<quantity id="4CAC9B78FFC3FFE2FFC6FECA5D2A3DBE" box="[1007,1093,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="20" unit="m" value="2400.0">
<elevation id="0079D1AEFFC3FFE2FFC6FECA5D2A3DBE" box="[1007,1093,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="19" pageNumber="20" unit="m" value="2400.0">2400 m</elevation>
</quantity>
,
</emphasis>
<collectorName id="26A1534BFFC3FFE2F879FED55DB33DBE" box="[1104,1244,367,394]" pageId="19" pageNumber="20">R. Arce Col.</collectorName>
(7
<collectionCode id="ED45AE58FFC3FFE2F928FECA5C263DBE" box="[1281,1353,367,393]" pageId="19" pageNumber="20">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCEEFE365B483EB2" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BEB369DFFC3FFE2FCEEFE365B483EB2" blockId="19.[151,1437,295,1438]" pageId="19" pageNumber="20">
<emphasis id="B920EA8FFFC3FFE2FCEEFE3658BB3D9A" bold="true" box="[199,468,403,429]" pageId="19" pageNumber="20">Differential Diagnosis.</emphasis>
This new species is a member of the
<taxonomicName id="4C544D1EFFC3FFE2FFACFE315D0E3D9A" authority="Subgroup" authorityName="Subgroup" box="[901,1121,403,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC3FFE2FFACFE315A833D9A" box="[901,1004,403,429]" italics="true" pageId="19" pageNumber="20">H. leechi</emphasis>
Subgroup
</taxonomicName>
of the
<taxonomicName id="4C544D1EFFC3FFE2F89DFE315C743D9A" box="[1204,1307,403,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="leechi">
<emphasis id="B920EA8FFFC3FFE2F89DFE315C743D9A" box="[1204,1307,403,429]" italics="true" pageId="19" pageNumber="20">H. leechi</emphasis>
</taxonomicName>
Group, the group of species that have a pronotal scintilla, and the males having metatibial brushes (
<bibRefCitation id="EFC54B6CFFC3FFE2F876FE125C4F3DE6" author="Perkins, P. D." box="[1119,1312,439,465]" pageId="19" pageNumber="20" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 36</bibRefCitation>
). Members will key to couplet
<quantity id="4CAC9B78FFC3FFE2FD45FE7E58F93DC2" box="[364,406,475,502]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="19" pageNumber="20" unit="in" value="5.0">5 in</quantity>
<bibRefCitation id="EFC54B6CFFC3FFE2FDB5FE7E5B003DC1" author="Perkins, P. D." box="[412,623,475,502]" pageId="19" pageNumber="20" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 65)</bibRefCitation>
. The species can be recognized by the relatively large size (ca.
<quantity id="4CAC9B78FFC3FFE2F900FE7E5CFF3DC2" box="[1321,1424,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.02" pageId="19" pageNumber="20" unit="mm" value="2.02">2.02 mm</quantity>
), the metaventral plaques very narrow, straight, slightly converging toward one another anteriorly, not raised, the metatibia of males arcuate, gradually increasing in width from base to apex, inner margin with moderately large brush of arcuate setae, and the aedeagus (
<figureCitation id="136F2A18FFC3FFE2FE50FDE25BA03E56" box="[633,719,583,609]" captionStart="FIGURE 10" captionStartId="18.[151,250,1792,1816]" captionTargetBox="[312,1275,181,1768]" captionTargetId="figure-35@18.[312,1275,181,1768]" captionTargetPageId="18" captionText="FIGURE 10. Hydraena (Hydraenopsis) guanajuatoensis new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145126" httpUri="https://zenodo.org/record/10145126/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
). Reliable determinations will include dissection of males and careful examination of the aedegus.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCEEFD2A58BE3962" pageId="19" pageNumber="20" type="description">
<paragraph id="8BEB369DFFC3FFE2FCEEFD2A5B463F92" blockId="19.[151,1437,295,1438]" pageId="19" pageNumber="20">
<emphasis id="B920EA8FFFC3FFE2FCEEFD2A583E3E9E" bold="true" box="[199,337,655,681]" pageId="19" pageNumber="20">Description</emphasis>
. Size:
<typeStatus id="54EF883FFFC3FFE2FD8DFD2A5B693E9E" box="[420,518,655,681]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 2.02/0.83; head width 0.44; pronotum 0.53/0.66, PA 0.50, PB 0.56; elytra 1.25/0.83. Habitus as illustrated (
<figureCitation id="136F2A18FFC3FFE2F800FD165D103EFA" box="[1065,1151,691,717]" captionStart="FIGURE 10" captionStartId="18.[151,250,1792,1816]" captionTargetBox="[312,1275,181,1768]" captionTargetId="figure-35@18.[312,1275,181,1768]" captionTargetPageId="18" captionText="FIGURE 10. Hydraena (Hydraenopsis) guanajuatoensis new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145126" httpUri="https://zenodo.org/record/10145126/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
). Dorsum brown to dark brown, palpi brown, tip of last palpomere darker. Dorsum of head and pronotum coarsely densely punctate, interstices shining; punctures of frons disc ca. 1–2xef, interstices ca. 1–2xpd; punctures of pronotal disc ca. 2–3xef, interstices narrow walls to ca. 2xpd; punctures of clypeus moderately fine, lateral areas microreticulate. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle ½, scintilla large; anterolateral, posterolateral and posteromedial pronotal foveae deep; anterior ½ of disc with shallow transverse impression.
</paragraph>
<paragraph id="8BEB369DFFC3FFE2FCEEFC0A5D1B3802" blockId="19.[151,1437,295,1438]" pageId="19" pageNumber="20">Elytra with summit of posterior declivity at or near posterior 1/3; lateral explanate margins wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, width ca. 1xpd or slightly less, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BEB369DFFC3FFE2FCEEFBE558BE3962" blockId="19.[151,1437,295,1438]" pageId="19" pageNumber="20">Venter: Mentum very densely very finely punctulate, interstices weakly shining; postmentum very finely densely micropunctulate. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/0.25/1/3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 3/1, parallel, apex blunt, apex raised slightly above mesoventral intercoxal process. Plaques very narrow, straight, slightly converging toward one another anteriorly, not raised, located at sides of deep median depression. AIS width at straight posterior margin ca. 2x P2. Pro- and mesotibiae moderately slender, straight. Metatibia arcuate, gradually increasing in width from base to apex, inner margin with moderately large brush of arcuate setae. Abdominal apex symmetrical; last tergite with shallow apicomedian notch.</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCEEFAFA5D05394E" box="[199,1130,1375,1402]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BEB369DFFC3FFE2FCEEFAFA5D05394E" blockId="19.[151,1437,295,1438]" box="[199,1130,1375,1402]" pageId="19" pageNumber="20">
<emphasis id="B920EA8FFFC3FFE2FCEEFAFA580E394E" bold="true" box="[199,353,1375,1401]" pageId="19" pageNumber="20">Distribution.</emphasis>
Currently known only from the
<typeStatus id="54EF883FFFC3FFE2FEE5FAC55B93394D" box="[716,764,1376,1402]" pageId="19" pageNumber="20">type</typeStatus>
locality and one nearby locality.
</paragraph>
</subSubSection>
<subSubSection id="C34E6516FFC3FFE2FCEEFA265A9D39AA" box="[199,1010,1411,1438]" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BEB369DFFC3FFE2FCEEFA265A9D39AA" blockId="19.[151,1437,295,1438]" box="[199,1010,1411,1438]" pageId="19" pageNumber="20">
<emphasis id="B920EA8FFFC3FFE2FCEEFA26582139AA" bold="true" box="[199,334,1411,1437]" pageId="19" pageNumber="20">Etymology.</emphasis>
Named in reference to the known geographical distribution.
</paragraph>
</subSubSection>
</treatment>
</document>
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