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<mods:title id="F15A04376040B803C1C41242FF8F93A4">The cephalopods of the Boda Limestone, Late Ordovician, of Dalarna, Sweden</mods:title>
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<treatment id="AD4D9054CD796A79F06C4384FB10FBBB" ID-DOI="http://doi.org/10.5281/zenodo.3815100" ID-GBIF-Taxon="163977244" ID-Zenodo-Dep="3815100" LSID="urn:lsid:plazi:treatment:AD4D9054CD796A79F06C4384FB10FBBB" httpUri="http://treatment.plazi.org/id/AD4D9054CD796A79F06C4384FB10FBBB" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="6DFE72C9CD796A7BF06C4384FB82FB75" box="[524,1063,1197,1224]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="255B2142CD796A7BF06C4384FB82FB75" blockId="29.[524,1063,1197,1259]" box="[524,1063,1197,1224]" pageId="29" pageNumber="30">
<heading id="7E13962ECD796A7BF06C4384FB82FB75" box="[524,1063,1197,1224]" centered="true" fontSize="11" level="2" pageId="29" pageNumber="30" reason="2">
<taxonomicName id="E2E45AC1CD796A7BF06C4384FB82FB75" authority="(Eichwald, 1860)" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[524,1063,1197,1224]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD796A7BF06C4384FCF1FB7A" bold="true" box="[524,852,1197,1224]" italics="true" pageId="29" pageNumber="30">Dawsonoceras fenestratum</emphasis>
(
<bibRefCitation id="41755CB3CD796A7BF1044384FBBAFB75" author="Eichwald E. de" box="[868,1055,1197,1224]" pageId="29" pageNumber="30" refId="ref47262" refString="Eichwald E. de. 1860. Lethaea Rossica ou Paleontologie de la Russie. Schweizerbart, Stuttgart." type="book" year="1860">Eichwald, 1860</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD796A7BF0F443FBFC3AFB56" box="[660,927,1233,1259]" pageId="29" pageNumber="30" type="description">
<paragraph id="255B2142CD796A7BF0F443FBFC3AFB56" blockId="29.[524,1063,1197,1259]" box="[660,927,1233,1259]" pageId="29" pageNumber="30">
<figureCitation id="BDDF3DC7CD796A7BF0F443FBFCA6FB56" box="[660,771,1233,1259]" captionStart="Fig" captionStartId="26.[190,233,1753,1779]" captionTargetBox="[243,1344,265,1707]" captionTargetId="figure@26.[243,1344,265,1707]" captionTargetPageId="26" captionText="Fig. 13. Polished median sections of Orthocerida of the Boda Limestone. A. Dawsonoceras fenestratum (Eichwald, 1860), PMU 26706, Kallholn. B-C. Striatocycloceras isbergi sp. nov. B. PMU 26681, Kallholn. C. PMU 26661, Osmundsberget. D. Isorthoceras angelini sp. nov., PMU 26784, Kallholn. E. Nathorstoceras kallholnense sp. nov., PMU 26729, holotype, Kallholn. F. Nathorstoceras adnatum sp. nov., PMU 26733, holotype, Kallholn. G. Pleurorthoceras osmundsbergense sp. nov., NRM-PZ Mo, 190102c, Osmundsberget.H.Ordogeisonocerasfoerstei(Strand,1934)comb.nov.,PMU26763,Kallholn. I. Geisonoceras wegelini (Angelin in Angelin &amp; Lindström, 1880) comb. nov., PMU 26740, Kallholn. Scale bars: A-D, I = 1 mm; E-H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3814251" httpUri="https://zenodo.org/record/3814251/files/figure.png" pageId="29" pageNumber="30">Figs 13A</figureCitation>
,
<figureCitation id="BDDF3DC7CD796A7BF17043FBFCE6FB56" box="[784,835,1233,1259]" captionStart="Fig" captionStartId="27.[189,232,1827,1853]" captionTargetBox="[200,1393,1425,1772]" captionTargetId="graphics@27.[350,920,1435,1742]" captionTargetPageId="27" captionText="Fig. 14. Reconstructions of the details of the septal necks and siphuncular segments of Orthocerida of the Boda Limestone. Note the different septal thicknesses. A. Striatocycloceras isbergi sp. nov., PMU 26681, Kallholn. B. Dawsonoceras fenestratum (Eichwald, 1860), PMU 26706, Kallholn. C. Pleurorthoceras osmundsbergense sp. nov., NRM-PZ Mo 150102c, Osmundsberget. D. Isorthoceras angelini sp. nov., PMU 26784, Kallholn." figureDoi="http://doi.org/10.5281/zenodo.3814253" httpUri="https://zenodo.org/record/3814253/files/figure.png" pageId="29" pageNumber="30">14B</figureCitation>
,
<figureCitation id="BDDF3DC7CD796A7BF12F43FBFCCEFB56" box="[847,875,1233,1259]" captionStart="Fig" captionStartId="30.[189,232,1770,1796]" captionTargetBox="[189,1395,932,1726]" captionTargetId="figure@30.[189,1400,932,1726]" captionTargetPageId="30" captionText="Fig. 16. Dawsonoceratidae and Proteoceratidae of the Boda Limestone. A-B. Dawsonoceras fenestratum (Eichwald, 1860), Kallholn. A. PMU 26702, nearly mature specimen with adorally decreasing angle of expansion. B. PMU 26712, juvenile specimen. C. Gorbyoceras sp. A, PMU 26772, Osmundsberget. D. Goryboceras sp. B, PMU 26773, Kallholn. E. Dawsonoceras stumburi sp. nov., PMU 26717, holotype, Kallholn. F. Gorbyoceras alternestriatum (Strand, 1934) comb. nov., PMU 26771, Unskarsheden. Scale bars: A-C, E = 10 mm; D = 10 mm; F = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3814257" httpUri="https://zenodo.org/record/3814257/files/figure.png" pageId="29" pageNumber="30">16</figureCitation>
A-B
</paragraph>
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<subSubSection id="6DFE72C9CD796A7BF2DD4238FE30FA2F" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="255B2142CD796A7BF2DD4238FC74FA91" blockId="29.[189,977,1298,1324]" box="[189,977,1298,1324]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="05F4066CCD796A7BF2DD4238FC74FA91" box="[189,977,1298,1324]" pageId="29" pageNumber="30">
<taxonomicName id="E2E45AC1CD796A7BF2DD4238FC8DFA91" authority="Eichwald, 1860: 1231" authorityName="Eichwald" authorityPageNumber="1231" authorityYear="1860" box="[189,808,1298,1324]" class="Cephalopoda" family="Nautilidae" genus="Cycloceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fenestratum">
<treatmentCitation id="A4450753CD796A7BF2DD4238FD7AFA91" author="Eichwald E. de" box="[189,735,1298,1324]" page="1231" pageId="29" pageNumber="30" year="1860">
<emphasis id="1790FD50CD796A7BF2DD4238FEE4FA91" box="[189,321,1298,1324]" italics="true" pageId="29" pageNumber="30">Cycloceras</emphasis>
<treatmentCitation id="A4450753CD796A7BF3284238FD7AFA91" author="Eichwald E. de" box="[328,735,1298,1324]" page="" pageId="29" pageNumber="30" year="1860">
<emphasis id="1790FD50CD796A7BF3284238FE76FA91" box="[328,467,1298,1324]" italics="true" pageId="29" pageNumber="30">fenestratum</emphasis>
<bibRefCitation id="41755CB3CD796A7BF3BA4238FD7AFA91" author="Eichwald E. de" box="[474,735,1298,1324]" pageId="29" pageNumber="30" refId="ref47262" refString="Eichwald E. de. 1860. Lethaea Rossica ou Paleontologie de la Russie. Schweizerbart, Stuttgart." type="book" year="1860">Eichwald, 1860: 1231</bibRefCitation>
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, 1232
</taxonomicName>
, pl. 48: 14a-b.
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</paragraph>
<paragraph id="255B2142CD796A7BF2DD427FFE30FA2F" blockId="29.[189,1398,1363,1426]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="05F4066CCD796A7BF2DD427FFE30FA2F" pageId="29" pageNumber="30">
<taxonomicName id="E2E45AC1CD796A7BF2DD427FFDA5FAD3" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[189,512,1364,1390]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD796A7BF2DD427FFDA5FAD3" box="[189,512,1364,1390]" italics="true" pageId="29" pageNumber="30">Dawsonoceras fenestratum</emphasis>
</taxonomicName>
<treatmentCitation id="A4450753CD796A7BF041427EFCCFFAD3" author="Kroger B. &amp; Isakar M." box="[545,874,1363,1390]" page="156" pageId="29" pageNumber="30" year="2006">
<bibRefCitation id="41755CB3CD796A7BF041427EFCCFFAD3" author="Kroger B. &amp; Isakar M." box="[545,874,1363,1390]" pageId="29" pageNumber="30" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar 2006: 156</bibRefCitation>
</treatmentCitation>
, figs 8b, 10b, 12c, f-g (with synonymy). ―
<treatmentCitation id="A4450753CD796A7BF2DD425DFE37FA2F" author="Kroger B." box="[189,402,1399,1426]" page="597" pageId="29" pageNumber="30" year="2007">
<bibRefCitation id="41755CB3CD796A7BF2DD425DFE37FA2F" author="Kroger B." box="[189,402,1399,1426]" pageId="29" pageNumber="30" pagination="591 - 608" refId="ref48327" refString="Kroger B. 2007. Concentrations of juvenile and small adult cephalopods in the Hirnantian cherts (Late Ordovician) of Porkuni, Estonia. Acta Palaeontologica Polonica 52: 591 - 608." type="journal article" year="2007">Kröger 2007: 597</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD796A7BF2DD4292FDB5F90F" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="255B2142CD796A7BF2DD4292FE93FA6F" blockId="29.[189,310,1464,1490]" box="[189,310,1464,1490]" pageId="29" pageNumber="30">
<heading id="7E13962ECD796A7BF2DD4292FE93FA6F" bold="true" box="[189,310,1464,1490]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="1790FD50CD796A7BF2DD4292FE93FA6F" bold="true" box="[189,310,1464,1490]" pageId="29" pageNumber="30">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD796A7BF2DD42C2FDB5F90F" blockId="29.[189,1399,1511,1714]" pageId="29" pageNumber="30">
<taxonomicName id="E2E45AC1CD796A7BF2DD42C2FECEF9BC" authorityName="Hyatt" authorityYear="1884" box="[189,363,1512,1537]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD796A7BF2DD42C2FECEF9BC" box="[189,363,1512,1537]" italics="true" pageId="29" pageNumber="30">Dawsonoceras</emphasis>
</taxonomicName>
with angle of expansion of less than ten degrees in juvenile stages. Tubular adult body chamber has diameter of approximately
<quantity id="E21C8CA7CD796A7BF0F44120FD4FF998" box="[660,746,1546,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="30" unit="mm" value="25.0">25 mm</quantity>
; five to six annulations occur at distance similar to shell diameter; approximately 20 longitudinal lirae occur around shell circumference, approximately fifteen growth lines or transverse striae occur per cycle of annulations; septal necks achoanitic; siphuncle subcentral, expanded within chambers. Mural and hyposeptal cameral deposits are known. (Adopted from
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.)
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD796A7BF2DD41F3FAD2F809" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="255B2142CD796A7BF2DD41F3FE5EF94F" blockId="29.[189,507,1752,1779]" box="[189,507,1752,1779]" pageId="29" pageNumber="30">
<heading id="7E13962ECD796A7BF2DD41F3FE5EF94F" bold="true" box="[189,507,1752,1779]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="1790FD50CD796A7BF2DD41F3FE5EF94F" bold="true" box="[189,507,1752,1779]" pageId="29" pageNumber="30">
<typeStatus id="FA5F9FE0CD796A7BF2DD41F3FF5FF94E" box="[189,250,1753,1779]" pageId="29" pageNumber="30">Type</typeStatus>
locality and horizon
</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD796A7BF2DD402DFC84F89C" blockId="29.[189,801,1799,1826]" box="[189,801,1799,1826]" pageId="29" pageNumber="30">
Mõnuste (Kirna), western
<collectingCountry id="5DF361D2CD796A7BF393402DFDEBF89C" box="[499,590,1799,1825]" name="Estonia" pageId="29" pageNumber="30">Estonia</collectingCountry>
, Late Ordovician.
</paragraph>
<paragraph id="255B2142CD796A7BF2DD4062FE0FF8DF" blockId="29.[189,426,1864,1890]" box="[189,426,1864,1890]" pageId="29" pageNumber="30">
<heading id="7E13962ECD796A7BF2DD4062FE0FF8DF" bold="true" box="[189,426,1864,1890]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="1790FD50CD796A7BF2DD4062FE0FF8DF" bold="true" box="[189,426,1864,1890]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD796A7BF2DD405DFAD2F809" blockId="29.[189,1399,1911,1973]" pageId="29" pageNumber="30">
Two specimens,
<collectionCode id="43F5B987CD796A7BF3EC405DFE69F82C" box="[396,460,1911,1937]" country="Sweden" httpUri="http://grbio.org/cool/32ji-dbbk" name="Paleontological Museum of Uppsala" pageId="29" pageNumber="30">PMU</collectionCode>
26700 and 26701, Osmundsberget, Siljan District; fifteen specimens (
<collectionCode id="43F5B987CD796A7BF755405DFAD2F82C" box="[1333,1399,1911,1937]" country="Sweden" httpUri="http://grbio.org/cool/32ji-dbbk" name="Paleontological Museum of Uppsala" pageId="29" pageNumber="30">PMU</collectionCode>
2670226716), Kallholn, Siljan District; all from the Boda Limestone, Boda Core Member, late Katian.
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD7A6A78F2DD4621FC12FD0E" pageId="30" pageNumber="31" type="description">
<paragraph id="255B2142CD7A6A78F2DD4621FEEBFE98" blockId="30.[189,334,267,293]" box="[189,334,267,293]" pageId="30" pageNumber="31">
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<emphasis id="1790FD50CD7A6A78F2DD4621FEEBFE98" bold="true" box="[189,334,267,293]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD7A6A78F2DD4613FC12FD0E" blockId="30.[189,1399,313,692]" pageId="30" pageNumber="31">
The external characters were described in detail by
<bibRefCitation id="41755CB3CD7A6A78F1534610FBC5FEEE" author="Kroger B. &amp; Isakar M." box="[819,1120,313,340]" pageId="30" pageNumber="31" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger and Isakar (2006</bibRefCitation>
; see also
<figureCitation id="BDDF3DC7CD7A6A78F6B84613FA96FEE9" box="[1240,1331,313,340]" captionStart="Fig" captionStartId="30.[189,232,1770,1796]" captionTargetBox="[189,1395,932,1726]" captionTargetId="figure@30.[189,1400,932,1726]" captionTargetPageId="30" captionText="Fig. 16. Dawsonoceratidae and Proteoceratidae of the Boda Limestone. A-B. Dawsonoceras fenestratum (Eichwald, 1860), Kallholn. A. PMU 26702, nearly mature specimen with adorally decreasing angle of expansion. B. PMU 26712, juvenile specimen. C. Gorbyoceras sp. A, PMU 26772, Osmundsberget. D. Goryboceras sp. B, PMU 26773, Kallholn. E. Dawsonoceras stumburi sp. nov., PMU 26717, holotype, Kallholn. F. Gorbyoceras alternestriatum (Strand, 1934) comb. nov., PMU 26771, Unskarsheden. Scale bars: A-C, E = 10 mm; D = 10 mm; F = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3814257" httpUri="https://zenodo.org/record/3814257/files/figure.png" pageId="30" pageNumber="31">Figs 16</figureCitation>
A-B). In specimen PMU 26706 additional details of the siphuncle are preserved. Specimen PMU 26706 is a fragment of a phragmocone with a diameter of
<quantity id="E21C8CA7CD7A6A78F08046AAFCFBFE27" box="[736,862,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="30" pageNumber="31" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">1821 mm</quantity>
and a length of
<quantity id="E21C8CA7CD7A6A78F67846AAFBC8FE27" box="[1048,1133,384,410]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="30" pageNumber="31" unit="mm" value="28.0">28 mm</quantity>
. The chamber distance is relatively constant at ca.
<quantity id="E21C8CA7CD7A6A78F0674689FDF5FE03" box="[519,592,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="30" pageNumber="31" unit="mm" value="4.0">4 mm</quantity>
. The siphuncle is slightly eccentric at the adoral end and ca.
<quantity id="E21C8CA7CD7A6A78F7514689FAD2FE03" box="[1329,1399,419,446]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="mm" value="8.0">8 mm</quantity>
distant from the conch margin (SPR 0.38). The siphuncular diameter is
<quantity id="E21C8CA7CD7A6A78F64946ECFB22FE5C" box="[1065,1159,454,481]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="mm" value="1.3">1.3 mm</quantity>
at the adapical end (0.07 of the corresponding conch cross section); there the septal perforation is only
<quantity id="E21C8CA7CD7A6A78F6FE46C0FB5CFDB9" box="[1182,1273,490,516]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="mm" value="0.8">0.8 mm</quantity>
wide. The siphuncular segments are barrel-shaped; rapidly increasing in diameter close to the septal perforation, the side directed toward the conch center bends inward slightly, forming a shallow concave connecting ring segment (
<figureCitation id="BDDF3DC7CD7A6A78F3094579FE79FDD0" box="[361,476,595,621]" captionStart="Fig" captionStartId="26.[190,233,1753,1779]" captionTargetBox="[243,1344,265,1707]" captionTargetId="figure@26.[243,1344,265,1707]" captionTargetPageId="26" captionText="Fig. 13. Polished median sections of Orthocerida of the Boda Limestone. A. Dawsonoceras fenestratum (Eichwald, 1860), PMU 26706, Kallholn. B-C. Striatocycloceras isbergi sp. nov. B. PMU 26681, Kallholn. C. PMU 26661, Osmundsberget. D. Isorthoceras angelini sp. nov., PMU 26784, Kallholn. E. Nathorstoceras kallholnense sp. nov., PMU 26729, holotype, Kallholn. F. Nathorstoceras adnatum sp. nov., PMU 26733, holotype, Kallholn. G. Pleurorthoceras osmundsbergense sp. nov., NRM-PZ Mo, 190102c, Osmundsberget.H.Ordogeisonocerasfoerstei(Strand,1934)comb.nov.,PMU26763,Kallholn. I. Geisonoceras wegelini (Angelin in Angelin &amp; Lindström, 1880) comb. nov., PMU 26740, Kallholn. Scale bars: A-D, I = 1 mm; E-H = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3814251" httpUri="https://zenodo.org/record/3814251/files/figure.png" pageId="30" pageNumber="31">Figs 13A</figureCitation>
,
<figureCitation id="BDDF3DC7CD7A6A78F38B4579FDBAFDD0" box="[491,543,595,621]" captionStart="Fig" captionStartId="27.[189,232,1827,1853]" captionTargetBox="[200,1393,1425,1772]" captionTargetId="graphics@27.[350,920,1435,1742]" captionTargetPageId="27" captionText="Fig. 14. Reconstructions of the details of the septal necks and siphuncular segments of Orthocerida of the Boda Limestone. Note the different septal thicknesses. A. Striatocycloceras isbergi sp. nov., PMU 26681, Kallholn. B. Dawsonoceras fenestratum (Eichwald, 1860), PMU 26706, Kallholn. C. Pleurorthoceras osmundsbergense sp. nov., NRM-PZ Mo 150102c, Osmundsberget. D. Isorthoceras angelini sp. nov., PMU 26784, Kallholn." figureDoi="http://doi.org/10.5281/zenodo.3814253" httpUri="https://zenodo.org/record/3814253/files/figure.png" pageId="30" pageNumber="31">14B</figureCitation>
). The asymmetrical shape of the connecting ring was noted in material previously described by
<bibRefCitation id="41755CB3CD7A6A78F388455DFCABFD2C" author="Kroger B. &amp; Isakar M." box="[488,782,630,657]" pageId="30" pageNumber="31" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar (2006)</bibRefCitation>
. The septal necks are short and suborthochoanitic. Shallow hyposeptal deposits occur at the adapical five chambers.
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD7A6A79F2DD45F1FDF3FCC9" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph id="255B2142CD7A6A78F2DD45F1FE8AFD48" blockId="30.[189,303,731,757]" box="[189,303,731,757]" pageId="30" pageNumber="31">
<heading id="7E13962ECD7A6A78F2DD45F1FE8AFD48" bold="true" box="[189,303,731,757]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="1790FD50CD7A6A78F2DD45F1FE8AFD48" bold="true" box="[189,303,731,757]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD7A6A78F2DD4423FAD2FCD7" blockId="30.[189,1399,777,874]" pageId="30" pageNumber="31">
This species was revised by
<bibRefCitation id="41755CB3CD7A6A78F07B4420FC9FFC99" author="Kroger B. &amp; Isakar M." box="[539,826,777,804]" pageId="30" pageNumber="31" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar (2006)</bibRefCitation>
and its main features were described based on specimens from the Porkuni Formation of
<collectingCountry id="5DF361D2CD7A6A78F0AA4406FC83FCFB" box="[714,806,812,838]" name="Estonia" pageId="30" pageNumber="31">Estonia</collectingCountry>
. Here, additional data regarding the variation of the external characters and features of the siphuncle are noted. The occurrence of hyposeptal cameral
</paragraph>
<caption id="719B71CACD7A6A78F2DD41C0FD55F809" ID-DOI="http://doi.org/10.5281/zenodo.3814257" ID-Zenodo-Dep="3814257" httpUri="https://zenodo.org/record/3814257/files/figure.png" pageId="30" pageNumber="31" startId="30.[189,232,1770,1796]" targetBox="[189,1395,932,1726]" targetPageId="30">
<paragraph id="255B2142CD7A6A78F2DD41C0FD55F809" blockId="30.[189,1400,1770,1972]" pageId="30" pageNumber="31">
<emphasis id="1790FD50CD7A6A78F2DD41C0FEB2F8B9" bold="true" box="[189,279,1770,1796]" pageId="30" pageNumber="31">Fig. 16.</emphasis>
<taxonomicName id="E2E45AC1CD7A6A78F37D41C0FE53F8B9" authorityName="Flower" authorityYear="1962" box="[285,502,1770,1796]" class="Cephalopoda" family="Dawsonoceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="family">Dawsonoceratidae</taxonomicName>
and
<taxonomicName id="E2E45AC1CD7A6A78F04B41C0FD44F8B9" authorityName="Flower" authorityYear="1962" box="[555,737,1770,1796]" class="Cephalopoda" family="Proteoceratidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="family">Proteoceratidae</taxonomicName>
of the Boda Limestone.
<emphasis id="1790FD50CD7A6A78F19A41C0FB88F8B9" bold="true" box="[1018,1069,1770,1796]" pageId="30" pageNumber="31">A -B</emphasis>
.
<taxonomicName id="E2E45AC1CD7A6A78F65941C1FE29F895" authority="(Eichwald, 1860)" baseAuthorityName="Eichwald" baseAuthorityYear="1860" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD7A6A78F65941C1FADDF8B9" box="[1081,1400,1770,1796]" italics="true" pageId="30" pageNumber="31">Dawsonoceras fenestratum</emphasis>
(
<bibRefCitation id="41755CB3CD7A6A78F2A54027FE26F895" author="Eichwald E. de" box="[197,387,1805,1832]" pageId="30" pageNumber="31" refId="ref47262" refString="Eichwald E. de. 1860. Lethaea Rossica ou Paleontologie de la Russie. Schweizerbart, Stuttgart." type="book" year="1860">Eichwald, 1860</bibRefCitation>
)
</taxonomicName>
, Kallholn.
<emphasis id="1790FD50CD7A6A78F06D4027FD87F89A" bold="true" box="[525,546,1805,1831]" pageId="30" pageNumber="31">A</emphasis>
. PMU 26702, nearly mature specimen with adorally decreasing angle of expansion.
<emphasis id="1790FD50CD7A6A78F326401AFEFFF8F7" bold="true" box="[326,346,1840,1866]" pageId="30" pageNumber="31">B</emphasis>
. PMU 26712, juvenile specimen.
<emphasis id="1790FD50CD7A6A78F099401AFCABF8F7" bold="true" box="[761,782,1840,1866]" pageId="30" pageNumber="31">C</emphasis>
.
<taxonomicName id="E2E45AC1CD7A6A78F140401AFC1CF8F7" authorityName="Shimizu &amp; Obata" authorityYear="1935" box="[800,953,1840,1866]" class="Cephalopoda" family="Proteoceratidae" genus="Gorbyoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD7A6A78F140401AFC1CF8F7" box="[800,953,1840,1866]" italics="true" pageId="30" pageNumber="31">Gorbyoceras</emphasis>
</taxonomicName>
sp. A, PMU 26772, Osmundsberget.
<emphasis id="1790FD50CD7A6A78F2DD407EFF77F8D3" bold="true" box="[189,210,1876,1902]" pageId="30" pageNumber="31">D</emphasis>
.
<emphasis id="1790FD50CD7A6A78F2BE4079FED2F8D0" box="[222,375,1875,1901]" italics="true" pageId="30" pageNumber="31">Goryboceras</emphasis>
sp. B, PMU 26773, Kallholn.
<emphasis id="1790FD50CD7A6A78F0AD407EFD44F8D3" bold="true" box="[717,737,1876,1902]" pageId="30" pageNumber="31">E</emphasis>
.
<taxonomicName id="E2E45AC1CD7A6A78F08D407EFBADF8D0" authority="Kröger, 2013" authorityName="Kröger" authorityYear="2013" box="[749,1032,1875,1901]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="stumburi" status="sp. nov.">
<emphasis id="1790FD50CD7A6A78F08D407EFBADF8D0" box="[749,1032,1875,1901]" italics="true" pageId="30" pageNumber="31">Dawsonoceras stumburi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CA3402BCD7A6A78F66D407EFBC2F8D3" box="[1037,1127,1876,1902]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
, PMU 26717, holotype, Kallholn.
<emphasis id="1790FD50CD7A6A78F351405DFEE6F82C" bold="true" box="[305,323,1911,1937]" pageId="30" pageNumber="31">F</emphasis>
.
<taxonomicName id="E2E45AC1CD7A6A78F331405CFCF6F82C" authority="(Strand, 1934)" authorityName="Kröger" authorityYear="2013" baseAuthorityName="Strand" baseAuthorityYear="1934" box="[337,851,1910,1937]" class="Cephalopoda" family="Proteoceratidae" genus="Gorbyoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="alternestriatum" status="comb. nov.">
<emphasis id="1790FD50CD7A6A78F331405CFD00F82D" box="[337,677,1910,1936]" italics="true" pageId="30" pageNumber="31">Gorbyoceras alternestriatum</emphasis>
(
<bibRefCitation id="41755CB3CD7A6A78F0D3405CFCEEF82C" author="Strand T." box="[691,843,1910,1937]" pageId="30" pageNumber="31" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand, 1934</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="0CA3402BCD7A6A78F13A405CFC78F82C" box="[858,989,1910,1937]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
, PMU 26771, Unskarsheden. Scale bars: A-C, E = 10 mm; D = 10 mm; F = 10 mm.
</paragraph>
</caption>
<paragraph id="255B2142CD7B6A79F2DD4627FD47FEF7" blockId="31.[189,1398,269,331]" pageId="31" pageNumber="32">
deposits in specimens of
<taxonomicName id="E2E45AC1CD7B6A79F38B4624FBA6FE95" authority="(Eichwald, 1860)" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[491,1027,269,296]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD7B6A79F38B4624FD3CFE9A" box="[491,665,270,295]" italics="true" pageId="31" pageNumber="32">Dawsonoceras</emphasis>
<emphasis id="1790FD50CD7B6A79F0C24627FC88FE9A" box="[674,813,269,295]" italics="true" pageId="31" pageNumber="32">fenestratum</emphasis>
(
<bibRefCitation id="41755CB3CD7B6A79F15E4627FC5EFE95" author="Eichwald E. de" box="[830,1019,269,296]" pageId="31" pageNumber="32" refId="ref47262" refString="Eichwald E. de. 1860. Lethaea Rossica ou Paleontologie de la Russie. Schweizerbart, Stuttgart." type="book" year="1860">Eichwald, 1860</bibRefCitation>
)
</taxonomicName>
from the Boda Limestone (see below) is included within the diagnosis herein.
</paragraph>
<paragraph id="255B2142CD7B6A79F2DD465BFE43FD3F" blockId="31.[189,1399,369,642]" pageId="31" pageNumber="32">
The conch measurements reveal a clear tendency of a decrease in angle of expansion with increasing conch diameter (
<bibRefCitation id="41755CB3CD7B6A79F3E146BFFD2EFE12" author="Kroger B. &amp; Isakar M." box="[385,651,404,431]" pageId="31" pageNumber="32" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar 2006</bibRefCitation>
). The mature body chamber of
<taxonomicName id="E2E45AC1CD7B6A79F19B46BFFB0FFE12" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[1019,1194,405,431]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD7B6A79F19B46BFFBB5FE13" box="[1019,1040,405,430]" italics="true" pageId="31" pageNumber="32">D</emphasis>
.
<emphasis id="1790FD50CD7B6A79F67F46BFFB0FFE12" box="[1055,1194,405,431]" italics="true" pageId="31" pageNumber="32">fenestratum</emphasis>
</taxonomicName>
is nearly tubular, whereas in juvenile conchs the maximum expansion reaches 10° (PMU 26708 with cross section diameters of
<quantity id="E21C8CA7CD7B6A79F33B46F1FE74FE48" box="[347,465,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.4" metricValueMin="0.9" pageId="31" pageNumber="32" unit="mm" value="11.5" valueMax="14.0" valueMin="9.0">914 mm</quantity>
; compare
<figureCitation id="BDDF3DC7CD7B6A79F02F46F1FD01FE4B" box="[591,676,475,502]" captionStart="Fig" captionStartId="30.[189,232,1770,1796]" captionTargetBox="[189,1395,932,1726]" captionTargetId="figure@30.[189,1400,932,1726]" captionTargetPageId="30" captionText="Fig. 16. Dawsonoceratidae and Proteoceratidae of the Boda Limestone. A-B. Dawsonoceras fenestratum (Eichwald, 1860), Kallholn. A. PMU 26702, nearly mature specimen with adorally decreasing angle of expansion. B. PMU 26712, juvenile specimen. C. Gorbyoceras sp. A, PMU 26772, Osmundsberget. D. Goryboceras sp. B, PMU 26773, Kallholn. E. Dawsonoceras stumburi sp. nov., PMU 26717, holotype, Kallholn. F. Gorbyoceras alternestriatum (Strand, 1934) comb. nov., PMU 26771, Unskarsheden. Scale bars: A-C, E = 10 mm; D = 10 mm; F = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3814257" httpUri="https://zenodo.org/record/3814257/files/figure.png" pageId="31" pageNumber="32">Fig. 16</figureCitation>
A-B). This is in agreement with the measurements listed by
<bibRefCitation id="41755CB3CD7B6A79F2DD46D5FE79FDA4" author="Kroger B. &amp; Isakar M." box="[189,476,510,537]" pageId="31" pageNumber="32" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar (2006)</bibRefCitation>
. Additionally, the relative distance of the transverse annuli is highly variable, but lacks a clear ontogenetic trend and varies between 0.16 and 0.3 of the corresponding conch diameter (mean 0.25, n=12). The number of longitudinal lirae is fixed in each specimen, but varies among different specimens from 20 to 31.
</paragraph>
<paragraph id="255B2142CD7B6A79F2DD4583FDF3FCC9" blockId="31.[189,1399,681,884]" pageId="31" pageNumber="32">
These external features are well within the range of variability of the Trenton dawsonoceratid
<taxonomicName id="E2E45AC1CD7B6A79F6914583FD8FFD5B" authority="Whiteaves, 1898" authorityName="Whiteaves" authorityYear="1898" class="Liliopsida" family="Orchidaceae" genus="Orthoceras" kingdom="Plantae" order="Asparagales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="species" species="beauportense">
<emphasis id="1790FD50CD7B6A79F6914583FAD2FD7E" box="[1265,1399,681,707]" italics="true" pageId="31" pageNumber="32">Orthoceras</emphasis>
<emphasis id="1790FD50CD7B6A79F2DD45E6FEFEFD5B" box="[189,347,716,742]" italics="true" pageId="31" pageNumber="32">beauportense</emphasis>
Whiteaves, 1898
</taxonomicName>
, from which the internal characters are unknown. It is probable that this species should be synonymized with
<taxonomicName id="E2E45AC1CD7B6A79F00B45DAFCBDFCB7" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[619,792,752,778]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD7B6A79F00B45DAFD25FCB4" box="[619,640,752,777]" italics="true" pageId="31" pageNumber="32">D</emphasis>
.
<emphasis id="1790FD50CD7B6A79F0ED45DAFCBDFCB7" box="[653,792,752,778]" italics="true" pageId="31" pageNumber="32">fenestratum</emphasis>
</taxonomicName>
. Chen &amp;
<bibRefCitation id="41755CB3CD7B6A79F1E645DAFBACFCB7" author="Zou X-P." box="[902,1033,752,778]" pageId="31" pageNumber="32" pagination="33 - 111" refId="ref46872" refString="Chen, J-Y. &amp; Zou X-P. 1984. Ordovician cephalopods from the Ordos area, China. Memoirs of the Nanjing Institute of Geology and Palaeontology 20: 33 - 111." type="journal article" year="1984">Zou (1984)</bibRefCitation>
reported a similar species from the late Katian Sheshan Formation of North
<collectingCountry id="5DF361D2CD7B6A79F0AE4439FCB2FC90" box="[718,791,787,813]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
, which they assign to
<taxonomicName id="E2E45AC1CD7B6A79F6414439FE30FCED" authority="(Whiteaves, 1898)" baseAuthorityName="Whiteaves" baseAuthorityYear="1898" class="Cephalopoda" family="Nautilidae" genus="Anaspyroceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="beauportense">
<emphasis id="1790FD50CD7B6A79F6414439FB75FC90" box="[1057,1232,787,813]" italics="true" pageId="31" pageNumber="32">Anaspyroceras</emphasis>
<emphasis id="1790FD50CD7B6A79F6B84439FAD3FC90" box="[1240,1398,787,813]" italics="true" pageId="31" pageNumber="32">beauportense</emphasis>
(Whiteaves, 1898)
</taxonomicName>
but provided no details regarding internal characters. This specimen probably should also be assigned to
<taxonomicName id="E2E45AC1CD7B6A79F3C14470FDF5FCCE" baseAuthorityName="Eichwald" baseAuthorityYear="1860" box="[417,592,857,884]" class="Cephalopoda" family="Dawsonoceratidae" genus="Dawsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="fenestratum">
<emphasis id="1790FD50CD7B6A79F3C14470FE13FCCE" box="[417,438,858,883]" italics="true" pageId="31" pageNumber="32">D</emphasis>
.
<emphasis id="1790FD50CD7B6A79F3A54473FDF5FCCE" box="[453,592,857,883]" italics="true" pageId="31" pageNumber="32">fenestratum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD7B6A79F2DD44B0FB10FBBB" pageId="31" pageNumber="32" type="multiple">
<paragraph id="255B2142CD7B6A79F2DD44B0FDD8FC08" blockId="31.[189,637,922,949]" box="[189,637,922,949]" pageId="31" pageNumber="32">
<heading id="7E13962ECD7B6A79F2DD44B0FDD8FC08" bold="true" box="[189,637,922,949]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="1790FD50CD7B6A79F2DD44B0FDD8FC08" bold="true" box="[189,637,922,949]" pageId="31" pageNumber="32">Stratigraphic and geographic range</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD7B6A79F2DD44E3FB10FBBB" blockId="31.[189,1398,969,1030]" pageId="31" pageNumber="32">
Nabala/Vormsi Regional Stages,
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(questionable, see
<bibRefCitation id="41755CB3CD7B6A79F11744E0FBD8FC59" author="Kroger B. &amp; Isakar M." box="[887,1149,969,996]" pageId="31" pageNumber="32" pagination="139 - 165" refId="ref48361" refString="Kroger B. &amp; Isakar M. 2006. Revision of annulated orthoceridan cephalopods of the Baltoscandic Ordovician. Fossil Record 9: 139 - 165. http: // dx. doi. org / 10.1002 / mmng. 200600005" type="journal article" year="2006">Kröger &amp; Isakar 2006</bibRefCitation>
); Porkuni Formation,
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; Boda Limestone, Siljan District,
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; late Katian-Hirnantian, Ordovician.
</paragraph>
</subSubSection>
</treatment>
</document>