543 lines
78 KiB
XML
543 lines
78 KiB
XML
<document ID-CLB-Dataset="55905" ID-DOI="http://dx.doi.org/10.3897/zookeys.1097.83916" ID-GBIF-Dataset="e036b25d-83e3-4247-9c3b-41ad2a84b72a" ID-PMC="PMC9043006" ID-Pensoft-Pub="1313-2970-1097-1" ID-Pensoft-UUID="6593D2730A295F46B63A738B12E43A22" ID-PubMed="35837583" ID-ZooBank="66F17436DF364FFDB2D3021F14D40D62" ModsDocID="1313-2970-1097-1" checkinTime="1650371395093" checkinUser="pensoft" docAuthor="Antic, Dragan, Vagalinski, Boyan, Stoev, Pavel & Akkari, Nesrine" docDate="2022" docId="B925ACBEB25B532FA3DC5C4A5F3C16EA" docLanguage="en" docName="ZooKeys 1097: 1-46" docOrigin="ZooKeys 1097" docPubDate="2022-04-19" docSource="http://dx.doi.org/10.3897/zookeys.1097.83916" docTitle="Rhodopodesmus niveus gen. nov. et 2022, sp. nov." docType="treatment" docUuid="CC18A689-89BD-4197-922D-9CB3D8B3EA77" docUuidSource="ZooBank" docVersion="6" id="6593D2730A295F46B63A738B12E43A22" lastPageNumber="1" masterDocId="6593D2730A295F46B63A738B12E43A22" masterDocTitle="A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="1" updateTime="1732687488807" updateUser="ExternalLinkService">
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<mods:titleInfo id="F36CDB5814D9FB56EAF5CB873EE4E315">
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<mods:title id="96D638A950C411D34F9FC7DFCA9DF609">A review of the cavernicolous Trichopolydesmidae (Diplopoda, Polydesmida) from the Carpathian-Balkan arch and the Rhodope Mountains, with descriptions of two new genera and three new species</mods:title>
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<mods:namePart id="7767A8E0E5035BBF534D82630E31E023">Antic, Dragan</mods:namePart>
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<mods:nameIdentifier id="77B6C4F3CDE5E5DF9E06D7E1D2CB472B" type="ORCID">https://orcid.org/0000-0002-1231-4213</mods:nameIdentifier>
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<mods:affiliation id="6822114B8506FFB3AB3A5B374A4B27AD">University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski trg 16, 11000 Belgrade, Serbia & Serbian Biospeleological Society, Trg Dositeja Obradovica 2, 21000 Novi Sad, Serbia</mods:affiliation>
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<mods:nameIdentifier id="5A2DB81A27939DF719F5EDCE0857A7E4" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="EA985C4A638B4F17107DDF3AFD7509D1">Vagalinski, Boyan</mods:namePart>
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<mods:affiliation id="0DABC1E4835B2A7C66BA1B169B8CAFEC">Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
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<mods:namePart id="9BA2BEE8EFBC57AB4D3BA3386FF62267">Stoev, Pavel</mods:namePart>
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<mods:nameIdentifier id="D26E809C1D58961E75EDD324189B289B" type="ORCID">https://orcid.org/0000-0002-5702-5677</mods:nameIdentifier>
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<mods:affiliation id="726CF7EE72558D44EC2A14D8B2CBC1DE">National Museum of Natural History, Bulgarian Academy of Sciences, Tsar Osvoboditel Blvd. 1, Sofia 1000, Bulgaria & Pensoft Publishers, Sofia, Bulgaria</mods:affiliation>
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<mods:namePart id="783A678270B085CAEE4BEFB4391CCC48">Akkari, Nesrine</mods:namePart>
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<mods:nameIdentifier id="1B9010CF72F590002E0F22B60D1DBC3B" type="ORCID">https://orcid.org/0000-0001-5019-4833</mods:nameIdentifier>
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<mods:title id="CC93F7D671671B57697AE042B68A8C3E">ZooKeys</mods:title>
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<mods:date id="11EC4D66E320F89D3B4E21E4264CF81D">2022</mods:date>
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<mods:number id="95DF577C5951C0C59AB5FB9B26A9607D">2022-04-19</mods:number>
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<mods:number id="B699FEF79EC1C67393DA85FA3447AAAD">1097</mods:number>
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<mods:url id="FF38B437A78A97900F153733D1A33A5A">http://dx.doi.org/10.3897/zookeys.1097.83916</mods:url>
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<mods:classification id="4F960984E8C2ECAB5FB03DA90C1938F0">journal article</mods:classification>
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<mods:identifier id="BB92BB9CF900414D8A7C867A01BE4D33" type="DOI">http://dx.doi.org/10.3897/zookeys.1097.83916</mods:identifier>
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<treatment id="B925ACBEB25B532FA3DC5C4A5F3C16EA" ID-GBIF-Taxon="194660824" LSID="urn:lsid:zoobank.org:act:CC18A689-89BD-4197-922D-9CB3D8B3EA77" httpUri="http://treatment.plazi.org/id/B925ACBEB25B532FA3DC5C4A5F3C16EA" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Diplopoda" scope_family="Trichopolydesmidae" scope_order="Polydesmida">
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<subSubSection id="5EE6470F8386946C15C1EC20B9374A86" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="D3B3F746A73AA6E66A66FF17C6C84CDC" pageId="0" pageNumber="1">
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<taxonomicName id="EB57A1876ABFE3A073394AC90DBC6EAB" ID-CoL="8ZDSX" LSID="http://zoobank.org/CC18A689-89BD-4197-922D-9CB3D8B3EA77" authority="gen. nov. et" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus" status="sp. nov.">Rhodopodesmus niveus gen. nov. et</taxonomicName>
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<taxonomicNameLabel id="56ECC1771BDCD65449871B17B3A4D021" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="84B793BAA91A76DBDF0FF9AEAFDC4F55" pageId="0" pageNumber="1" type="description">
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<paragraph id="22AF5A74086D1FD43F0B1C8A14E71E01" pageId="0" pageNumber="1">
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<figureCitation id="72CD8C8342F13353CB78C1F58DE5ABFB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">Figs 14</figureCitation>
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<figureCitation id="4B4DE93A1B789C0AC8563ABDC1327913" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">, 15</figureCitation>
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<figureCitation id="607731594DAC9F7D1B091140B6176F27" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ left gonopod (NMNHS- 10823) A mesal view B lateral view C disto-lateral view D antero-lateral view E postero-distal view F anterior view. Abbreviations: a acropodite, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, pf prefemorite, psp proximal solenomeral process, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure16" httpUri="https://binary.pensoft.net/fig/674294" pageId="0" pageNumber="1">, 16</figureCitation>
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<figureCitation id="0B64C4E6AD8C95A4CB1E416939A4984A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">, 17K</figureCitation>
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||
<figureCitation id="A1BE3E3372889AB4F40B878CD4AEEFC8" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">, 18</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="5B88A55D7EDE0A4460E50F2A3B495047" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="671C10220D9881AB19ECB2892CFD45A6" pageId="0" pageNumber="1">
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||
<taxonomicName id="36E52D234052EF01ECFD073D857F19D4" authority="Antić & Vagalinski & Stoev & Akkari, 2022" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="sp. nov.">Bacillidesmus</taxonomicName>
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sp. nov.-
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<bibRefCitation id="4A8EB5658B1B339FC83A83C8288A0680" author="Vagalinski, B" editor="Beron, P" journalOrPublisher="4. Biodiversity of Western Rhodopes (Bulgaria and Greece) II. Pensoft and National Museum of Natural History, Sofia" pageId="0" pageNumber="1" pagination="129 - 143" refId="B62" refString="Vagalinski, B, Stoev, P, 2011. An annotated checklist of the myriapods of the Western Rhodopes Mts (Bulgaria and Greece). In: Beron, P, Ed., Biodiversity of Bulgaria. 4. Biodiversity of Western Rhodopes (Bulgaria and Greece) II. Pensoft and National Museum of Natural History, Sofia: 129 - 143" title="An annotated checklist of the myriapods of the Western Rhodopes Mts (Bulgaria and Greece)." volumeTitle="Biodiversity of Bulgaria." year="2011">Vagalinski and Stoev 2011</bibRefCitation>
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||
: 135.
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||
</paragraph>
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||
<paragraph id="6E7A13E78785E3597D39ACFADD167FF4" pageId="0" pageNumber="1">
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||
<taxonomicName id="5B524FB414561F69DA79464DAD331AC2" authority="Antić & Vagalinski & Stoev & Akkari, 2022" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus" status="sp. nov.">Bacillidesmus</taxonomicName>
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sp. [nov.]-
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<bibRefCitation id="112BFFD7526FCFA4DEF81C559555C004" author="Beron, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" refId="B10" refString="Beron, P, 2015. Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." title="Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." year="2015">Beron 2015</bibRefCitation>
|
||
: 80.
|
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</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="39C5F84DAC728B6EC256E75875428C1A" pageId="0" pageNumber="1" type="diagnosis">
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<paragraph id="B53C368E692388B4B08551685A5C0F97" pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph id="3D987F09AEEB5D821BF8A55B11B8F179" pageId="0" pageNumber="1">As for the monospecific genus.</paragraph>
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</subSubSection>
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<subSubSection id="AF62D6657144CADD3A7CC7444EBBF780" pageId="0" pageNumber="1" type="name">
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||
<paragraph id="BD3B6B7D817D447C7C1B0B3A84316F3B" pageId="0" pageNumber="1">Name.</paragraph>
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<paragraph id="D0644B8A712F5BA3EB9CCD5B07557A32" pageId="0" pageNumber="1">
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The specific name is a Latin adjective;
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<emphasis id="85F18EBC5DAB73A6EA03EA697EAA49DA" italics="true" pageId="0" pageNumber="1">niveus</emphasis>
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refers to the snow-white body color of the living specimens. Furthermore, the name of the type locality, cave Snezhanka, in Bulgarian means Snow White, the heroine from the fairy tale of the Brothers Grimm.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5A47FA09AF1B4DDC7B2109031DC8A29" pageId="0" pageNumber="1" type="materials_examined">
|
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<paragraph id="D4A15C0EBBF24907B6F1801C8B0973F0" pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph id="D37AB29CA5288A5B75133C42FD214268" pageId="0" pageNumber="1">
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<materialsCitation id="460E3F9BC8BA32062901D8925FE5B9B4" collectingDate="2020-10-26" collectorName="D. Antic, B. Vagalinski." country="Bulgaria" county="Peshtera Municipality" latitude="42.00222" location="Snezhanka Cave" longLatPrecision="1" longitude="24.27764" municipality="Peshtera" specimenCount="♂" stateProvince="Pazardzhik District" typeStatus="Holotype">
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<emphasis id="4C520BAC2DDCF4CBD06FDC468E48117F" bold="true" italics="true" pageId="0" pageNumber="1">
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<typeStatus id="418A2FCE74C544F2C7A6E14F8C483ECC">Holotype</typeStatus>
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</emphasis>
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<specimenCount id="6DB614AF6DB1B3B3A54F9837AD1D3045">♂</specimenCount>
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(NMNHS-10823),
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<collectingCountry id="06B15C2B575DD0AB14B99B615F8A1E76" name="Bulgaria">Bulgaria</collectingCountry>
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,
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<collectingRegion id="B7E692078B3713E139F6B5E1562BBB59" country="Bulgaria" name="Pazardzhik">Pazardzhik District</collectingRegion>
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,
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||
<collectingCounty id="58C4AF5D458B2583F902EA34571E736B">Peshtera Municipality</collectingCounty>
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,
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<collectingMunicipality id="EC6A9B349CE9CDDCF1AEA8FF1633B760">Peshtera</collectingMunicipality>
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||
,
|
||
<location id="14836C2557B00B5A5B1505FC8907E230" LSID="urn:lsid:plazi:treatment:B925ACBEB25B532FA3DC5C4A5F3C16EA:14836C2557B00B5A5B1505FC8907E230" country="Bulgaria" county="Peshtera Municipality" latitude="42.00222" longLatPrecision="1" longitude="24.27764" municipality="Peshtera" name="Snezhanka Cave" stateProvince="Pazardzhik District">Snezhanka Cave</location>
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,
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<geoCoordinate id="57BAD6EEC9792FDF2BF866ACA6C7BEFF" degrees="42.00222" direction="north" orientation="latitude" precision="1" value="42.00222">N 42.00222</geoCoordinate>
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,
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<geoCoordinate id="ADDD302E23B5AD8198EE6CC3B7B6C6D6" degrees="24.27764" direction="east" orientation="longitude" precision="1" value="24.27764">E 24.27764</geoCoordinate>
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,
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<collectingDate id="7AD4791B5EFA1F56A47305565AF434D3" value="2020-10-26">26.X.2020</collectingDate>
|
||
, leg.
|
||
<collectorName id="C6C0AF494F4839B0C96F5C6E4EFBB9AA">
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D.
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<normalizedToken id="3C906E5035CCDD28B3F55AB791D96D66" originalValue="Antić">Antic</normalizedToken>
|
||
</collectorName>
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||
&
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||
<collectorName id="A11AFF5939FE3C8C584D6778E4E5CCED">B. Vagalinski.</collectorName>
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</materialsCitation>
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</paragraph>
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<paragraph id="BEBA28022A52F8C071E9576DAD12B1AF" pageId="0" pageNumber="1">
|
||
<materialsCitation id="1F501763A189463F1C357D007C2AC9FD" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="D. Antic & B. Vagalinski." country="Bulgaria" county="Peshtera Municipality" latitude="42.00222" location="Snezhanka Cave" longLatPrecision="1" longitude="24.27764" municipality="Peshtera" specimenCount="3" specimenCount-female="3" stateProvince="Pazardzhik" typeStatus="Paratypes">
|
||
<emphasis id="30D2A500843D23EB657799DC8E5B07A9" bold="true" pageId="0" pageNumber="1">
|
||
<emphasis id="73DD2E74C10FD105B29B5033D6DA9FDF" italics="true" pageId="0" pageNumber="1">
|
||
<typeStatus id="D71970426D1541C6FFB70B261376F20D">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="F7D5C0472A0A97E4D1DFD41EBB838947" type="female">3 ♀♀</specimenCount>
|
||
(one used for SEM, NMNHS-10824-10826), same data as for holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="B7C3499235E182CDD0F2ECBA16FB80E7" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="D. Antic & B. Vagalinski." country="Bulgaria" county="Peshtera Municipality" latitude="42.00222" location="Snezhanka Cave" longLatPrecision="1" longitude="24.27764" municipality="Peshtera" specimenCount="1" specimenCount-female="1" stateProvince="Pazardzhik" typeStatus="paratype">
|
||
<specimenCount id="3C7C6DC7E462B80A4367A623E6871EA9" type="female">1 ♀</specimenCount>
|
||
(used for SEM, NHMW MY10259) same data as for holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8982DE056B1F5A664D029B881C1FA4A" pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph id="DDC52CAD6A5E56CFACD7A9A77C5EED2A" pageId="0" pageNumber="1">Additional material.</paragraph>
|
||
<paragraph id="3C234D6D416898CFFBE888DC0F17EBA7" pageId="0" pageNumber="1">
|
||
<specimenCount id="9225F2004FF71750CFBED82ED71C0750" type="male">1 ♂</specimenCount>
|
||
(fragments, one gonopod available, NMNHS-10827),
|
||
<specimenCount id="DFF0918E10EC08E6C04BD2BF48FCE70E" type="female">1 ♀</specimenCount>
|
||
(fragments, NMNHS-10828), both fragments, 1 whole juvenile (NMNHS-10829), same cave but
|
||
<date id="55A4C264AB367FD1D3AF7D9D39C18875" value="2005-09-18">18.IX.2005</date>
|
||
, leg. P. Beron.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E54A836267F8AE5D3B98C36236A81341" pageId="0" pageNumber="1" type="description">
|
||
<paragraph id="A163EDA32B48B6C5240F5C8D99EA0A09" pageId="0" pageNumber="1">Description.</paragraph>
|
||
<paragraph id="3CF237AF6C22D10D8B6EE45B637651B4" pageId="0" pageNumber="1">
|
||
<emphasis id="253BA54ADDB4686E4A71913A09D355EF" bold="true" italics="true" pageId="0" pageNumber="1">Number of body rings and measurements</emphasis>
|
||
: Body with 19 rings (including telson) in adults, moniliform (Fig.
|
||
<figureCitation id="0FDFCFEF6453B65C6C5E90583C7180C7" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">14</figureCitation>
|
||
). Holotype male 5.3 mm long, width of midbody pro- and metazonae 0.35 mm and 0.50 mm, respectively. Paratype females 4.3-5.4 mm long, width of midbody pro- and metazonae 0.35-0.40 mm and 0.45-0.60 mm, respectively.
|
||
</paragraph>
|
||
<caption id="D7C0D2DD2F8EB7A11763EF88315F3091" doi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1" start="Figure 14" startId="F14">
|
||
<paragraph id="5AE358E441FDD04D0D5589462C00839F" pageId="0" pageNumber="1">
|
||
<emphasis id="F97BA0FE45B441D65B4C5C716B15A073" bold="true" pageId="0" pageNumber="1">Figure 14.</emphasis>
|
||
<taxonomicName id="B91E9553BDAB9198DB46DE61226D8BA1" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="1011D81D0EBB403BEA4172035516A961" italics="true" pageId="0" pageNumber="1">Rhodopodesmus niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov., holotype ♂ (
|
||
<emphasis id="EEC98D8ECC14B4448BEB21ECFD29270B" bold="true" pageId="0" pageNumber="1">A-D</emphasis>
|
||
, NMNHS-10823) and paratype ♀ (
|
||
<emphasis id="B08CFB245A53C875A9270EF956A01B35" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
, NHMW MY10259) habitus
|
||
<emphasis id="741FE4C11F2E89588639FA19723A267E" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
in situ, dorsal view (photo H. Reip)
|
||
<emphasis id="4A8FA0970B19164F53A8927FC911E3EB" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
lateral view
|
||
<emphasis id="55DA4C641A68936ED7149DB835F5906D" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
anterior part of body, lateral view
|
||
<emphasis id="0AF39690F38A8A25AD213DB7799AA9FF" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
posterior part of body, lateral view
|
||
<emphasis id="43F253BFCC3E994C0C45DE5891FA6224" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
lateral view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BA33970414F5231821D071BFCEC8B868" pageId="0" pageNumber="1">
|
||
<emphasis id="FD5AD72C490535C733D47B21202B43CA" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
|
||
: Living animals snow white (Fig.
|
||
<figureCitation id="AB94F12ED703CFB30EBB1E21FA40171F" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">14A</figureCitation>
|
||
), slightly translucent. Yellowish in alcohol (Fig.
|
||
<figureCitation id="FA1DBDC998FC8AA1B2485C1D409F15E3" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">14B-E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="81D4A9969A43A974AC7E73CB304AF41E" pageId="0" pageNumber="1">
|
||
<emphasis id="3FE67A1D517B6E4718FEF8ABF27421CA" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
: Broader than collum, setose; epicranial suture poorly developed; isthmus between antennae ≈ 1.7
|
||
<normalizedToken id="2516BF52ACFB175D8ADA011DB3E2AEDB" originalValue="×">x</normalizedToken>
|
||
broader than diameter of antennal socket (Fig.
|
||
<figureCitation id="D17C0B7B54FFEDD5FAC91655B17866FD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15A, D</figureCitation>
|
||
). Labrum with three labral teeth, and with 3+3 labral and five supralabral setae (Fig.
|
||
<figureCitation id="DF8B8C2312513AB5A27C4214E0096F42" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15A</figureCitation>
|
||
). Gnathochilarium without peculiarities. Antennae rather short, clavate (Figs
|
||
<figureCitation id="0D91BB8854F6E740CE776FC230E2BF45" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">14A, C</figureCitation>
|
||
,
|
||
<figureCitation id="0C3868B4CE2F2BE5DA101732F9A804BD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15B, C</figureCitation>
|
||
). Antennomere length 6> 2 = 3 = 4> 5> 7 = 1. Antennae 0.8 mm long in holotype male; length/breadth ratios of antennomeres 1-7: 1 (1), 2 (2), 3 (2), 4 (2), 5 (1), 6 (1.5) and 7 (1). Antennomere 6 with four sensilla trichodea and with strongly developed disto-dorsal pit with numerous long sensilla basiconica partially exposed outside the pit (Fig.
|
||
<figureCitation id="58E7F1F3EE437DA4F18B1DA769854835" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15C</figureCitation>
|
||
). Antennomere 7 with one sensillum trichoideum and a small bulge with three sensilla basiconica spiniformia (Fig.
|
||
<figureCitation id="12ABE8DB72FB24E7F8224EE85CC31D10" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15B, C</figureCitation>
|
||
). Four apical cones (Fig.
|
||
<figureCitation id="BA683605DB996D9ED66C75E53585EEAE" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="FA58F7B8B30A5EC252E12664767225CB" doi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1" start="Figure 15" startId="F15">
|
||
<paragraph id="74E36A453E633CE875527006AC57791A" pageId="0" pageNumber="1">
|
||
<emphasis id="5C8B60FBE479EC852F91B091B2DEB77E" bold="true" pageId="0" pageNumber="1">Figure 15.</emphasis>
|
||
<taxonomicName id="7F27349F057A1ED70F7A8120D5806E80" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="319F095163CAC334CBE604B74AE76A29" italics="true" pageId="0" pageNumber="1">Rhodopodesmus niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY10259)
|
||
<emphasis id="912785E6E79E8C1A4FCE3EA56FA92F4D" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
head, anterior view
|
||
<emphasis id="1DD2B5AF0336E94D0B1C46D369F38B0A" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
right antenna, lateral view
|
||
<emphasis id="ECE8627A296A802A19AF6F2B0CFAF818" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia)
|
||
<emphasis id="B5FD85A74A974E93D74A5E55E791148F" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
anterior part of body, dorsal view
|
||
<emphasis id="261D48EF2EE238F4130C17FD7F318B86" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
body rings 8 and 9, dorsal view
|
||
<emphasis id="EFC76D372097496AAF95B23E94F554A7" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
anterior part of body, dorso-lateral view
|
||
<emphasis id="EEA66F06485ECC8D6EBA4C5734C9BF00" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
right ozopore 9, lateral view
|
||
<emphasis id="17095A02F4CEC7A3E6679D4396F07CF8" bold="true" pageId="0" pageNumber="1">H, I</emphasis>
|
||
body rings 17-19, dorsal and lateral views, respectively
|
||
<emphasis id="2E4E3935EE253A36F598F101F7520178" bold="true" pageId="0" pageNumber="1">J</emphasis>
|
||
body rings 8-10. Scale bars: 0.1 mm (
|
||
<emphasis id="F6BF55FD862C1B545168B39EC92FA0BC" bold="true" pageId="0" pageNumber="1">A, B, D-F, H-J</emphasis>
|
||
), 0.05 mm (
|
||
<emphasis id="8E9A03E24BA108F8EBAEB9531936116D" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), 0.02 mm (
|
||
<emphasis id="3F3C8BC6279B2E6205A74B4F83141CD4" bold="true" pageId="0" pageNumber="1">G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DEACE51732C0FC5EBA9DA0F3DAF3875F" pageId="0" pageNumber="1">
|
||
<emphasis id="32FE5000BF5BECB5AF95D59F8E8E7CCB" bold="true" italics="true" pageId="0" pageNumber="1">Collum</emphasis>
|
||
: Semi-circular, with one or two lateral incisions and ≈ 6 irregular rows of medium sized and trichoid setae (Fig.
|
||
<figureCitation id="671378B29619D60223D9E2ECB5D9DDBA" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2EC8C1799BE52D3FD18D722B31C19E73" pageId="0" pageNumber="1">
|
||
<emphasis id="A40F8BA94AAAE84505F2E9B18D38100F" bold="true" italics="true" pageId="0" pageNumber="1">Body rings</emphasis>
|
||
: Tegument shining, texture alveolate, reticulate and scaly. Rings densely setose (Fig.
|
||
<figureCitation id="34948CE57F8A53DDA1B229DA5DA4F933" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ (A-D, NMNHS- 10823) and paratype ♀ (E, NHMW MY 10259) habitus A in situ, dorsal view (photo H. Reip) B lateral view C anterior part of body, lateral view D posterior part of body, lateral view E lateral view. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1097.83916.figure14" httpUri="https://binary.pensoft.net/fig/674292" pageId="0" pageNumber="1">14B-E</figureCitation>
|
||
). Setae medium sized and trichoid, originating from small tubercules (Fig.
|
||
<figureCitation id="E94DC9B4E0A5CA3CB43E8F2AA4D2C349" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15D-F, H-J</figureCitation>
|
||
). Posteriormost tubercules mostly with a small thorn (Fig.
|
||
<figureCitation id="2E7B75FC1707AA4454884557556DF621" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15E</figureCitation>
|
||
). Rings 2-4 with ≈ 4 rows of setae (Fig.
|
||
<figureCitation id="C4F4B3A35946F32058BE2A8C6BD6E952" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15D, F</figureCitation>
|
||
). Rings 5-18 with ≈ 6-8 irregular rows of setae (Fig.
|
||
<figureCitation id="837F9F931320787A23A466F563CE3FD3" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15D-F, H-J</figureCitation>
|
||
). Paraterga serrated, with 5-7 teeth (Fig.
|
||
<figureCitation id="817A008E22F9778E5A0360AAB850B735" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15E, H</figureCitation>
|
||
). Pore formula normal: 5, 7, 9, 10, 12, 13, 15-18. Poriferous metazonae with enlarged posterolateral cone bearing an ozopore and three medium-sized setae (Fig.
|
||
<figureCitation id="E689A5CB6E197E025BEB107A6476929B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15G</figureCitation>
|
||
). Epiproct triangular in dorsal view, directed slightly caudoventrad (Fig.
|
||
<figureCitation id="B31537A767133D0817082AEC2B59221C" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15H, I</figureCitation>
|
||
). Paraprocts semi-spherical, each with two long setae originating from small tubercules and ≈ 10 shorter setae without tubercules (Fig.
|
||
<figureCitation id="D28B90BA80C5F19058BCE907800A9960" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15I</figureCitation>
|
||
). Hypoproct trapeziform, with two long distal setae and numerous shorter setae throughout (Fig.
|
||
<figureCitation id="2D9B52248FD0A0853FC65CB0222522AC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15I</figureCitation>
|
||
). Sterna unmodified, poorly setose. Pleurosternal carinae absent, only a few small teeth present on rings 2 and 3 (Fig.
|
||
<figureCitation id="3FF149E8CB81845BCE9424BAE68FDDB8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rhodopodesmus niveus gen. nov. et sp. nov., paratype ♀ habitus (NHMW MY 10259) A head, anterior view B right antenna, lateral view C tip of right antenna, dorsal view (arrow indicates small bulge with sensilla basiconica spiniformia) D anterior part of body, dorsal view E body rings 8 and 9, dorsal view F anterior part of body, dorso-lateral view G right ozopore 9, lateral view H, I body rings 17 - 19, dorsal and lateral views, respectively J body rings 8 - 10. Scale bars: 0.1 mm (A, B, D-F, H-J), 0.05 mm (C), 0.02 mm (G)." figureDoi="10.3897/zookeys.1097.83916.figure15" httpUri="https://binary.pensoft.net/fig/674293" pageId="0" pageNumber="1">15F</figureCitation>
|
||
). Gonopod aperture large, subsemi-circular.
|
||
</paragraph>
|
||
<paragraph id="EA1A6E9B7687AF70E8A204EDAF5343ED" pageId="0" pageNumber="1">
|
||
<emphasis id="D5A2A7CD36E6150636785E9B48D53DF2" bold="true" italics="true" pageId="0" pageNumber="1">Walking legs</emphasis>
|
||
: Legs 1-3 in males with swollen femur, especially femur 2; coxa 2 with a short mesal apophysis (cf.
|
||
<bibRefCitation id="A29B9855212C433F373EFDC4B5CD29F6" author="Strasser, K" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="1" pagination="325 - 385" refId="B54" refString="Strasser, K, 1966a. Ueber Diplopoden Bulgariens. Annales Zoologici 23 (12): 325 - 385" title="Ueber Diplopoden Bulgariens." volume="23" year="1966 a">Strasser 1966a</bibRefCitation>
|
||
: 341, fig. 13). No other peculiarities.
|
||
</paragraph>
|
||
<paragraph id="CC7B817DB0809F7746FC60F7A13EC013" pageId="0" pageNumber="1">
|
||
<taxonomicName id="E457BE7C08961E32143A3A9D160C62B8" genus="Gonopods" lsidName="Gonopods" pageId="0" pageNumber="1" rank="genus">
|
||
<emphasis id="AD18336BCD4054F429A37ED2C0D92815" bold="true" italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="41E8EA101CA5DCA7BC2D832DE15F3FEF" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Rhodopodesmus niveus gen. nov. et sp. nov., holotype ♂ left gonopod (NMNHS- 10823) A mesal view B lateral view C disto-lateral view D antero-lateral view E postero-distal view F anterior view. Abbreviations: a acropodite, ca cannula, cx coxa, dp distal projection of solenophore, dsp distal solenomeral process, ll lamella of solenophore, pf prefemorite, psp proximal solenomeral process, s solenomere, sb solenomeral branch, sf seminal fossa, sg seminal groove, sph solenophore. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure16" httpUri="https://binary.pensoft.net/fig/674294" pageId="0" pageNumber="1">16</figureCitation>
|
||
,
|
||
<figureCitation id="EE90F1BEC7D47E2C1C200C03B3035FE3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Gonopods of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and Rhodopes A-C Bacillidesmus filiformis (Latzel, 1884) lectotype ♂ (NHWM MY 3754) A after Attems (1898) B? left gonopod,? mesal view, after Attems (1940) C? left gonopod,? mesal view, present study D Banatodesmus jeanneli (Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after Tabacaru (1980) E, F Napocodesmus florentzae Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after Tabacaru (1975) G, H Trichopolydesmus eremitis Verhoeff, 1898 ♂ from Closani cave, right gonopod, antero-mesal and mesal views, respectively, after Ceuca (1958) I Balkanodesminus bulgaricus (Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS- 10813), left gonopod, mesal view J Balkanodesminus serbicus gen. nov. et sp. nov. paratype ♂ (NHMW MY 10263), left gonopod, mesal view K Rhodopodesmus niveus gen. nov. et sp. nov. holotype ♂ (NMNHS- 10823), left gonopod, mesal view L Balkanodesminus dentatus (Strasser, 1966 a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after Strasser (1966 a) M Balkanodesminus dentatoides gen. nov. et sp. nov. paratype ♂ (NHMW MY 10258), right gonopod, mesal view N Balkanodesminus dentatoides gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY 10267), left gonopod, mesal view. Scale bars: 0.05 mm." figureDoi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1">17K</figureCitation>
|
||
): Coxa (cx) large, semi-circular in ventral view, with deep gonocoel mesally; anterior third much lower than rest of coxa, shield-like, thus lateral, swollen and alveolate part rectangular in lateral view; with ≈ 15 setae. Cannula (ca) long, C-shaped. Telopodite long compared to coxa, consisting of a transverse, setose prefemorite (pf) and more or less C-shaped (in lateral and mesal views) acropodite (a) longitudinally divided in distal half into two branches, solenomeral branch (sb) and solenophore (sph). Solenomeral branch positioned mesally, transversely divided into three processes, besides solenomere (s) and distal solenomeral process (dsp), there is an additional, proximal solenomeral process (psp), more or less spatulate and forming C-shaped rift with solenomere. Both solenomere (s) and distal solenomeral process (dsp) long, slender, of same length, forming acute angle at bifurcation. Solenophore (sph) longer than solenomeral branch, characterized by a lateral lamella (ll) and a distal projection (dp). Lateral lamella (ll) with triangular lobe. Distal projection (dp) long, thin and twisted. Seminal groove (sg) starts from seminal fossa (sf) mesally on prefemorite, extends along mesal side of acropodite up to bifurcation of solenomeral branch and solenophore, then passes on lateral side of solenomeral branch, further proximally on solenomere, ending distally.
|
||
</paragraph>
|
||
<caption id="60BBED6E48ACE82535D27A1417558ACC" doi="10.3897/zookeys.1097.83916.figure16" httpUri="https://binary.pensoft.net/fig/674294" pageId="0" pageNumber="1" start="Figure 16" startId="F16">
|
||
<paragraph id="2798D258611B37CAA7D02FFBADD943EB" pageId="0" pageNumber="1">
|
||
<emphasis id="46EB8BC703C9D6A3300AF26AA1B1BF5B" bold="true" pageId="0" pageNumber="1">Figure 16.</emphasis>
|
||
<taxonomicName id="9F2C44AFDB3B3C38FD930B933C5A0AA5" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="FD483EB73474B5A706585FB4A58EF818" italics="true" pageId="0" pageNumber="1">Rhodopodesmus niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov., holotype ♂ left gonopod (NMNHS-10823)
|
||
<emphasis id="19F359773FE88AB54E8828E72D766678" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
mesal view
|
||
<emphasis id="B6F8543EF0E794C5A4C0086C8C361EC5" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
lateral view
|
||
<emphasis id="FC83F052F3C748D4312A09B458F4BED9" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
disto-lateral view
|
||
<emphasis id="E8A5518C89430AA0AFA1A66D6C7C5CC0" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
antero-lateral view
|
||
<emphasis id="5132671C51A9B098D545E1E39F14884B" bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
postero-distal view
|
||
<emphasis id="21B201F1CACC84559F9234312385359B" bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
anterior view. Abbreviations:
|
||
<emphasis id="CDFEE822BBB4DF0B1DDD8C19C631B752" bold="true" pageId="0" pageNumber="1">a</emphasis>
|
||
acropodite,
|
||
<emphasis id="ED7762BC638449C9AEEB0A1F745D2A40" bold="true" pageId="0" pageNumber="1">ca</emphasis>
|
||
cannula,
|
||
<emphasis id="B80B3FBCA8687F79857417B50B234EA5" bold="true" pageId="0" pageNumber="1">cx</emphasis>
|
||
coxa,
|
||
<emphasis id="4B9806109299ACE907A4458330B76D84" bold="true" pageId="0" pageNumber="1">dp</emphasis>
|
||
distal projection of solenophore,
|
||
<emphasis id="355025E516AFD1654706EC31E5D89291" bold="true" pageId="0" pageNumber="1">dsp</emphasis>
|
||
distal solenomeral process,
|
||
<emphasis id="B82143A391EB30925F5EE32B90AF15A1" bold="true" pageId="0" pageNumber="1">ll</emphasis>
|
||
lamella of solenophore,
|
||
<emphasis id="AE615857621C5CA7045EC1467D7128CC" bold="true" pageId="0" pageNumber="1">pf</emphasis>
|
||
prefemorite,
|
||
<emphasis id="B2304673B24C7E23E743034C347AE292" bold="true" pageId="0" pageNumber="1">psp</emphasis>
|
||
proximal solenomeral process,
|
||
<emphasis id="6D48AE1FEC6524ABE6BC24CC90CB05A7" bold="true" pageId="0" pageNumber="1">s</emphasis>
|
||
solenomere,
|
||
<emphasis id="8E4040863F6B1AAC2CB32FAD350B9A59" bold="true" pageId="0" pageNumber="1">sb</emphasis>
|
||
solenomeral branch,
|
||
<emphasis id="E6718DCC51096BD976D0876DD881479B" bold="true" pageId="0" pageNumber="1">sf</emphasis>
|
||
seminal fossa,
|
||
<emphasis id="A0FC4DB2807D0CC5C7CDBD0FE8AC3C26" bold="true" pageId="0" pageNumber="1">sg</emphasis>
|
||
seminal groove,
|
||
<emphasis id="58AC6F0818335815A906090D275D624C" bold="true" pageId="0" pageNumber="1">sph</emphasis>
|
||
solenophore. Scale bars: 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="0FABCE616A93E40AF4C3A0E3408B8B62" pageId="0" pageNumber="1" type="habitat">
|
||
<paragraph id="B14650C91DCA0972DB2F7E01CE3E39E0" pageId="0" pageNumber="1">Habitat.</paragraph>
|
||
<paragraph id="D335435CD9A6D335750A543AE43EB39B" pageId="0" pageNumber="1">
|
||
Snezhanka Cave consists of a single gallery forming six distinct halls with total length of 348 m. The entrance is located at 865 m a.s.l. The cave is rich in diverse sinter formations and sinter ponds. It was established as a natural monument in 1961, and has served as show cave since 1968 (
|
||
<bibRefCitation id="E12F4CFE21B7F402C072660208770FBF" author="Petrov, B" journalOrPublisher="UNDP, ' Rhodope project'" pageId="0" pageNumber="1" refId="B42" refString="Petrov, B, Stoev, P, 2007. The underground world of the mountain Rhodopes. UNDP, 'Rhodope project'" title="The underground world of the mountain Rhodopes." year="2007">Petrov and Stoev 2007</bibRefCitation>
|
||
). Most of the
|
||
<normalizedToken id="A290C48B6451FE1B2D2C5C0B7CA21E28" originalValue="cave’s">cave's</normalizedToken>
|
||
invertebrate fauna known at present includes either trogloxenes or troglophiles (
|
||
<bibRefCitation id="BAAFF39E6559CDE5555B9D027033065D" author="Beron, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" refId="B10" refString="Beron, P, 2015. Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." title="Cave Fauna of Bulgaria. East-West and National Museum of Natural History Publishers, 434 pp." year="2015">Beron 2015</bibRefCitation>
|
||
), with the exception of the local endemic
|
||
<taxonomicName id="7A8AE525266CDAFE7585567C7C2113E2" authorityName="Giachino; B. Gueorguiev & Vailati" authorityYear="2011" class="Insecta" family="Carabidae" genus="Paralovricia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paralovricia beroni" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="beroni">
|
||
<emphasis id="103AB2BE365E8DC37899CDE9759B61BF" italics="true" pageId="0" pageNumber="1">Paralovricia beroni</emphasis>
|
||
</taxonomicName>
|
||
Giachino,
|
||
<normalizedToken id="31A6B98C9367E39C59ADF22DFBFF725C" originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||
& Vailati, 2011 (
|
||
<taxonomicName id="0B6BD3CC0A2ED94AE2F7D032FD47C319" class="Insecta" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
||
,
|
||
<taxonomicName id="6FE8720C77B64C52A82A403074DF69F0" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
), which is considered a probable hypogean, although not typical troglobitic species (
|
||
<bibRefCitation id="CD3931BB131A29F6141D1A1DFB4EAAE8" DOI="https://doi.org/10.3897/zookeys.117.1612" author="Giachino, PM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="59 - 72" refId="B18" refString="Giachino, PM, Gueorguiev, B, Vailati, D, 2011. A new remarkable subterranean beetle of the Rhodopes: Paralovricia gen. n. beroni sp. n. belonging to Lovriciina new subtribe (Coleoptera, Carabidae, Trechinae, Bembidiini). ZooKeys 117: 59 - 72, DOI: https://doi.org/10.3897/zookeys.117.1612" title="A new remarkable subterranean beetle of the Rhodopes: Paralovricia gen. n. beroni sp. n. belonging to Lovriciina new subtribe (Coleoptera, Carabidae, Trechinae, Bembidiini)." url="https://doi.org/10.3897/zookeys.117.1612" volume="117" year="2011">Giachino et al. 2011</bibRefCitation>
|
||
). Another myriapod known from this cave is
|
||
<taxonomicName id="08F98EB06375CA25DDA61530FD73EEBB" authorityName="Verhoeff" authorityYear="1926" class="Chilopoda" family="Lithobiidae" genus="Lithobius" kingdom="Animalia" lsidName="Lithobius lakatnicensis" order="Lithobiomorpha" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lakatnicensis">
|
||
<emphasis id="2ACFF0C41B017891F55045177B1C5F68" italics="true" pageId="0" pageNumber="1">Lithobius lakatnicensis</emphasis>
|
||
</taxonomicName>
|
||
Verhoeff, 1926.
|
||
</paragraph>
|
||
<caption id="C5D6F35DB7808FDE2ED32C9444719530" doi="10.3897/zookeys.1097.83916.figure17" httpUri="https://binary.pensoft.net/fig/674295" pageId="0" pageNumber="1" start="Figure 17" startId="F17">
|
||
<paragraph id="A9E095807F9520BD48B184E4804F5332" pageId="0" pageNumber="1">
|
||
<emphasis id="5F6B0F85F3333760CB8DAE10BF03BE82" bold="true" pageId="0" pageNumber="1">Figure 17.</emphasis>
|
||
<taxonomicName id="013F7041B14FA43DD93DE651E08B8A9D" genus="Gonopods" lsidName="Gonopods" pageId="0" pageNumber="1" rank="genus">Gonopods</taxonomicName>
|
||
of cavernicolous
|
||
<taxonomicName id="728010EB6FBBEC19C2EF5F835430306A" authorityName="Verhoeff" authorityYear="1910" class="Diplopoda" family="Trichopolydesmidae" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
|
||
from the Carpathian-Balkan arch and Rhodopes
|
||
<emphasis id="30BD3524F6009A3C66555ACB5BBD4A71" bold="true" pageId="0" pageNumber="1">A-C</emphasis>
|
||
<taxonomicName id="04C9CE0718814C273C5ACF76A19F0B56" baseAuthorityName="Latzel" baseAuthorityYear="1884" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus filiformis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="filiformis">
|
||
<emphasis id="F33B9A3D11C4D8678558060BEA703BA5" italics="true" pageId="0" pageNumber="1">Bacillidesmus filiformis</emphasis>
|
||
</taxonomicName>
|
||
(Latzel, 1884) lectotype ♂ (NHWM MY3754)
|
||
<emphasis id="10F14F4AB7FC4ADEC5C487F373267D7C" bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
after
|
||
<bibRefCitation id="9E42FAA1622F09D4A033DECE0E679094" author="Attems, C" journalOrPublisher="Denkschriften der Akademie der Wissenschaften Wien, mathematisch-naturwissenschaftliche Klassen," pageId="0" pageNumber="1" pagination="221 - 482" publicationUrl="http://biodiversitylibrary.org/page/27925127" refId="B4" refString="Attems, C, 1898. System der Polydesmiden. I. Theil. Denkschriften der Akademie der Wissenschaften Wien, mathematisch-naturwissenschaftliche Klassen, 67: 221 - 482, http://biodiversitylibrary.org/page/27925127" title="System der Polydesmiden. I. Theil." url="http://biodiversitylibrary.org/page/27925127" volume="67" year="1898">Attems (1898)</bibRefCitation>
|
||
<emphasis id="0E877A5350557EB8B3B9524A6971F279" bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
?left gonopod,?mesal view, after
|
||
<bibRefCitation id="81D4D32E7F22E79B4A1E3F86A902FA95" DOI="https://doi.org/10.1515/9783111609645" author="Attems, C" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 577" refId="B8" refString="Attems, C, 1940. Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Periodontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich 70: 1 - 577, DOI: https://doi.org/10.1515/9783111609645" title="Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Periodontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae." url="https://doi.org/10.1515/9783111609645" volume="70" year="1940">Attems (1940)</bibRefCitation>
|
||
<emphasis id="0305E2E5FD7FAA3C6C27C05D89874478" bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
?left gonopod,?mesal view, present study
|
||
<emphasis id="133E646541B2422A12F0DA63481D8B5D" bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName id="5510C3064ECBAEFAF422CE1F1B4655CA" baseAuthorityName="Tabacaru" baseAuthorityYear="1980" class="Diplopoda" family="Trichopolydesmidae" genus="Banatodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Banatodesmus jeanneli" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="jeanneli">
|
||
<emphasis id="FAD879FE59971D075827C88989A0A78E" italics="true" pageId="0" pageNumber="1">Banatodesmus jeanneli</emphasis>
|
||
</taxonomicName>
|
||
(Tabacaru, 1980) syntype ♂, right gonopod, mesal view, after
|
||
<bibRefCitation id="E074049F9D83551EC835FBEFE6B1FC70" author="Tabacaru, I" journalOrPublisher="Travaux de l'Institut de Speologie " Emile Racovitza "" pageId="0" pageNumber="1" pagination="155 - 161" refId="B58" refString="Tabacaru, I, 1980. Trichopolydesmus (Banatodesmus) jeanneli n. sg., n. sp. Travaux de l'Institut de Speologie "Emile Racovitza" 24: 155 - 161" title="Trichopolydesmus (Banatodesmus) jeanneli n. sg., n. sp." volume="24" year="1980">Tabacaru (1980)</bibRefCitation>
|
||
<emphasis id="748117B1EEDF410CD5C0F6E35AB33F6A" bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
<taxonomicName id="2BA1FC588ADEAEA592452E749A23A5A2" authorityName="Tabacaru" authorityYear="1975" class="Diplopoda" family="Trichopolydesmidae" genus="Napocodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Napocodesmus florentzae" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="florentzae">
|
||
<emphasis id="15E77352E5F56D5EB702D76E35E23F01" italics="true" pageId="0" pageNumber="1">Napocodesmus florentzae</emphasis>
|
||
</taxonomicName>
|
||
Tabacaru, 1975 holotype ♂, right gonopod, mesal and anterior views, respectively, after
|
||
<bibRefCitation id="EA31E8DF2368A3C77D3C731228202AF8" author="Tabacaru, I" journalOrPublisher="Travaux de l'Institut de Speologie " Emile Racovitza "" pageId="0" pageNumber="1" pagination="71 - 82" refId="B57" refString="Tabacaru, I, 1975. Napocodesmus florentzae n. sp. Travaux de l'Institut de Speologie "Emile Racovitza" 14: 71 - 82" title="Napocodesmus florentzae n. sp." volume="14" year="1975">Tabacaru (1975)</bibRefCitation>
|
||
<emphasis id="FA3040770FF9F7D8CBED591D779DE46D" bold="true" pageId="0" pageNumber="1">G, H</emphasis>
|
||
<taxonomicName id="98A4148830EF7206DBB02297025CECE2" authorityName="Verhoeff" authorityYear="1898" class="Diplopoda" family="Trichopolydesmidae" genus="Trichopolydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichopolydesmus eremitis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="eremitis">
|
||
<emphasis id="3BE1C9721209F644DAA024BDFDCBF67E" italics="true" pageId="0" pageNumber="1">Trichopolydesmus eremitis</emphasis>
|
||
</taxonomicName>
|
||
Verhoeff, 1898 ♂ from
|
||
<normalizedToken id="E6EE1D166061403E8C658F26683B44BE" originalValue="Cloşani">Closani</normalizedToken>
|
||
cave, right gonopod, antero-mesal and mesal views, respectively, after
|
||
<bibRefCitation id="FD760F5F887CE60AD460D8B0DF920905" author="Ceuca, T" journalOrPublisher="Studii si Cercetări de Biologie" pageId="0" pageNumber="1" pagination="335 - 343" refId="B14" refString="Ceuca, T, 1958. Contributii la cunoasterea Diplopodelor din fauna R. P. Romine. III. Diplopode cavernicole. Studii si Cercetări de Biologie 2: 335 - 343" title="Contributii la cunoasterea Diplopodelor din fauna R. P. Romine. III. Diplopode cavernicole." volume="2" year="1958">Ceuca (1958)</bibRefCitation>
|
||
<emphasis id="7D78E311625592F6E71EF00B7E5472FA" bold="true" pageId="0" pageNumber="1">I</emphasis>
|
||
<taxonomicName id="A95DAB6F4F52FBA54C18E27107E2F72A" authorityName="gen. nov., comb. nov. ex Bacillidesmus" baseAuthorityName="Strasser" baseAuthorityYear="1962" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus bulgaricus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="bulgaricus">
|
||
<emphasis id="AFEA7AEBF538481FB261DA4CA7F4B70B" italics="true" pageId="0" pageNumber="1">Balkanodesminus bulgaricus</emphasis>
|
||
</taxonomicName>
|
||
(Strasser, 1962) gen. nov., comb. nov. ♂ from Ponora Cave (NMNHS-10813), left gonopod, mesal view
|
||
<emphasis id="A5A6F1826912959E6DE6D66D8EE001C9" bold="true" pageId="0" pageNumber="1">J</emphasis>
|
||
<taxonomicName id="77F51CFE31A604FFCCE7EFF9D3139F12" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus serbicus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="serbicus">
|
||
<emphasis id="22FA6F8F97DDBA2B70F8B18249B60AC4" italics="true" pageId="0" pageNumber="1">Balkanodesminus serbicus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. paratype ♂ (NHMW MY10263), left gonopod, mesal view
|
||
<emphasis id="37439810FF5EF2C7AE2601D8E8AD7EFE" bold="true" pageId="0" pageNumber="1">K</emphasis>
|
||
<taxonomicName id="C71631319525F5905D7FAB7E338AA1D7" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="CFBD4749C38C0E21EAD2910A89969B9D" italics="true" pageId="0" pageNumber="1">Rhodopodesmus niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. holotype ♂ (NMNHS-10823), left gonopod, mesal view
|
||
<emphasis id="111B8CBF3B4ED1FA140673CD5F16A50E" bold="true" pageId="0" pageNumber="1">L</emphasis>
|
||
<taxonomicName id="112A4CA5B9F7561C6D414F61EE5EFE24" authorityName="gen. nov., comb. nov." authorityYear="2022" baseAuthorityName="Strasser" baseAuthorityYear="1966" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatus">
|
||
<emphasis id="7C14BDAB8563360D091BFC5093713D02" italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatus</emphasis>
|
||
</taxonomicName>
|
||
(Strasser, 1966a) gen. nov., comb. nov., stat. nov. syntype ♂, right gonopod, mesal view, after
|
||
<bibRefCitation id="4135F2001CB5F4F3953C5078E9A69650" author="Strasser, K" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="1" pagination="325 - 385" refId="B54" refString="Strasser, K, 1966a. Ueber Diplopoden Bulgariens. Annales Zoologici 23 (12): 325 - 385" title="Ueber Diplopoden Bulgariens." volume="23" year="1966 a">Strasser (1966a)</bibRefCitation>
|
||
<emphasis id="A59D45C0132293EFEC970041CE7B5733" bold="true" pageId="0" pageNumber="1">M</emphasis>
|
||
<taxonomicName id="91276A347D3B940DEE6ED70C4F3F60B6" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
|
||
<emphasis id="50339A7D3D8078FDDF96ED7EABD8AAD0" italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. paratype ♂ (NHMW MY10258), right gonopod, mesal view
|
||
<emphasis id="90CEAB8F81DEDF1E6E44293D4B637522" bold="true" pageId="0" pageNumber="1">N</emphasis>
|
||
<taxonomicName id="8AE7C9A263F0D602E5845DD9B96C05E4" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus dentatoides" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="dentatoides">
|
||
<emphasis id="672EDCC15B213D6C404C627D5C56E938" italics="true" pageId="0" pageNumber="1">Balkanodesminus dentatoides</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. ♂ from Sopotska Cave (NHMW MY10267), left gonopod, mesal view. Scale bars: 0.05 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="00EF57FB20EC00D04D79202ADD1C4075" pageId="0" pageNumber="1">
|
||
All five recently collected specimens of
|
||
<taxonomicName id="CC45792A231B260BA2780127F83EDF09" authorityName="gen. nov. et" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus niveus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="niveus">
|
||
<emphasis id="CA58BD5EBF618701521830ADBD8B41DC" italics="true" pageId="0" pageNumber="1">Rhodopodesmus niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. by D.A. and B.V. were found in the middle part of the cave at two spots, and all were in rotten wood.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BF9F9CB94C098B9AAB3FA7329D68B31F" pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph id="F14534C05EA0EE7EFF3EDE270377A818" pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph id="277CB837D6BDC1428D52FFC9743A1BFD" pageId="0" pageNumber="1">
|
||
So far, known only from its type locality, the Snezhanka Cave, Bulgaria (see also under Remarks) (Fig.
|
||
<figureCitation id="611D15FE4AD7CCA2BA6A9367AD692F9C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">18</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DFCBEA7DEF04E8054C0FAEB93985E056" doi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1" start="Figure 18" startId="F18">
|
||
<paragraph id="6029FCE7733302B24C58359B501E8F29" pageId="0" pageNumber="1">
|
||
<emphasis id="4AF75F5BCFEC2F3D75C33A0C6BBFE3DB" bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
|
||
Distribution of cavernicolous
|
||
<taxonomicName id="6E9C318B207DE1A8279DCB633A434F55" authorityName="Verhoeff" authorityYear="1910" class="Diplopoda" family="Trichopolydesmidae" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
|
||
from the Carpathian-Balkan arch and the Rhodope Mountains.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="1A517B61E8B314B0C4109685899D1DFE" pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph id="755CB792A4656E947A2B2386C6A278D6" pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph id="A17047B303A97AE881AE9D170D492AA9" pageId="0" pageNumber="1">
|
||
As mentioned above, it seems that the most similar genus to
|
||
<taxonomicName id="9733E7423F35F5E4C19BC92032E04C0F" genus="Rhodopodsemus" lsidName="Rhodopodsemus" pageId="0" pageNumber="1" rank="genus">
|
||
<emphasis id="418A91FDADAC8579C325A067E0C7CAA5" italics="true" pageId="0" pageNumber="1">Rhodopodsemus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. is
|
||
<taxonomicName id="1771BE94CD212B420933D1E58935C352" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FFC29774FDE6DD14C43E73D08FD18AB" italics="true" pageId="0" pageNumber="1">Balkanodesminus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., which makes sense due to their distributions. These two genera share not only similarities in certain habitus features but also in the gonopods. Both include small-bodied species with 19 rings in adults, with sensilla basiconica on antennomere 6 partially exposed outside the pit, hypoproct with more than two long distal setae and paraprocts with more than 2+2 long setae. The conformation of the gonopods is very similar, where the acropodite is longitudinally divided into two branches, with the solenomeral branch transversely divided into long and relatively slender solenomere and well-developed distal process in both genera. Based on this, both genera differ from other European
|
||
<taxonomicName id="13DC9418437F3D043978D647B6C5A2A5" authorityName="Verhoeff" authorityYear="1910" class="Diplopoda" family="Trichopolydesmidae" kingdom="Animalia" lsidName="" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Trichopolydesmidae</taxonomicName>
|
||
. However, in
|
||
<taxonomicName id="304957A79D24402FE7D0D1AC5D9CAA20" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="24AE483E54768940851B1CA50BC87F01" italics="true" pageId="0" pageNumber="1">Rhodopodesmus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. the solenomeral branch is trifid, where in addition to solenomere and distal process, in the base of this branch there is another, proximal process, which is more or less spatulate. These two genera also differ in some details of the gonopod coxa.
|
||
<taxonomicName id="2EBF582105C34F32737666FA92DFD2BD" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="09C57419D4CD46BC45E8AD0F04482E9E" italics="true" pageId="0" pageNumber="1">Rhodopodesmus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. has a very deep gonocoel, i.e., the anterior third of the coxa is much lower than the rest it, in the form of a shield, so that the rest of the coxa has a more or less rectangular shape laterally, while in
|
||
<taxonomicName id="B86F6B4AF14F3900E42E2AE629F4C5BE" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9EFC048DFB5EB64146B7EA2C7DB86D7A" italics="true" pageId="0" pageNumber="1">Balkanodesminus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. it is semi-circular. Also, the coxa in
|
||
<taxonomicName id="8134AF0902CCCA8CF1B5969EF59438A7" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="163E6F7A254CD50239D74ACEA7433B2E" italics="true" pageId="0" pageNumber="1">Rhodopodesmus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. has circa 15 setae, while the same in species of
|
||
<taxonomicName id="832F88B2C01751FF30F1472F8523FBE8" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Balkanodesminus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Balkanodesminus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="57FA1F36C10CEBED9DC383FBFEF13134" italics="true" pageId="0" pageNumber="1">Balkanodesminus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. bears only 3.
|
||
</paragraph>
|
||
<paragraph id="4B448E165FD12CFBED520C3C72636AD4" pageId="0" pageNumber="1">
|
||
It is worth mentioning that specimens of
|
||
<taxonomicName id="E8EA446100700051049672583ED7DCDC" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Rhodopodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhodopodesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="77389799E6BF1DF55C1069C8188BF8D8" italics="true" pageId="0" pageNumber="1">Rhodopodesmus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. were found in two more caves in the Rhodopes, viz., one female in Dupkata (= Dupcheto) Cave near Rakitovo and one male and one female in Skoka Cave near Ribnovo. Unfortunately, the material from Skoka Cave (including one male) could not be relocated in NMNHS for this study, and we still do not know whether it is
|
||
<taxonomicName id="27654F7AA4ECD9F6F0BAB691B438F21E" lsidName="R. niveus" pageId="0" pageNumber="1" rank="species" species="niveus">
|
||
<emphasis id="FBB721A3DAAEAEF100E6DFD760036062" italics="true" pageId="0" pageNumber="1">R. niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. or a new species. The female from Dupkata Cave probably belongs to
|
||
<taxonomicName id="5EF48391FA6E0DFBBF1F5A9019346431" lsidName="R. niveus" pageId="0" pageNumber="1" rank="species" species="niveus">
|
||
<emphasis id="E511D2E963182C3C4316D2F09C2B0E6C" italics="true" pageId="0" pageNumber="1">R. niveus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. et sp. nov. considering its proximity to the type locality of this species. Both localities are mapped (Fig.
|
||
<figureCitation id="6ADAA6691204864770327D8406694890" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Distribution of cavernicolous Trichopolydesmidae from the Carpathian-Balkan arch and the Rhodope Mountains." figureDoi="10.3897/zookeys.1097.83916.figure18" httpUri="https://binary.pensoft.net/fig/674296" pageId="0" pageNumber="1">18</figureCitation>
|
||
, yellow squares), and already noted under
|
||
<taxonomicName id="E20561E33F4A003F403328DFE3E084B9" authorityName="Antić & Vagalinski & Stoev & Akkari" authorityYear="2022" class="Diplopoda" family="Trichopolydesmidae" genus="Bacillidesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bacillidesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="3A5F4D8838D048804AADE85FCBB49434" italics="true" pageId="0" pageNumber="1">Bacillidesmus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by
|
||
<bibRefCitation id="5DC0ACB5E3A6B9054A762D65954B7296" author="Vagalinski, B" editor="Beron, P" journalOrPublisher="4. Biodiversity of Western Rhodopes (Bulgaria and Greece) II. Pensoft and National Museum of Natural History, Sofia" pageId="0" pageNumber="1" pagination="129 - 143" refId="B62" refString="Vagalinski, B, Stoev, P, 2011. An annotated checklist of the myriapods of the Western Rhodopes Mts (Bulgaria and Greece). In: Beron, P, Ed., Biodiversity of Bulgaria. 4. Biodiversity of Western Rhodopes (Bulgaria and Greece) II. Pensoft and National Museum of Natural History, Sofia: 129 - 143" title="An annotated checklist of the myriapods of the Western Rhodopes Mts (Bulgaria and Greece)." volumeTitle="Biodiversity of Bulgaria." year="2011">Vagalinski and Stoev (2011)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |