436 lines
47 KiB
XML
436 lines
47 KiB
XML
<document id="83451071DE7985E554DE492F57A5561B" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFF99FFBBFCBEFD2B5B653DE6" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ochthebius (Gymnochthebius) falli" docType="treatment" docVersion="2" lastPageNumber="75" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="74" updateTime="1700168056660" updateUser="ExternalLinkService">
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<mods:title id="B85BA00BC29944FF71A9C0BC3F7432A4">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="7860F84975759562FA2D371F2E00B514">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="408DF824AA49A0CB48A5E35B9C778AD9">text</mods:typeOfResource>
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<mods:title id="BA5034D72B5E330A1B5B7099A0FE5571">Zootaxa</mods:title>
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<mods:part id="782905EB7D6FC316927502B059A9DDD4">
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<mods:date id="E74D666840E839092FD3224225720C39">2023</mods:date>
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<mods:number id="72E3BDBB931825D50525C277EAAFA4A0">2023-11-07</mods:number>
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<mods:number id="F99115F9D0619B3B32548AB3E6819F6A">5367</mods:number>
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<mods:url id="DBF21C5DFDF38A17C6386C41B055B11F">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
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<mods:classification id="E4976369B9CF0B60B08E93CA632F7F62">journal article</mods:classification>
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<mods:identifier id="B8544240B85038ACC09B91309FC4C82D" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
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<mods:identifier id="8737566A1F6B99908BC187F8B6231B07" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="BA2D758722A1A95131BE472BC94E5CEB" type="Zenodo-Dep">10145106</mods:identifier>
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<treatment id="03FD878BFF99FFBBFCBEFD2B5B653DE6" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6" httpUri="http://treatment.plazi.org/id/03FD878BFF99FFBBFCBEFD2B5B653DE6" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
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<subSubSection id="C34E6516FF99FFB8FCBEFD2B5D6C3E9F" box="[151,1027,654,681]" pageId="73" pageNumber="74" type="nomenclature">
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<paragraph id="8BEB369DFF99FFB8FCBEFD2B5D6C3E9F" blockId="73.[151,1027,654,717]" box="[151,1027,654,681]" pageId="73" pageNumber="74">
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<heading id="D0A381F1FF99FFB8FCBEFD2B5D6C3E9F" bold="true" box="[151,1027,654,681]" fontSize="11" level="2" pageId="73" pageNumber="74" reason="4">
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<emphasis id="B920EA8FFF99FFB8FCBEFD2B5D6C3E9F" bold="true" box="[151,1027,654,681]" pageId="73" pageNumber="74">
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<taxonomicName id="4C544D1EFF99FFB8FCBEFD2B5BD23E9F" ID-CoL="3HR59" authority="(Perkins)" baseAuthorityName="Perkins" box="[151,701,654,681]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
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<emphasis id="B920EA8FFF99FFB8FCBEFD2B584D3E9E" bold="true" box="[151,290,654,681]" italics="true" pageId="73" pageNumber="74">Ochthebius</emphasis>
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(
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<emphasis id="B920EA8FFF99FFB8FD1AFD2B5B6F3E9E" bold="true" box="[307,512,654,681]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
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)
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<emphasis id="B920EA8FFF99FFB8FE38FD2B5B2D3E9E" bold="true" box="[529,578,654,681]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
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(Perkins)
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</taxonomicName>
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Species Status Reinstated
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FF99FFB8FCBEFD1658413EFA" box="[151,302,691,717]" pageId="73" pageNumber="74" type="description">
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<paragraph id="8BEB369DFF99FFB8FCBEFD1658413EFA" blockId="73.[151,1027,654,717]" box="[151,302,691,717]" pageId="73" pageNumber="74">
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(
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<figureCitation id="136F2A18FF99FFB8FCB6FD1658493EFA" box="[159,294,691,717]" captionStart-0="FIGURES34–35" captionStart-1="FIGURE 36" captionStartId-0="58.[151,264,1792,1816]" captionStartId-1="59.[151,250,1629,1653]" captionTargetBox-0="[186,1395,184,1753]" captionTargetBox-1="[151,1436,181,1605]" captionTargetId-0="figure-35@58.[166,1421,181,1768]" captionTargetId-1="figure-18@59.[151,1436,181,1605]" captionTargetPageId-0="58" captionTargetPageId-1="59" captionText-0="FIGURES34–35.Habitusandmalegenitalia.34.Ochthebius(Gymnochthebius)falli(Perkins).35.Ochthebius(Gymnochthebius) fossatus LeConte." captionText-1="FIGURE 36. Geographical distribution of Ochthebius (Gymnochthebius) falli (Perkins)." figureDoi-0="http://doi.org/10.5281/zenodo.10145174" figureDoi-1="http://doi.org/10.5281/zenodo.10145176" httpUri-0="https://zenodo.org/record/10145174/files/figure.png" httpUri-1="https://zenodo.org/record/10145176/files/figure.png" pageId="73" pageNumber="74">Figs. 34–36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C34E6516FF99FFB8FCBEFD525AAC3F19" pageId="73" pageNumber="74" type="reference_group">
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<paragraph id="8BEB369DFF99FFB8FCBEFD525B403F38" blockId="73.[151,963,759,814]" box="[151,559,759,783]" pageId="73" pageNumber="74">
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<treatmentCitationGroup id="AB4411B3FF99FFB8FCBEFD525B403F38" box="[151,559,759,783]" pageId="73" pageNumber="74">
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<treatmentCitation id="0AF5108CFF99FFB8FCBEFD525B403F38" author="Perkins, P. D." box="[151,559,759,783]" page="281" pageId="73" pageNumber="74" year="1980">
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<taxonomicName id="4C544D1EFF99FFB8FCBEFD525B403F38" ID-CoL="3HR59" authority="Perkins 1980: 281" authorityName="Perkins" authorityPageNumber="281" authorityYear="1980" box="[151,559,759,783]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli">
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<emphasis id="B920EA8FFF99FFB8FCBEFD52581E3F38" box="[151,369,759,783]" italics="true" pageId="73" pageNumber="74">Gymnochthebius falli</emphasis>
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<bibRefCitation id="EFC54B6CFF99FFB8FD5EFD525B403F38" author="Perkins, P. D." box="[375,559,759,783]" pageId="73" pageNumber="74" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 281</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BEB369DFF99FFB8FCBEFCB35AAC3F19" blockId="73.[151,963,759,814]" box="[151,963,790,814]" pageId="73" pageNumber="74">
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<taxonomicName id="4C544D1EFF99FFB8FCBEFCB35BEA3F19" ID-CoL="6SFSW" authority="LeConte" authorityName="LeConte" box="[151,645,790,814]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
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<emphasis id="B920EA8FFF99FFB8FCBEFCB358653F19" box="[151,266,790,814]" italics="true" pageId="73" pageNumber="74">Ochthebius</emphasis>
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(
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<emphasis id="B920EA8FFF99FFB8FD30FCB358AC3F19" box="[281,451,790,814]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
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)
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<emphasis id="B920EA8FFF99FFB8FDFBFCB35B4B3F19" box="[466,548,790,814]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
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LeConte
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</taxonomicName>
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;
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<bibRefCitation id="EFC54B6CFF99FFB8FEA6FCB35B9A3F19" author="Jach, M. A." box="[655,757,790,814]" pageId="73" pageNumber="74" pagination="301 - 308" refId="ref49627" refString="Jach, M. A. (1994) Descriptions of new species of Hydraenidae (Coleoptera) from Texas, Indiana, and Oklahoma, with faunistic and taxonomic notes on the family in Texas. The Coleopterists Bulletin, 48 (4), 301 - 308." type="journal article" year="1994">Jäch 1994</bibRefCitation>
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(invalid synonymy)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E6516FF99FFB8FCBEFCF95D4C3935" pageId="73" pageNumber="74" type="description">
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||
<paragraph id="8BEB369DFF99FFB8FCBEFCF958353879" blockId="73.[151,1437,860,2039]" pageId="73" pageNumber="74">
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||
<materialsCitation id="3B3C3CC0FF99FFB8FCBEFCF958383879" country="Mexico" county="The" location="The" municipality="The" pageId="73" pageNumber="74" specimenCount="3" specimenCount-male="1" stateProvince="Oaxaca">
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||
<collectingCounty id="628A4E11FF99FFB8FCBEFCF959AC3F41" box="[151,195,860,886]" pageId="73" pageNumber="74">The</collectingCounty>
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following collection/deposition data include the first records for
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<collectingCountry id="F343760DFF99FFB8FF82FCF95D6B3F41" box="[939,1028,860,886]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
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||
.
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||
<collectingMunicipality id="6B8FACE7FF99FFB8F839FCF95D533F41" box="[1040,1084,860,886]" pageId="73" pageNumber="74">The</collectingMunicipality>
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data also includes several first museum depositories for the species.
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<location id="8E8B6046FF99FFB8FE63FC255B193FAD" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FE63FC255B193FAD" box="[586,630,896,922]" country="Mexico" county="The" municipality="The" name="The" pageId="73" pageNumber="74" stateProvince="Oaxaca">The</location>
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new records increase the number of identified specimens in museum collections from 40 to 60.
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<location id="8E8B6046FF99FFB8FDEAFC0158803F89" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FDEAFC0158803F89" box="[451,495,932,958]" country="Mexico" county="The" municipality="The" name="The" pageId="73" pageNumber="74" stateProvince="Oaxaca">The</location>
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range of this species (
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<figureCitation id="136F2A18FF99FFB8FEC4FC015A2F3F89" box="[749,832,932,958]" captionStart="FIGURE 36" captionStartId="59.[151,250,1629,1653]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-18@59.[151,1436,181,1605]" captionTargetPageId="59" captionText="FIGURE 36. Geographical distribution of Ochthebius (Gymnochthebius) falli (Perkins)." figureDoi="http://doi.org/10.5281/zenodo.10145176" httpUri="https://zenodo.org/record/10145176/files/figure.png" pageId="73" pageNumber="74">Fig. 36</figureCitation>
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) extends northward from
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<collectingRegion id="4990F87FFF99FFB8F84EFC015DD13F89" box="[1127,1214,932,958]" country="Mexico" name="Oaxaca" pageId="73" pageNumber="74">Oaxaca</collectingRegion>
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,
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<collectingCountry id="F343760DFF99FFB8F8E3FC015C4F3F89" box="[1226,1312,932,958]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
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to
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<location id="8E8B6046FF99FFB8F96CFC0059873FD4" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8F96CFC0059873FD4" country="Mexico" county="The" municipality="The" name="western Kansas" pageId="73" pageNumber="74" stateProvince="Oaxaca">western Kansas</location>
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and
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<location id="8E8B6046FF99FFB8FD36FC6D58903FD5" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FD36FC6D58903FD5" box="[287,511,968,994]" country="Mexico" county="The" municipality="The" name="southern Idaho. The" pageId="73" pageNumber="74" stateProvince="Oaxaca">southern Idaho. The</location>
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<specimenCount id="9D52FD14FF99FFB8FE2FFC6C5B043FD5" box="[518,619,968,995]" count="1" pageId="73" pageNumber="74" type="male">one male</specimenCount>
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specimen known from the northernmost locality, Idaho, was collected in a thermal spring in 1957. The
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<specimenCount id="9D52FD14FF99FFB8FDC1FC485BE23831" box="[488,653,1004,1031]" count="2" pageId="73" pageNumber="74" type="generic">two specimens</specimenCount>
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from one site in Arizona were collected in 1920. It is possible that the species has been extirpated from these localities. The records from
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<location id="8E8B6046FF99FFB8FFA8FBB45D70381C" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FFA8FBB45D70381C" box="[897,1055,1041,1067]" country="Mexico" county="The" municipality="The" name="western Texas" pageId="73" pageNumber="74" stateProvince="Oaxaca">western Texas</location>
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||
were collected relatively recently, from 1991–2000
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||
</materialsCitation>
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||
.
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</paragraph>
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||
<paragraph id="8BEB369DFF99FFB8FCEEFBFC5D4C3935" blockId="73.[151,1437,860,2039]" pageId="73" pageNumber="74">
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||
<materialsCitation id="3B3C3CC0FF99FFB8FCEEFBFC5C053845" box="[199,1386,1111,1139]" collectingDate="1963-08-03" collectingDateMax="1963-08-05" collectingDateMin="1963-08-03" collectionCode="NMNH" collectorName="P. J. Spangler" country="Mexico" location="New" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Mexico">
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||
<location id="8E8B6046FF99FFB8FCEEFBFC59933844" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FCEEFBFC59933844" box="[199,252,1113,1139]" country="Mexico" name="New" pageId="73" pageNumber="74" stateProvince="Mexico">New</location>
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||
records:
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||
<emphasis id="B920EA8FFF99FFB8FD4FFBFD58D23845" bold="true" box="[358,445,1112,1138]" pageId="73" pageNumber="74">
|
||
<collectingCountry id="F343760DFF99FFB8FD4FFBFD58D23845" box="[358,445,1112,1138]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
|
||
</emphasis>
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||
:
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||
<emphasis id="B920EA8FFF99FFB8FDE4FBFD5BC73845" bold="true" box="[461,680,1111,1138]" pageId="73" pageNumber="74">
|
||
<collectingRegion id="4990F87FFF99FFB8FDE4FBFD5BCB3845" box="[461,676,1111,1138]" country="Mexico" name="Mexico" pageId="73" pageNumber="74">Estado de México</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<collectingRegion id="4990F87FFF99FFB8FE86FBFD5A343845" box="[687,859,1112,1138]" country="Mexico" name="Aguascalientes" pageId="73" pageNumber="74">Aguascalientes</collectingRegion>
|
||
,
|
||
<date id="FFEA105DFF99FFB8FF4EFBFD5A903844" box="[871,1023,1112,1139]" pageId="73" pageNumber="74" value="1963-08-03" valueMax="1963-08-05" valueMin="1963-08-03">
|
||
<collectingDate id="EFAEE9B5FF99FFB8FF4EFBFD5A903844" box="[871,1023,1112,1139]" pageId="73" pageNumber="74" value="1963-08-03" valueMax="1963-08-05" valueMin="1963-08-03">3–5.viii.1963</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A1534BFF99FFB8F815FBFC5DBA3845" box="[1084,1237,1112,1139]" pageId="73" pageNumber="74">P. J. Spangler</collectorName>
|
||
(10
|
||
<collectionCode id="ED45AE58FF99FFB8F923FBFC5C0F3844" box="[1290,1376,1113,1139]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="73" pageNumber="74" type="Museum">NMNH</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3C3CC0FF99FFB8F951FBFD5A6438A1" collectingDate="1989-03-02" collectionCode="IEXA" collectorName="R. Arce" country="Mexico" location="Almoloya-Apan" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Mexico">
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||
km 15
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||
<location id="8E8B6046FF99FFB8FC95FBD9581A38A1" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FC95FBD9581A38A1" box="[188,373,1148,1174]" country="Mexico" name="Almoloya-Apan" pageId="73" pageNumber="74" stateProvince="Mexico">Almoloya-Apán</location>
|
||
,
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||
<date id="FFEA105DFF99FFB8FDADFBD9589D38A1" box="[388,498,1148,1174]" pageId="73" pageNumber="74" value="1989-03-02">
|
||
<collectingDate id="EFAEE9B5FF99FFB8FDADFBD9589D38A1" box="[388,498,1148,1174]" pageId="73" pageNumber="74" value="1989-03-02">2.iii.1989</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A1534BFF99FFB8FE28FBD85B3538A1" box="[513,602,1148,1175]" pageId="73" pageNumber="74">R. Arce</collectorName>
|
||
col. (3
|
||
<collectionCode id="ED45AE58FF99FFB8FE92FBD95A6C38A1" box="[699,771,1148,1174]" pageId="73" pageNumber="74">IEXA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3C3CC0FF99FFB8FF27FBD958FF388D" collectingDate="1990-10-09" collectionCode="LUM" collectorName="R. Baranowski" country="Mexico" location="3 km E Santiago Yolomecatl" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Oaxaca">
|
||
<emphasis id="B920EA8FFF99FFB8FF27FBD95A0038A1" bold="true" box="[782,879,1148,1174]" pageId="73" pageNumber="74">
|
||
<collectingRegion id="4990F87FFF99FFB8FF27FBD95A0538A1" box="[782,874,1148,1174]" country="Mexico" name="Oaxaca" pageId="73" pageNumber="74">Oaxaca</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<location id="8E8B6046FF99FFB8FF53FBD95DA338A1" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FF53FBD95DA338A1" box="[890,1228,1148,1175]" country="Mexico" name="3 km E Santiago Yolomecatl" pageId="73" pageNumber="74" stateProvince="Oaxaca">
|
||
<quantity id="4CAC9B78FF99FFB8FF53FBD95AD838A1" box="[890,951,1148,1174]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="73" pageNumber="74" unit="km" value="3.0">3 km</quantity>
|
||
E Santiago Yolomecatl
|
||
</location>
|
||
,
|
||
<date id="FFEA105DFF99FFB8F8F2FBD95C5038A1" box="[1243,1343,1148,1174]" pageId="73" pageNumber="74" value="1990-10-09">
|
||
<collectingDate id="EFAEE9B5FF99FFB8F8F2FBD95C5038A1" box="[1243,1343,1148,1174]" pageId="73" pageNumber="74" value="1990-10-09">9.x.1990</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A1534BFF99FFB8F9AAFBD85872388D" pageId="73" pageNumber="74">R. Baranowski</collectorName>
|
||
(1
|
||
<collectionCode id="ED45AE58FF99FFB8FD6CFB0558E7388D" box="[325,392,1184,1210]" pageId="73" pageNumber="74">LUM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3C3CC0FF99FFB8FDB4FB0558CB38E9" collectingDate="1991-07-17" collectionCode="TAMU" collectorName="R. C. Vogtsberger" country="United States of America" county="Brewster" location="Buttrill Springs" municipality="Big Bend National Park" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Texas">
|
||
<emphasis id="B920EA8FFF99FFB8FDB4FB055884388D" bold="true" box="[413,491,1184,1210]" pageId="73" pageNumber="74">
|
||
<collectingCountry id="F343760DFF99FFB8FDB4FB055884388D" box="[413,491,1184,1210]" name="United States of America" pageId="73" pageNumber="74">U.S.A.</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="B920EA8FFF99FFB8FDD3FB055B2A388D" bold="true" box="[506,581,1184,1210]" pageId="73" pageNumber="74">
|
||
<collectingRegion id="4990F87FFF99FFB8FDD3FB055B50388D" box="[506,575,1184,1210]" country="United States of America" name="Texas" pageId="73" pageNumber="74">Texas</collectingRegion>
|
||
,
|
||
</emphasis>
|
||
<collectingCounty id="628A4E11FF99FFB8FE64FB045BDB388C" box="[589,692,1185,1211]" pageId="73" pageNumber="74">Brewster</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6B8FACE7FF99FFB8FEE8FB055ABB388D" box="[705,980,1184,1210]" pageId="73" pageNumber="74">Big Bend National Park</collectingMunicipality>
|
||
,
|
||
<location id="8E8B6046FF99FFB8FFC8FB055DFC388D" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FFC8FB055DFC388D" box="[993,1171,1184,1210]" country="United States of America" county="Brewster" municipality="Big Bend National Park" name="Buttrill Springs" pageId="73" pageNumber="74" stateProvince="Texas">Buttrill Springs</location>
|
||
,
|
||
<date id="FFEA105DFF99FFB8F889FB055C4E388D" box="[1184,1313,1184,1210]" pageId="73" pageNumber="74" value="1991-07-17">
|
||
<collectingDate id="EFAEE9B5FF99FFB8F889FB055C4E388D" box="[1184,1313,1184,1210]" pageId="73" pageNumber="74" value="1991-07-17">17.vii.1991</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A1534BFF99FFB8F949FB04587238E9" pageId="73" pageNumber="74">R. C. Vogtsberger</collectorName>
|
||
(2
|
||
<collectionCode id="ED45AE58FF99FFB8FD6EFB6158F238E9" box="[327,413,1220,1246]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;M University" pageId="73" pageNumber="74" type="Herbarium">TAMU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3C3CC0FF99FFB8FD98FB615D9338E9" box="[433,1276,1220,1247]" collectingDate="1994-06-04" collectingDateMax="1994-06-05" collectingDateMin="1994-06-04" collectionCode="TAMU" collectorName="E. G. Riley" country="United States of America" location="Park" municipality="Stillwell" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Texas">
|
||
<collectingMunicipality id="6B8FACE7FF99FFB8FD98FB615B7F38E9" box="[433,528,1220,1246]" pageId="73" pageNumber="74">Stillwell</collectingMunicipality>
|
||
RV
|
||
<location id="8E8B6046FF99FFB8FE6EFB615B1538E9" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FE6EFB615B1538E9" box="[583,634,1220,1246]" country="United States of America" municipality="Stillwell" name="Park" pageId="73" pageNumber="74" stateProvince="Texas">Park</location>
|
||
on hwy 2627,
|
||
<date id="FFEA105DFF99FFB8FF03FB615ADD38E8" box="[810,946,1220,1247]" pageId="73" pageNumber="74" value="1994-06-04" valueMax="1994-06-05" valueMin="1994-06-04">
|
||
<collectingDate id="EFAEE9B5FF99FFB8FF03FB615ADD38E8" box="[810,946,1220,1247]" pageId="73" pageNumber="74" value="1994-06-04" valueMax="1994-06-05" valueMin="1994-06-04">4–5.vi.1994</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A1534BFF99FFB8FFDBFB605D1A38E9" box="[1010,1141,1220,1247]" pageId="73" pageNumber="74">E. G. Riley</collectorName>
|
||
(3
|
||
<collectionCode id="ED45AE58FF99FFB8F8B7FB615D9B38E9" box="[1182,1268,1220,1246]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;M University" pageId="73" pageNumber="74" type="Herbarium">TAMU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3C3CC0FF99FFB8F920FB615D703935" collectingDate="2000-05-04" collectionCode="MCZ" collectorName="N. Ruidosa Rd & G. L. Challet" country="United States of America" county="Presidio Co." location="Drying Creek" pageId="73" pageNumber="74" specimenCount="1" stateProvince="Texas">
|
||
<collectingCounty id="628A4E11FF99FFB8F920FB615CF738E9" box="[1289,1432,1220,1246]" pageId="73" pageNumber="74">Presidio Co.</collectingCounty>
|
||
,
|
||
<location id="8E8B6046FF99FFB8FCBEFB4D585C3935" LSID="urn:lsid:plazi:treatment:03FD878BFF99FFBBFCBEFD2B5B653DE6:8E8B6046FF99FFB8FCBEFB4D585C3935" box="[151,307,1256,1282]" country="United States of America" county="Presidio Co." name="Drying Creek" pageId="73" pageNumber="74" stateProvince="Texas">Drying Creek</location>
|
||
,
|
||
<quantity id="4CAC9B78FF99FFB8FD17FB4D58E93935" box="[318,390,1256,1282]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="73" pageNumber="74" unit="mi" value="12.0">12 mi.</quantity>
|
||
<collectorName id="26A1534BFF99FFB8FDA4FB4C5B5C3935" box="[397,563,1256,1283]" pageId="73" pageNumber="74">N. Ruidosa Rd</collectorName>
|
||
2810,
|
||
<date id="FFEA105DFF99FFB8FEA9FB4D5B8D3935" box="[640,738,1256,1282]" pageId="73" pageNumber="74" value="2000-05-04">
|
||
<collectingDate id="EFAEE9B5FF99FFB8FEA9FB4D5B8D3935" box="[640,738,1256,1282]" pageId="73" pageNumber="74" value="2000-05-04">4.v.2000</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A1534BFF99FFB8FF37FB4D5ADF3935" box="[798,944,1256,1283]" pageId="73" pageNumber="74">G. L. Challet</collectorName>
|
||
(1
|
||
<collectionCode id="ED45AE58FF99FFB8FFFCFB4D5D783935" box="[981,1047,1256,1282]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="73" pageNumber="74" type="Museum">MCZ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FF99FFBBFCEEFAA95B653DE6" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="discussion">
|
||
<paragraph id="8BEB369DFF99FFB8FCEEFAA9586039EC" blockId="73.[151,1437,860,2039]" pageId="73" pageNumber="74">
|
||
<emphasis id="B920EA8FFF99FFB8FCEEFAA958233911" bold="true" box="[199,332,1292,1318]" pageId="73" pageNumber="74">Discussion.</emphasis>
|
||
When species are placed into synonymy it is contingent upon experienced taxonomists to clearly discuss and illustrate the characters that are known to be highly diagnostic in that taxon. In
|
||
<taxonomicName id="4C544D1EFF99FFB8F88CFA955C5B397D" authorityName="Mulsant" authorityYear="1844" box="[1189,1332,1328,1354]" class="Insecta" family="Hydraenidae" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="family">Hydraenidae</taxonomicName>
|
||
the male genitalia are widely known as the most highly diagnostic character to use to differentiate species. For this reason,
|
||
<bibRefCitation id="EFC54B6CFF99FFB8FCBEFADD581639A5" author="Perkins, P. D." box="[151,377,1400,1426]" pageId="73" pageNumber="74" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 278)</bibRefCitation>
|
||
illustrated the male genitalia of representatives of all forms that were placed into synonymy, including the junior synonyms of
|
||
<taxonomicName id="4C544D1EFF99FFB8FE38FA385A0D3981" box="[529,866,1436,1462]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FE38FA385B423981" box="[529,557,1437,1462]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FE15FA395B973981" box="[572,760,1436,1462]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FF21FA395A0D3981" box="[776,866,1436,1462]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
, including examples from
|
||
<collectingCountry id="F343760DFF99FFB8F8A6FA395DB93981" box="[1167,1238,1436,1462]" name="Brazil" pageId="73" pageNumber="74">Brazil</collectingCountry>
|
||
, Florida,
|
||
<collectingCountry id="F343760DFF99FFB8F969FA395CF73981" box="[1344,1432,1436,1462]" name="Mexico" pageId="73" pageNumber="74">Mexico</collectingCountry>
|
||
, and Texas.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFF99FFB8FCEEFA415A7F3A5D" blockId="73.[151,1437,860,2039]" pageId="73" pageNumber="74">
|
||
Contrastingly,
|
||
<bibRefCitation id="EFC54B6CFF99FFB8FD47FA41589A39C9" author="Jach, M. A." box="[366,501,1508,1534]" pageId="73" pageNumber="74" pagination="301 - 308" refId="ref49627" refString="Jach, M. A. (1994) Descriptions of new species of Hydraenidae (Coleoptera) from Texas, Indiana, and Oklahoma, with faunistic and taxonomic notes on the family in Texas. The Coleopterists Bulletin, 48 (4), 301 - 308." type="journal article" year="1994">Jäch (1994)</bibRefCitation>
|
||
, when publishing a supposed synonymy, did not illustrate any male genitalia, and significantly did not discuss the marked differences in the male genitalia of
|
||
<taxonomicName id="4C544D1EFF99FFB8F83FF9AC5C2C3A15" authorityName="Perkins" authorityYear="1980" box="[1046,1347,1544,1570]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8F83FF9AC5D5D3A15" box="[1046,1074,1545,1570]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8F86CF9AD5C6E3A15" box="[1093,1281,1544,1570]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8F93FF9AD5C2C3A15" box="[1302,1347,1544,1570]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D1EFF99FFB8F9A8F9AC58A93A71" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8F9A8F9AC5CF23A15" box="[1409,1437,1545,1570]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FC88F98958323A71" box="[161,349,1580,1606]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FD45F98958A93A71" box="[364,454,1580,1606]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
that were illustrated by
|
||
<bibRefCitation id="EFC54B6CFF99FFB8FEFAF9895ADC3A71" author="Perkins, P. D." box="[723,947,1580,1606]" pageId="73" pageNumber="74" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980: 278)</bibRefCitation>
|
||
. He also did not examine the
|
||
<typeStatus id="54EF883FFF99FFB8F8D2F9895C323A71" box="[1275,1373,1580,1606]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C544D1EFF99FFB8F9A8F98858F53A5D" authorityName="Perkins" authorityYear="1980" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8F9A8F9885CF23A71" box="[1409,1437,1581,1606]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FC88F9F558323A5D" box="[161,349,1616,1642]" italics="true" pageId="73" pageNumber="74">Gymnochthebius</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FD44F9F558F53A5D" box="[365,410,1616,1642]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
. These are significant omissions.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFF99FFB8FCEEF9D05A583AE0" blockId="73.[151,1437,860,2039]" pageId="73" pageNumber="74">
|
||
Even though the differences in the male genitalia of the two species were illustrated adequately by line drawings in
|
||
<bibRefCitation id="EFC54B6CFF99FFB8FC9DF93D58333A85" author="Perkins, P. D." box="[180,348,1688,1714]" pageId="73" pageNumber="74" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins (1980)</bibRefCitation>
|
||
, additional documentation, in the form of photographic images, is given herein, in order to make the differences abundantly clear to all students of the group.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFF99FFBBFCEEF945580A3CE2" blockId="73.[151,1437,860,2039]" lastBlockId="74.[151,1437,151,465]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
|
||
Aedeagal characters differentiating the two species: (1) In
|
||
<taxonomicName id="4C544D1EFF99FFB8FF76F9445A8A3ACD" authorityName="Perkins" authorityYear="1980" box="[863,997,1760,1786]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FF76F9445A143ACD" box="[863,891,1761,1786]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FFA2F9455AC83ACD" box="[907,935,1760,1786]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FF91F9455A8A3ACD" box="[952,997,1760,1786]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2A18FF99FFB8FFDCF9455D273ACD" box="[1013,1096,1760,1786]" captionStart="FIGURES34–35" captionStartId="58.[151,264,1792,1816]" captionTargetBox="[186,1395,184,1753]" captionTargetId="figure-35@58.[166,1421,181,1768]" captionTargetPageId="58" captionText="FIGURES34–35.Habitusandmalegenitalia.34.Ochthebius(Gymnochthebius)falli(Perkins).35.Ochthebius(Gymnochthebius) fossatus LeConte." figureDoi="http://doi.org/10.5281/zenodo.10145174" httpUri="https://zenodo.org/record/10145174/files/figure.png" pageId="73" pageNumber="74">Fig. 34</figureCitation>
|
||
) the internal tubule is straight over the midlength, then has an abrupt bend in the distal ¼. In
|
||
<taxonomicName id="4C544D1EFF99FFB8FF6EF8A05A993B29" box="[839,1014,1796,1822]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FF6EF8A05A0C3B29" box="[839,867,1797,1822]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FF5BF8A15AE23B29" box="[882,909,1796,1822]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FFB5F8A15A993B29" box="[924,1014,1796,1822]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="136F2A18FF99FFB8F82CF8A15D3A3B28" box="[1029,1109,1796,1823]" captionStart="FIGURES34–35" captionStartId="58.[151,264,1792,1816]" captionTargetBox="[186,1395,184,1753]" captionTargetId="figure-35@58.[166,1421,181,1768]" captionTargetPageId="58" captionText="FIGURES34–35.Habitusandmalegenitalia.34.Ochthebius(Gymnochthebius)falli(Perkins).35.Ochthebius(Gymnochthebius) fossatus LeConte." figureDoi="http://doi.org/10.5281/zenodo.10145174" httpUri="https://zenodo.org/record/10145174/files/figure.png" pageId="73" pageNumber="74">Fig. 35</figureCitation>
|
||
) the internal tubule is arcuate for the entire length, without a bend in the distal ½. (2) The lumen aperture of
|
||
<taxonomicName id="4C544D1EFF99FFB8FFD5F88C5D103B75" authorityName="Perkins" authorityYear="1980" box="[1020,1151,1832,1858]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FFD5F88C5D783B75" box="[1020,1047,1833,1858]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8F80EF88D5D2D3B75" box="[1063,1090,1832,1858]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8F87BF88D5D103B75" box="[1106,1151,1832,1858]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
has a more oval-elongate shape, whereas the aperture is as wide as long in
|
||
<taxonomicName id="4C544D1EFF99FFB8FEE6F8E85AEB3B51" box="[719,900,1868,1894]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FEE6F8E85B843B51" box="[719,747,1869,1894]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FED4F8E95A773B51" box="[765,792,1868,1894]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FF03F8E95AEB3B51" box="[810,900,1868,1894]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
. (3) The strongly sclerotized distal ends of the main-piece of
|
||
<taxonomicName id="4C544D1EFF99FFB8FD11F8D458D43BBD" authorityName="Perkins" authorityYear="1980" box="[312,443,1904,1930]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FD11F8D4583B3BBD" box="[312,340,1905,1930]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FD4AF8D558113BBD" box="[355,382,1904,1930]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FDA7F8D558D43BBD" box="[398,443,1904,1930]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
are more narrowly separated than those of
|
||
<taxonomicName id="4C544D1EFF99FFB8FFB5F8D45D233BBD" box="[924,1100,1904,1930]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FFB5F8D45AD83BBD" box="[924,951,1905,1930]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FFEEF8D55A8D3BBD" box="[967,994,1904,1930]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FFDBF8D55D233BBD" box="[1010,1100,1904,1930]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
. (4) The main-piece, in dorsal or ventral views, compared to the length of the aedeagus, is wider in
|
||
<taxonomicName id="4C544D1EFF99FFB8FFA3F8305D633B99" authorityName="Perkins" authorityYear="1980" box="[906,1036,1940,1966]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FFA3F8305AC93B99" box="[906,934,1941,1966]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FF9CF8315ABF3B99" box="[949,976,1940,1966]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FFF6F8315D633B99" box="[991,1036,1940,1966]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
(l/w ca. 3.9) than in
|
||
<taxonomicName id="4C544D1EFF99FFB8F8C4F8305CF33B99" box="[1261,1436,1940,1966]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8F8C4F8305C663B99" box="[1261,1289,1941,1966]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8F931F8315C5C3B99" box="[1304,1331,1940,1966]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8F96BF8315CF33B99" box="[1346,1436,1940,1966]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
(l/w ca. 5.1). (5) The ratio of the width of the lumen opening and the maximum width of the main-piece is about 1:
|
||
<quantity id="4CAC9B78FF99FFB8FC8FF87959833BC1" box="[166,236,2012,2039]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" pageId="73" pageNumber="74" unit="in" value="5.5">5.5 in</quantity>
|
||
<taxonomicName id="4C544D1EFF99FFB8FCDAF878581A3BC1" authorityName="Perkins" authorityYear="1980" box="[243,373,2012,2038]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FCDAF87858613BC1" box="[243,270,2013,2038]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FD37F87958563BC1" box="[286,313,2012,2038]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FD61F879581A3BC1" box="[328,373,2012,2038]" italics="true" pageId="73" pageNumber="74">falli</emphasis>
|
||
</taxonomicName>
|
||
vs. about 1:
|
||
<quantity id="4CAC9B78FF99FFB8FDDCF8795B533BC1" box="[501,572,2012,2038]" metricMagnitude="-2" metricUnit="m" metricValue="6.604" pageId="73" pageNumber="74" unit="in" value="2.6">2.6 in</quantity>
|
||
<taxonomicName id="4C544D1EFF99FFB8FE6BF8785B993BC1" box="[578,758,2012,2038]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF99FFB8FE6BF8785B313BC1" box="[578,606,2013,2038]" italics="true" pageId="73" pageNumber="74">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF99FFB8FE44F8795BE63BC1" box="[621,649,2012,2038]" italics="true" pageId="73" pageNumber="74">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF99FFB8FEB1F8795B993BC1" box="[664,758,2012,2038]" italics="true" pageId="73" pageNumber="74">fossatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EA8FFF99FFB8FEDFF8795B953BC1" box="[758,762,2012,2038]" italics="true" pageId="73" pageNumber="74">.</emphasis>
|
||
The shapes and sizes of these structures are all integrated by behavior: the aedeagal tube is everted during copulation (
|
||
<bibRefCitation id="EFC54B6CFF9AFFBBFF0CFF325A843C86" author="Perkins, P. D." box="[805,1003,151,177]" pageId="74" pageNumber="75" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 20</bibRefCitation>
|
||
). These significant differences are not easily overlooked.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFF9AFFBBFCEEFF7A5B653DE6" blockId="74.[151,1437,151,465]" pageId="74" pageNumber="75">
|
||
The specimens used to illustrate these structural differences were collected at the same locality in
|
||
<collectingCountry id="F343760DFF9AFFBBF969FF7A5CF73CCE" box="[1344,1432,223,249]" name="Mexico" pageId="74" pageNumber="75">Mexico</collectingCountry>
|
||
, during the same collecting event. Although the aedeagi used to illustrate these structures are approximately the same length, the adults differ notably in size, the specimen of
|
||
<taxonomicName id="4C544D1EFF9AFFBBFF3EFE825AF13D76" authorityName="Perkins" authorityYear="1980" box="[791,926,295,321]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF9AFFBBFF3EFE825A5C3D77" box="[791,819,295,320]" italics="true" pageId="74" pageNumber="75">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF9AFFBBFF6DFE825A303D76" box="[836,863,295,321]" italics="true" pageId="74" pageNumber="75">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF9AFFBBFF58FE825AF13D76" box="[881,926,295,321]" italics="true" pageId="74" pageNumber="75">falli</emphasis>
|
||
</taxonomicName>
|
||
being larger (1.89 vs.
|
||
<quantity id="4CAC9B78FF9AFFBBF8B6FE835C6B3D76" box="[1183,1284,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="74" pageNumber="75" unit="mm" value="1.65">1.65 mm</quantity>
|
||
total length). There is considerable body size variation of
|
||
<taxonomicName id="4C544D1EFF9AFFBBFEB2FEEE5A3D3D52" box="[667,850,331,357]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF9AFFBBFEB2FEEE5BD83D53" box="[667,695,331,356]" italics="true" pageId="74" pageNumber="75">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF9AFFBBFEE3FEEE5B8A3D52" box="[714,741,331,357]" italics="true" pageId="74" pageNumber="75">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF9AFFBBFED1FEEE5A3D3D52" box="[760,850,331,357]" italics="true" pageId="74" pageNumber="75">fossatus</emphasis>
|
||
</taxonomicName>
|
||
over its extensive geographical range (see
|
||
<bibRefCitation id="EFC54B6CFF9AFFBBF961FEEE59BC3DBE" author="Perkins, P. D." pageId="74" pageNumber="75" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980</bibRefCitation>
|
||
). Specimens of
|
||
<taxonomicName id="4C544D1EFF9AFFBBFDAFFECA5B643DBE" authorityName="Perkins" authorityYear="1980" box="[390,523,367,393]" class="Insecta" family="Hydraenidae" genus="Gymnochthebius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="falli" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF9AFFBBFDAFFECA58CD3DBF" box="[390,418,367,392]" italics="true" pageId="74" pageNumber="75">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF9AFFBBFD9BFECA58A23DBE" box="[434,461,367,393]" italics="true" pageId="74" pageNumber="75">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF9AFFBBFDF7FECA5B643DBE" box="[478,523,367,393]" italics="true" pageId="74" pageNumber="75">falli</emphasis>
|
||
</taxonomicName>
|
||
average larger than the upper end of this size variation. Another difference in the figures is that the specimen of
|
||
<taxonomicName id="4C544D1EFF9AFFBBFDD5FE365BDE3D9A" box="[508,689,403,429]" class="Insecta" family="Hydraenidae" genus="Ochthebius" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="fossatus" subGenus="Gymnochthebius">
|
||
<emphasis id="B920EA8FFF9AFFBBFDD5FE365B783D9B" box="[508,535,403,428]" italics="true" pageId="74" pageNumber="75">H.</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFF9AFFBBFE00FE365B2A3D9A" box="[553,581,403,429]" italics="true" pageId="74" pageNumber="75">G.</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFF9AFFBBFE7EFE365BDE3D9A" box="[599,689,403,429]" italics="true" pageId="74" pageNumber="75">fossatus</emphasis>
|
||
</taxonomicName>
|
||
is teneral. This of course has no bearing on the decision that the specimens represent two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |