369 lines
47 KiB
XML
369 lines
47 KiB
XML
<document id="F0E26A095D04703B8E5D177D633BC1FD" ID-DOI="10.5281/zenodo.197810" ID-GBIF-Dataset="3fed616e-ea51-496b-b522-145b87775b5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="197810" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460279155734" checkinUser="plazi" docAuthor="Rahmadi, Cahyo, Harvey, Mark S. & Kojima, Jun-Ichi" docDate="2010" docId="AD1687F3FF8BFF8D0CEB4B802B1FFD33" docLanguage="en" docName="zt02612p021.pdf" docOrigin="Zootaxa 2612" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sarax mardua Rahmadi, Harvey & Kojima, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="18" masterDocId="512FFF8BFF87FF9C0C7C4D7D2E59FFA4" masterDocTitle="Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island" masterLastPageNumber="21" masterPageNumber="1" pageNumber="13" updateTime="1698249151004" updateUser="plazi">
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<mods:mods id="51100C0E35EDFA032A8BB503B3759976" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C4CA872AC9F12F453FF4897FF7585B3B">
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<mods:title id="09F73F493B9C9D2DFE565283B5FE9BA3">Whip spiders of the genus Sarax Simon 1892 (Amblypygi: Charinidae) from Borneo Island</mods:title>
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</mods:titleInfo>
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<mods:name id="83245E2C581E8CB527137A933D1FDE89" type="personal">
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<mods:role id="0CFDAC527CBD7551B8A14D5AE8A208DB">
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<mods:roleTerm id="2FBAB6E260F4B647C7535EFC2348792B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E08BCDCA27D554FC9F2210A3C3AEC9CB">Rahmadi, Cahyo</mods:namePart>
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</mods:name>
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<mods:name id="B064F53F06AC28F3478770CCC8324D39" type="personal">
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<mods:role id="D6C83E58E2019043E78151F67B8CEBD7">
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<mods:roleTerm id="005E1A747D6381BE556B56A8BB1D2831">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2A7925913DAC9421532EDE0C6F3EB55D">Harvey, Mark S.</mods:namePart>
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</mods:name>
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<mods:name id="1DEA93DC08A65BE994EB307F45203926" type="personal">
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<mods:role id="4CACE71A803353BEB54E07DA61038A17">
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<mods:roleTerm id="EF50D0A3B2745D86A7F693530EC8F798">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="40402D1D5B7FCC900FC3633B6E38F0A6">Kojima, Jun-Ichi</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9C2AD73500AED3701BEE3CFDB774ABED">text</mods:typeOfResource>
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<mods:titleInfo id="180653722E027DACC9A588FD335E4A8F">
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<mods:title id="2E04151A30031473165D3B1409A20D61">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="DB60D6A2AB426D2674F3EF45055D92AF">
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<mods:date id="044A7B722D80CBE13C07C8C3A9D09504">2010</mods:date>
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<mods:detail id="07E94235EE3613D7B8DB160FA95DA1A9" type="volume">
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<mods:number id="CBF092B2BB2B45CBF89E40588389D475">2612</mods:number>
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</mods:detail>
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<mods:extent id="752A9639D2DFE8518D4B3FB4C4806C12" unit="page">
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<mods:start id="8FADD23081FDD8B5607772E3A3E381B0">1</mods:start>
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<mods:end id="62749CA4142F3DA7F3681D88EFFECD62">21</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="0B8AAADF8C159713214D7C10098C86B9">journal article</mods:classification>
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<mods:identifier id="CB242A59A13130C2957450B933B81F48" type="DOI">10.5281/zenodo.197810</mods:identifier>
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<mods:identifier id="9530CC9DB6018FACACD10964A311CB24" type="GBIF-Dataset">3fed616e-ea51-496b-b522-145b87775b5a</mods:identifier>
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<mods:identifier id="9A47EEEED3ADCC15674322318578899E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="FA128EBC3BF32CE1624C3507B462A6B7" type="Zenodo-Dep">197810</mods:identifier>
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</mods:mods>
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<treatment id="AD1687F3FF8BFF8D0CEB4B802B1FFD33" ID-DOI="http://doi.org/10.5281/zenodo.6207617" ID-GBIF-Taxon="119396510" ID-Zenodo-Dep="6207617" LSID="urn:lsid:zoobank.org:act:0FE66A57-5AE1-4442-9F8C-E30743F28E3D" httpUri="http://treatment.plazi.org/id/AD1687F3FF8BFF8D0CEB4B802B1FFD33" lastPageId="17" lastPageNumber="18" pageId="12" pageNumber="13">
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<subSubSection id="6DA5656EFF8BFF900CEB4B802FD8F89E" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="250036E5FF8BFF900CEB4B802FF0F8B3" blockId="12.[151,425,1789,1850]" box="[151,425,1789,1815]" pageId="12" pageNumber="13">
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||
<heading id="7E488189FF8BFF900CEB4B802FF0F8B3" bold="true" box="[151,425,1789,1815]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
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||
<emphasis id="17CBEAF7FF8BFF900CEB4B802FF0F8B3" bold="true" box="[151,425,1789,1815]" pageId="12" pageNumber="13">
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<taxonomicName id="E2BF4D66FF8BFF900CEB4B802F1DF8B3" ID-CoL="79MF3" box="[151,324,1789,1815]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="mardua" status="sp. nov.">
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<emphasis id="17CBEAF7FF8BFF900CEB4B802F1DF8B3" bold="true" box="[151,324,1789,1815]" italics="true" pageId="12" pageNumber="13">Sarax mardua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0CF8578CFF8BFF900D304B802FF0F8B3" box="[332,425,1789,1815]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="250036E5FF8BFF900CEB4A5D2FD8F89E" blockId="12.[151,425,1789,1850]" box="[151,385,1824,1850]" pageId="12" pageNumber="13">
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(
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<figureCitation id="BD842A60FF8BFF900CE34A5D2F7BF89E" box="[159,290,1824,1850]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="12" pageNumber="13">Fig. 26–32</figureCitation>
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,
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<figureCitation id="BD842A60FF8BFF900D524A5D2F20F89E" box="[302,377,1824,1850]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="12" pageNumber="13">39–40</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF8BFF900CEB4A172A24F850" pageId="12" pageNumber="13" type="materials_examined">
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<paragraph id="250036E5FF8BFF900CEB4A172A24F850" blockId="12.[151,1437,1898,2036]" pageId="12" pageNumber="13">
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<emphasis id="17CBEAF7FF8BFF900CEB4A172FF8F820" bold="true" box="[151,417,1898,1924]" pageId="12" pageNumber="13">Material examined:</emphasis>
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Male
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<typeStatus id="FA048847FF8BFF900D8F4A172C3BF820" box="[499,610,1898,1924]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="43AEAE20FF8BFF900E084A172CE7F820" box="[628,702,1898,1924]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="12" pageNumber="13">MZB</collectionCode>
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.Ambl.151),
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<collectingCountry id="5DA87675FF8BFF900F274A172A50F820" box="[859,1033,1898,1924]" name="Indonesia" pageId="12" pageNumber="13">INDONESIA</collectingCountry>
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:
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<emphasis id="17CBEAF7FF8BFF90086A4A162AAEF820" box="[1046,1271,1899,1924]" italics="true" pageId="12" pageNumber="13">East Kalimantan:</emphasis>
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Kutai Timur Regency, Sangkulirang District, Pengadan Village, Gua
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<taxonomicName id="E2BF4D66FF8BFF900F074AED2DBAF80E" box="[891,995,1936,1962]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="12" pageNumber="19" phylum="Arthropoda" rank="species" species="mardua">Mardua</taxonomicName>
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, near Pengadan, GPS coordinates
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<geoCoordinate id="408B5022FF8BFF900CEB4AC82F62F86B" box="[151,315,1973,1999]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="1" value="1.2319806">1°13'55.13"N</geoCoordinate>
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,
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<geoCoordinate id="408B5022FF8BFF900D344AC82C5AF86B" box="[328,515,1973,1999]" direction="east" orientation="longitude" pageId="12" pageNumber="13" precision="1" value="117.73987">117°44'23.53"E</geoCoordinate>
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(approximate),
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<date id="51011025FF8BFF900EBF4AC82DDDF86B" box="[707,900,1973,1999]" pageId="12" pageNumber="13" value="2004-08-19">19 August 2004</date>
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, L. Deharveng & A. Bedos col., (KAL-119).
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<typeStatus id="FA048847FF8BFF900CEB4AA72F57F850" box="[151,270,2010,2036]" pageId="12" pageNumber="13" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="33B9FD6CFF8BFF900D644AA72F27F850" box="[280,382,2010,2036]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
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and
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<specimenCount id="33B9FD6CFF8BFF900DCB4AA72C6EF850" box="[439,567,2010,2036]" pageId="12" pageNumber="13" type="juvenile">2 juveniles</specimenCount>
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(MNHN-Am.12), same locality data as
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<typeStatus id="FA048847FF8BFF9008724AA72A20F850" box="[1038,1145,2010,2036]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA5656EFF8AFF910CB94DEA2F20FEE3" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="250036E5FF8AFF910CB94DEA2F20FEE3" blockId="13.[151,1437,151,327]" pageId="13" pageNumber="14">
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<emphasis id="17CBEAF7FF8AFF910CB94DEA2F0BFF15" bold="true" box="[197,338,151,177]" pageId="13" pageNumber="14">Diagnosis:</emphasis>
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<taxonomicName id="E2BF4D66FF8AFF910D264DE52C54FF15" box="[346,525,152,177]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mardua">
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<emphasis id="17CBEAF7FF8AFF910D264DE52C54FF15" box="[346,525,152,177]" italics="true" pageId="13" pageNumber="14">Sarax mardua</emphasis>
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</taxonomicName>
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has a large body-size (adult body length
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<quantity id="E2479B00FF8AFF91085D4DE52A97FF16" box="[1057,1230,151,178]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="10.1" metricValueMin="8.5" pageId="13" pageNumber="14" unit="mm" value="9.3" valueMax="10.1" valueMin="8.5">
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8.5
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<emphasis id="17CBEAF7FF8AFF9108344DEA2A0EFF15" bold="true" box="[1096,1111,151,177]" italics="true" pageId="13" pageNumber="14">–</emphasis>
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10.1 mm
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</quantity>
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), distinct sexual dimorphism, with the pedipalp shorter in the female than the male, and color in alcohol pale yellow. The median eye tubercle is low and small, and is completely divided into two parts each with an eye, legs elongate, trichobothrium
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<emphasis id="17CBEAF7FF8AFF910D324C752F3CFE85" box="[334,357,264,289]" italics="true" pageId="13" pageNumber="14">bt</emphasis>
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close to proximal margin of fourth basitibial segment, and trichobothrium
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<emphasis id="17CBEAF7FF8AFF9108AE4C752AB7FE85" box="[1234,1262,264,289]" italics="true" pageId="13" pageNumber="14">bc</emphasis>
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about midway between
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<emphasis id="17CBEAF7FF8AFF910D7C4C502F4EFEE2" box="[256,279,301,326]" italics="true" pageId="13" pageNumber="14">bf</emphasis>
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and
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<emphasis id="17CBEAF7FF8AFF910D334C502F28FEE2" box="[335,369,301,326]" italics="true" pageId="13" pageNumber="14">sbf</emphasis>
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.
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</paragraph>
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</subSubSection>
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<caption id="71C0666DFF8AFF910CEB4A172D28F84F" httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="13" pageNumber="14" targetBox="[191,1379,393,1845]" targetPageId="13">
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<paragraph id="250036E5FF8AFF910CEB4A172D28F84F" blockId="13.[151,1437,1898,2027]" pageId="13" pageNumber="14">
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<emphasis id="17CBEAF7FF8AFF910CEB4A172F04F826" bold="true" box="[151,349,1898,1922]" pageId="13" pageNumber="14">FIGURES 26–32</emphasis>
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.
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<taxonomicName id="E2BF4D66FF8AFF910D104A172C50F825" box="[364,521,1898,1921]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mardua" status="sp. nov.">
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<emphasis id="17CBEAF7FF8AFF910D104A172C50F825" box="[364,521,1898,1921]" italics="true" pageId="13" pageNumber="14">Sarax mardua</emphasis>
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</taxonomicName>
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<emphasis id="17CBEAF7FF8AFF910E6D4A172C32F826" bold="true" box="[529,619,1898,1922]" pageId="13" pageNumber="14">
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<taxonomicNameLabel id="0CF8578CFF8AFF910E6D4A172C32F826" box="[529,619,1898,1922]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am.12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am.12).
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</paragraph>
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</caption>
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<subSubSection id="6DA5656EFF89FF8D0CB94DEA2AB1FE86" lastPageId="17" lastPageNumber="18" pageId="14" pageNumber="15" type="description">
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<paragraph id="250036E5FF89FF920CB94DEA2CDBFF58" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
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<emphasis id="17CBEAF7FF89FF920CB94DEA2F31FF15" bold="true" box="[197,360,151,177]" pageId="14" pageNumber="15">Description:</emphasis>
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<emphasis id="17CBEAF7FF89FF920D0E4DE52FE2FF15" box="[370,443,152,177]" italics="true" pageId="14" pageNumber="15">Male:</emphasis>
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Color in alcohol: Carapace yellowish pale, centrally with brown marks, flange yellow; pedipalp yellowish-brown except for tarsus brown and spine yellow; legs yellowish-brown, but patella dark brown; abdomen pale yellow.
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</paragraph>
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<paragraph id="250036E5FF89FF920CB94C752B66FE13" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
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Carapace (
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<figureCitation id="BD842A60FF89FF920D344C752FFCFE86" box="[328,421,264,290]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 26</figureCitation>
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): Width about 1.4
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<emphasis id="17CBEAF7FF89FF920EFD4C7A2CC9FE85" bold="true" box="[641,656,263,289]" italics="true" pageId="14" pageNumber="15">–</emphasis>
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1.7 times its length; surface finely granulate, without setiferous tubercles; median sulcus deep in posterior one-third of carapace; 4 pairs of lateral sulci present; flange wide and bent upward; anterior margin rounded, with 5 frontal setae, anterior process visible from above. Median eye tubercle low and small, clearly divided into 2 parts each including eye, without apical setae; median eyes facing antero-laterally; lateral eyes close to lateral margin of carapace, small, with yellowish tapetum.
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</paragraph>
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<paragraph id="250036E5FF89FF920CB94CBF2E95FDD6" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
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Chelicera (
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<figureCitation id="BD842A60FF89FF920D394CBF2FC7FE78" box="[325,414,450,476]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 30</figureCitation>
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): Dorsal surface smooth, with 2 fine frontal setae and several fine setae. Basal segment with 4 teeth: lower-most tooth largest, upper-most tooth bicuspid, with upper cusp larger than lower cusp; inner surface with several setae arranged in vertical row; outer surface with small tooth opposite bicuspid tooth, ventrally with several setae near proximal margin. Movable article with 5 small teeth almost equal in size.
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</paragraph>
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<paragraph id="250036E5FF89FF920CB94F002D67FD46" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
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Sternum (
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<figureCitation id="BD842A60FF89FF920D414F002FC3FD33" box="[317,410,637,663]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 27</figureCitation>
|
||
): First sternite (= tritosternum) elongate, with paired apical and several other setae. Second and third sternites rounded and slightly doomed, with 4 and 1 seta, respectively, in addition to paired apical setae. Fourth sternite (= metasternum) with 4 setae.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF89FF920CB94F902F16FAB6" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
|
||
Pedipalp (
|
||
<figureCitation id="BD842A60FF89FF920D414F902FCEFCA3" box="[317,407,749,775]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 28</figureCitation>
|
||
): Strong and slender. Trochanter: antero-dorsal margin with 4 setiferous tubercles, and median spine; antero-ventral margin with ventro-anterior apophysis present on distal margin equipped with several setiferous tubercles. Femur: antero-dorsal margin with 4 major spines, 1 minor spine, several setiferous tubercles and small tubercles, length of spines:
|
||
<emphasis id="17CBEAF7FF89FF920F404E202DA9FCD3" bold="true" box="[828,1008,861,887]" pageId="14" pageNumber="15">F1>F2>F3>F4</emphasis>
|
||
; antero-ventral margin with 4 major spines
|
||
<emphasis id="17CBEAF7FF89FF920C974EFF2EACFC38" bold="true" box="[235,245,898,924]" pageId="14" pageNumber="15">(</emphasis>
|
||
length
|
||
<emphasis id="17CBEAF7FF89FF920D354EFF2C61FC38" bold="true" box="[329,568,898,924]" pageId="14" pageNumber="15">FI>FII>FIII>FIV)</emphasis>
|
||
, 1 minor spine, several setiferous tubercle and small tubercles; 1 spine present dorsally of
|
||
<emphasis id="17CBEAF7FF89FF920DFE4EDA2FF9FC65" bold="true" box="[386,416,935,961]" pageId="14" pageNumber="15">FI</emphasis>
|
||
and as long as half of
|
||
<emphasis id="17CBEAF7FF89FF920ECB4EDA2C8CFC65" bold="true" box="[695,725,935,961]" pageId="14" pageNumber="15">FI</emphasis>
|
||
. Patella: antero-dorsal margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF89FF920CEB4EB02F3EFC43" bold="true" box="[151,359,973,999]" pageId="14" pageNumber="15">P1>P2>P3>P4>)</emphasis>
|
||
, distributed in distal half length of patella, 1 minor spine, several setiferous tubercles and small tubercles, 1 minor spine as long as 1/3 of
|
||
<emphasis id="17CBEAF7FF89FF920EBE4E8F2CBAFBA8" bold="true" box="[706,739,1010,1036]" pageId="14" pageNumber="15">P1</emphasis>
|
||
length between
|
||
<emphasis id="17CBEAF7FF89FF920FDF4E8F2D9AFBA8" bold="true" box="[931,963,1010,1036]" pageId="14" pageNumber="15">P1</emphasis>
|
||
and distal margin of patella and, 1 spine between
|
||
<emphasis id="17CBEAF7FF89FF920D7E496A2F7BFB95" bold="true" box="[258,290,1047,1073]" pageId="14" pageNumber="15">P4</emphasis>
|
||
and proximal margin; antero-ventral margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF89FF9208ED496A2B61FB95" bold="true" box="[1169,1336,1047,1073]" pageId="14" pageNumber="15">PI>PII>PIII)</emphasis>
|
||
, several setiferous tubercles and small tubercles. Tibia: antero-dorsal margin with outer surface smooth and with 2 major spines, proximal spine about half as long as distal spine; antero-ventral margin with 1 major spine close to distal margin of tibia. Tarsus completely divided (claw clearly demarcated by articulation), with 2 spines on antero-dorsal margin: proximal spine short about 1/3 as long as distal spine, the spines separated each other by about two times basal diameter of proximal spine; cleaning organ ventrally with about 30 modified hairs; apotele present.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF89FF920CB948602CF5F99D" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
|
||
Legs (
|
||
<figureCitation id="BD842A60FF89FF920D7348602F30FA93" box="[271,361,1309,1335]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 29</figureCitation>
|
||
): Femora of legs I
|
||
<emphasis id="17CBEAF7FF89FF920E3848602C0AFA93" bold="true" box="[580,595,1309,1335]" italics="true" pageId="14" pageNumber="15">–</emphasis>
|
||
IV with small tubercles bearing setae. Tibia and tarsus of leg I with 23 and 41 segments, respectively; tibiae of legs
|
||
<date id="51011025FF89FF920ED3483F2D6BFAF8" box="[687,818,1346,1372]" pageId="14" pageNumber="15">II and III 2</date>
|
||
-segmented; basitibia of leg
|
||
<date id="51011025FF89FF9208F9483F2AE5FAF8" box="[1157,1212,1346,1372]" pageId="14" pageNumber="15">IV 4</date>
|
||
-segmented, fourth segment with 1 trichobothrium (value in parentheses: ratio of the distance from the trichobothrium to the proximal margin of the segment against the length of the segment),
|
||
<emphasis id="17CBEAF7FF89FF920FB648F12DBBFA01" box="[970,994,1420,1445]" italics="true" pageId="14" pageNumber="15">bt</emphasis>
|
||
(0.40); distitibiae of legs II
|
||
<emphasis id="17CBEAF7FF89FF92095348F62B67FA01" bold="true" box="[1327,1342,1419,1445]" italics="true" pageId="14" pageNumber="15">–</emphasis>
|
||
IV each with 18 trichobothria
|
||
<emphasis id="17CBEAF7FF89FF920DEB48CD2FF4FA6D" box="[407,429,1456,1481]" italics="true" pageId="14" pageNumber="15">bf</emphasis>
|
||
(0.06),
|
||
<emphasis id="17CBEAF7FF89FF920E7748CD2C74FA6D" box="[523,557,1456,1481]" italics="true" pageId="14" pageNumber="15">sbf</emphasis>
|
||
(0.33),
|
||
<emphasis id="17CBEAF7FF89FF920EF748CD2CFEFA6D" box="[651,679,1456,1481]" italics="true" pageId="14" pageNumber="15">bc</emphasis>
|
||
(0.20),
|
||
<emphasis id="17CBEAF7FF89FF920F7848CD2D42FA6D" box="[772,795,1456,1481]" italics="true" pageId="14" pageNumber="15">bt</emphasis>
|
||
close to proximal margin of basitibial fourth segment,
|
||
<emphasis id="17CBEAF7FF89FF920CEB48A82EEAFA4A" box="[151,179,1493,1518]" italics="true" pageId="14" pageNumber="15">bc</emphasis>
|
||
about midway between
|
||
<emphasis id="17CBEAF7FF89FF920DA748A82FA7FA4A" box="[475,510,1493,1518]" italics="true" pageId="14" pageNumber="15">sbf</emphasis>
|
||
and
|
||
<emphasis id="17CBEAF7FF89FF920E4648A82C08FA4A" box="[570,593,1493,1518]" italics="true" pageId="14" pageNumber="15">bf</emphasis>
|
||
(
|
||
<figureCitation id="BD842A60FF89FF920E1F48A82CE7FA4B" box="[611,702,1493,1519]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 29</figureCitation>
|
||
). Tarsi of legs
|
||
<date id="51011025FF89FF920F0E48A82D95FA4B" box="[882,972,1493,1519]" pageId="14" pageNumber="15">
|
||
II
|
||
<emphasis id="17CBEAF7FF89FF920FFA48A82DCCFA4B" bold="true" box="[902,917,1493,1519]" italics="true" pageId="14" pageNumber="15">–</emphasis>
|
||
IV 4
|
||
</date>
|
||
-segmented; first segment greater than length of subsequent three segments combined; second segment with light-yellow transverse line; fourth segment without oblique slit; pulvilli present.
|
||
</paragraph>
|
||
<paragraph id="250036E5FF89FF920CB94B3E2FF2F968" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
|
||
Genitalia: Covered ventrally by genital operculum, proximal margin slightly concave; lateral margin yellowish-brown and median part white, medial lobes larger than lateral lobes separated with brown marks on basal lateral lobes (
|
||
<figureCitation id="BD842A60FF89FF920D0B4BF02F89F903" box="[375,464,1677,1703]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="14" pageNumber="15">Fig. 40</figureCitation>
|
||
). In dorsal view, anterior margin of gonopods rounded with brown bands extending to mid-length (
|
||
<figureCitation id="BD842A60FF89FF920D3A4BCF2FC4F968" box="[326,413,1714,1740]" captionStart="FIGURES 33 – 40" captionStartId="15.[151,269,1903,1927]" captionTargetBox="[282,1316,214,1873]" captionTargetId="figure@15.[269,1332,194,1895]" captionTargetPageId="15" captionText="FIGURES 33 – 40. Male genitalia of Sarax species from Borneo: 33. S. cavernicola, dorsal view; 34. same, ventral view; 35. S. yayukae from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37. S. sangkulirangensis new species, dorsal view; 38. ventral view; 39. S. mardua from Gua Mardua, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="14" pageNumber="15">Fig. 39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF89FF920CB94BAA2DBEF856" blockId="14.[151,1437,151,2034]" pageId="14" pageNumber="15">
|
||
<emphasis id="17CBEAF7FF89FF920CB94BAA2F70F954" box="[197,297,1751,1776]" italics="true" pageId="14" pageNumber="15">Female:</emphasis>
|
||
Distinct sexual dimorphism. Pedipalps short and stout (
|
||
<figureCitation id="BD842A60FF89FF920FB04BAA2A7CF955" box="[972,1061,1751,1777]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 31</figureCitation>
|
||
), shorter than male. Pedipalpal femur on antero-dorsal margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF89FF920F244B862DBFF8B1" bold="true" box="[856,998,1787,1813]" pageId="14" pageNumber="15">F2>F1>F3)</emphasis>
|
||
, antero-ventral margin with 3 spines (length
|
||
<emphasis id="17CBEAF7FF89FF920C8B4A5D2FFDF89E" bold="true" box="[247,420,1824,1850]" pageId="14" pageNumber="15">FI>FII>FIII)</emphasis>
|
||
, between FI and proximal margin of femora with 1 spine about half of
|
||
<emphasis id="17CBEAF7FF89FF92095C4A5D2B66F89E" bold="true" box="[1312,1343,1824,1850]" pageId="14" pageNumber="15">FI</emphasis>
|
||
length. Pedipalpal patella on antero-dorsal margin with 4 major spines (length
|
||
<emphasis id="17CBEAF7FF89FF920FAA4A382ACDF8FB" bold="true" box="[982,1172,1861,1887]" pageId="14" pageNumber="15">P1>P2>P3>P4)</emphasis>
|
||
, the spines located for most of patella length, the most distal spines between
|
||
<emphasis id="17CBEAF7FF89FF920F5D4A142D18F827" bold="true" box="[801,833,1897,1923]" pageId="14" pageNumber="15">P1</emphasis>
|
||
and distal margin about half of
|
||
<emphasis id="17CBEAF7FF89FF9208B84A142ABCF827" bold="true" box="[1220,1253,1897,1923]" pageId="14" pageNumber="15">P1</emphasis>
|
||
length; anteroventral margin with 3 major spines (length
|
||
<emphasis id="17CBEAF7FF89FF920EF34AF32D6DF80C" bold="true" box="[655,820,1934,1960]" pageId="14" pageNumber="15">PI>PII>PIII)</emphasis>
|
||
. Tibia: antero-dorsal margin with 2 spines, proximal spines less than half distal spine. Pedipalp tarsus with two long spines, the distal spine longer than proximal one. Genitalia: with paired gonopods, soft and small, tube-like (
|
||
<figureCitation id="BD842A60FF89FF920FFE4AA52D83F856" box="[898,986,2008,2034]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="14" pageNumber="15">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="71C0666DFF88FF930CEB4A122F32F842" httpUri="https://zenodo.org/record/197817/files/figure.png" pageId="15" pageNumber="16" targetBox="[282,1316,214,1873]" targetPageId="15">
|
||
<paragraph id="250036E5FF88FF930CEB4A122F32F842" blockId="15.[151,1437,1903,2022]" pageId="15" pageNumber="16">
|
||
<emphasis id="17CBEAF7FF88FF930CEB4A122F04F823" bold="true" box="[151,349,1903,1927]" pageId="15" pageNumber="16">FIGURES 33–40.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="E2BF4D66FF88FF930E634A122C04F822" box="[543,605,1903,1926]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="17CBEAF7FF88FF930E634A122C04F822" box="[543,605,1903,1926]" italics="true" pageId="15" pageNumber="16">Sarax</emphasis>
|
||
</taxonomicName>
|
||
species from Borneo: 33.
|
||
<taxonomicName id="E2BF4D66FF88FF930F0A4A122A57F822" box="[886,1038,1903,1926]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cavernicola">
|
||
<emphasis id="17CBEAF7FF88FF930F0A4A122A57F822" box="[886,1038,1903,1926]" italics="true" pageId="15" pageNumber="16">S. cavernicola</emphasis>
|
||
</taxonomicName>
|
||
, dorsal view; 34. same, ventral view; 35.
|
||
<taxonomicName id="E2BF4D66FF88FF930CBC4AF22F6CF802" box="[192,309,1935,1958]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="yayukae">
|
||
<emphasis id="17CBEAF7FF88FF930CBC4AF22F6CF802" box="[192,309,1935,1958]" italics="true" pageId="15" pageNumber="16">S. yayukae</emphasis>
|
||
</taxonomicName>
|
||
from West and Central Kalimantan, dorsal view; 36. same, ventral view; 37.
|
||
<taxonomicName id="E2BF4D66FF88FF9308FA4AF22B3EF802" box="[1158,1383,1935,1958]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sangkulirangensis" status="sp. nov.">
|
||
<emphasis id="17CBEAF7FF88FF9308FA4AF22B3EF802" box="[1158,1383,1935,1958]" italics="true" pageId="15" pageNumber="16">S. sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="0CF8578CFF88FF9309134AF22EBEF862" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
|
||
, dorsal view; 38. ventral view; 39.
|
||
<taxonomicName id="E2BF4D66FF88FF930E234AD22C95F862" box="[607,716,1967,1990]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mardua">
|
||
<emphasis id="17CBEAF7FF88FF930E234AD22C95F862" box="[607,716,1967,1990]" italics="true" pageId="15" pageNumber="16">S. mardua</emphasis>
|
||
</taxonomicName>
|
||
from Gua
|
||
<taxonomicName id="E2BF4D66FF88FF930F3C4AD22DCFF862" box="[832,918,1967,1990]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="15" pageNumber="19" phylum="Arthropoda" rank="species" species="mardua">Mardua</taxonomicName>
|
||
, East Kalimantan, dorsal view, 40. Ventral view. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="71C0666DFF97FF8C0CEB4BC22F42F8B2" httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="16" pageNumber="17" targetBox="[200,1385,221,1687]" targetPageId="16">
|
||
<paragraph id="250036E5FF97FF8C0CEB4BC22F42F8B2" blockId="16.[151,1436,1727,1814]" pageId="16" pageNumber="17">
|
||
<emphasis id="17CBEAF7FF97FF8C0CEB4BC22F3BF973" bold="true" box="[151,354,1727,1751]" pageId="16" pageNumber="17">FIGURES 41–42.</emphasis>
|
||
Map showing the distribution of
|
||
<taxonomicName id="E2BF4D66FF97FF8C0EA94BBD2D4CF973" box="[725,789,1728,1751]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="17CBEAF7FF97FF8C0EA94BBD2D4CF973" box="[725,789,1728,1751]" italics="true" pageId="16" pageNumber="17">Sarax</emphasis>
|
||
</taxonomicName>
|
||
species: 41. The distribution in Southeast Asia, except for
|
||
<taxonomicName id="E2BF4D66FF97FF8C0CEB4B9D2E8CF953" box="[151,213,1760,1783]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="17CBEAF7FF97FF8C0CEB4B9D2E8CF953" box="[151,213,1760,1783]" italics="true" pageId="16" pageNumber="17">Sarax</emphasis>
|
||
</taxonomicName>
|
||
species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="250036E5FF97FF8C0CB94A382BC5F850" blockId="16.[151,1437,1861,2036]" pageId="16" pageNumber="17">Measurements (in mm): male (n=1) [female (n=1)]; values for segments of the appendages are their lengths. Body length (excluding chelicera) 10.06 [8.47]. Carapace: median length 3.82 [3.35]; width 5.59 [4.47]; median eyes to anterior margin of carapace 0.08 [0.08]; distance between lateral eyes 2.71 [2.19]; lateral eye to anterior margin of carapace 0.54 [0.47]; lateral eye to lateral margin of carapace 0.16 [0.21]. Pedipalps: trochanter 1.43 [1.01]; femur 4.27 [3.31]; patella 5.18 [3.35]; tibia 1.87 [1.09]; tarsus 2.00 [1.49].</paragraph>
|
||
<paragraph id="250036E5FF96FF8D0CEB4DE52AB1FE86" blockId="17.[151,1437,152,663]" pageId="17" pageNumber="18">Leg I: femur 15.39 [11.39]; patella 0.79[0.70]. Leg II: femur 8.78 [6.79], patella 1.38 [0.86]; basitibia 9.45[5.63]; distitibia 5.08 [3.61]; metatarsus + tarsus 3.61 [3.07]. Leg III: femur 10.07 [7.37]; patella 1.41 [0.84]; basitibia 9.29 [7.10]; distitibia 4.81 [3.98]; metatarsus+ tarsus 3.56 [2.93]. Leg IV: femur 8.99 [6.87]; patella 1.07 [0.78]; basitibia 9.65 [7.74]; distitibia 4.38 [3.52]; metatarsus + tarsus 3.64 [3.02].</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF96FF8D0CB94C502CC0FEC8" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="250036E5FF96FF8D0CB94C502CC0FEC8" blockId="17.[151,1437,152,663]" pageId="17" pageNumber="18">
|
||
<emphasis id="17CBEAF7FF96FF8D0CB94C502F04FEE3" bold="true" box="[197,349,301,327]" pageId="17" pageNumber="18">Etymology:</emphasis>
|
||
The species name is derived from the name of the cave “Gua
|
||
<taxonomicName id="E2BF4D66FF96FF8D08114C502A89FEE3" box="[1133,1232,301,327]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="19" phylum="Arthropoda" rank="species" species="mardua">Mardua</taxonomicName>
|
||
” where the
|
||
<typeStatus id="FA048847FF96FF8D091B4C502BC5FEE3" box="[1383,1436,301,327]" pageId="17" pageNumber="18">type</typeStatus>
|
||
material was collected (Gua means “cave”).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF96FF8D0CB94C0A2C20FDA6" pageId="17" pageNumber="18" type="distribution">
|
||
<paragraph id="250036E5FF96FF8D0CB94C0A2F65FE13" blockId="17.[151,1437,152,663]" pageId="17" pageNumber="18">
|
||
<emphasis id="17CBEAF7FF96FF8D0CB94C0A2F31FE35" bold="true" box="[197,360,375,401]" pageId="17" pageNumber="18">Distribution:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF96FF8D0D134C052C4CFE35" box="[367,533,376,401]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mardua">
|
||
<emphasis id="17CBEAF7FF96FF8D0D134C052C4CFE35" box="[367,533,376,401]" italics="true" pageId="17" pageNumber="18">Sarax mardua</emphasis>
|
||
</taxonomicName>
|
||
has only been found in the Sangkulirang Karst, East Kalimantan (
|
||
<collectingCountry id="5DA87675FF96FF8D09624C052BCDFE36" box="[1310,1428,376,402]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
|
||
) (
|
||
<figureCitation id="BD842A60FF96FF8D0CE34CE02F69FE13" box="[159,304,413,439]" captionStart="FIGURES 41 – 42" captionStartId="16.[151,272,1727,1751]" captionTargetBox="[200,1385,221,1687]" captionTargetId="figure@16.[191,1395,194,1706]" captionTargetPageId="16" captionText="FIGURES 41 – 42. Map showing the distribution of Sarax species: 41. The distribution in Southeast Asia, except for Sarax species from India and New Guinea. 42. A detailed map showing the distribution in the Sangkulirang Karst, East Kalimantan." httpUri="https://zenodo.org/record/197818/files/figure.png" pageId="17" pageNumber="18">Figs. 41, 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="250036E5FF96FF8D0CB94CBF2C20FDA6" blockId="17.[151,1437,152,663]" pageId="17" pageNumber="18">
|
||
<emphasis id="17CBEAF7FF96FF8D0CB94CBF2FCBFE78" bold="true" box="[197,402,450,476]" pageId="17" pageNumber="18">Natural history:</emphasis>
|
||
The species is known only from Gua
|
||
<taxonomicName id="E2BF4D66FF96FF8D0F264CBF2DEFFE78" box="[858,950,450,476]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="19" phylum="Arthropoda" rank="species" species="mardua">Mardua</taxonomicName>
|
||
where it occurs with a mite harvestman,
|
||
<taxonomicName id="E2BF4D66FF96FF8D0CEB4C952F4CFDA5" box="[151,277,488,513]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="17CBEAF7FF96FF8D0CEB4C952F4CFDA5" box="[151,277,488,513]" italics="true" pageId="17" pageNumber="18">Stylocellus</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="E2BF4D66FF96FF8D0D314C952F9AFDA6" box="[333,451,488,514]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
|
||
,
|
||
<taxonomicName id="E2BF4D66FF96FF8D0DB34C952C34FDA6" box="[463,621,488,514]" class="Arachnida" family="Stylocellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6DA5656EFF96FF8D0CB94F702B1FFD33" pageId="17" pageNumber="18" type="discussion">
|
||
<paragraph id="250036E5FF96FF8D0CB94F702B1FFD33" blockId="17.[151,1437,152,663]" pageId="17" pageNumber="18">
|
||
<emphasis id="17CBEAF7FF96FF8D0CB94F702F1CFD83" bold="true" box="[197,325,525,551]" pageId="17" pageNumber="18">Remarks:</emphasis>
|
||
<taxonomicName id="E2BF4D66FF96FF8D0D324F702FA5FD82" box="[334,508,525,550]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mardua">
|
||
<emphasis id="17CBEAF7FF96FF8D0D324F702FA5FD82" box="[334,508,525,550]" italics="true" pageId="17" pageNumber="18">Sarax mardua</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="E2BF4D66FF96FF8D0F324F702A14FD82" box="[846,1101,525,550]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF96FF8D0F324F702A14FD82" box="[846,1101,525,550]" italics="true" pageId="17" pageNumber="18">S. sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
by the presence of distinct sexual dimorphism, and the reduced median eye tubercle which is completely divided into two parts (
|
||
<figureCitation id="BD842A60FF96FF8D094B4F4F2BD7FDE8" box="[1335,1422,562,588]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="17" pageNumber="18">Fig. 26</figureCitation>
|
||
). The arrangement of the trichobothria differs from
|
||
<taxonomicName id="E2BF4D66FF96FF8D0E884F252DB5FDD5" box="[756,1004,600,625]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF96FF8D0E884F252DB5FDD5" box="[756,1004,600,625]" italics="true" pageId="17" pageNumber="18">S. sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
where trichobothrium
|
||
<emphasis id="17CBEAF7FF96FF8D097D4F252B44FDD5" box="[1281,1309,600,625]" italics="true" pageId="17" pageNumber="18">bc</emphasis>
|
||
is slightly closer to
|
||
<emphasis id="17CBEAF7FF96FF8D0D7E4F002F7DFD32" box="[258,292,637,662]" italics="true" pageId="17" pageNumber="18">sbf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF96FF8D0DF84F002FC2FD32" box="[388,411,637,662]" italics="true" pageId="17" pageNumber="18">bf</emphasis>
|
||
(
|
||
<figureCitation id="BD842A60FF96FF8D0DD64F002C5BFD33" box="[426,514,637,663]" captionStart="FIGURES 26 – 32" captionStartId="13.[151,273,1898,1922]" captionTargetBox="[191,1379,393,1845]" captionTargetId="figure@13.[151,1436,381,1877]" captionTargetPageId="13" captionText="FIGURES 26 – 32. Sarax mardua sp. nov., male holotype unless stated otherwise: 26. Frontal area of carapace; 27. Sternal area of opisthosoma, ventral view; 28. Antero-dorsal view of left pedipalp; 29. Fourth basitibial segment and distibiae of right leg IV; 30. Antero-dorsal view of left pedipalp, paratype female (MNHN-Am. 12); 31. External view of left chelicerae; 32. Female gonopods, dorsal view, (MNHN-Am. 12)." httpUri="https://zenodo.org/record/197816/files/figure.png" pageId="17" pageNumber="18">Fig. 29</figureCitation>
|
||
); in
|
||
<taxonomicName id="E2BF4D66FF96FF8D0E4A4F002D71FD32" box="[566,808,637,662]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sangkulirangensis">
|
||
<emphasis id="17CBEAF7FF96FF8D0E4A4F002D71FD32" box="[566,808,637,662]" italics="true" pageId="17" pageNumber="18">S. sangkulirangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="17CBEAF7FF96FF8D0F4B4F002D0AFD32" box="[823,851,637,662]" italics="true" pageId="17" pageNumber="18">bc</emphasis>
|
||
is slightly closer to
|
||
<emphasis id="17CBEAF7FF96FF8D08434F002A0CFD32" box="[1087,1109,637,662]" italics="true" pageId="17" pageNumber="18">bf</emphasis>
|
||
than to
|
||
<emphasis id="17CBEAF7FF96FF8D08C94F002A81FD32" box="[1205,1240,637,662]" italics="true" pageId="17" pageNumber="18">sbf</emphasis>
|
||
(
|
||
<figureCitation id="BD842A60FF96FF8D08944F002B64FD33" box="[1256,1341,637,663]" captionStart="FIGURES 21 – 25" captionStartId="11.[151,274,1221,1245]" captionTargetBox="[204,1355,266,1164]" captionTargetId="figure@11.[151,1436,194,1200]" captionTargetPageId="11" captionText="FIGURES 21 – 25. Sarax sangkulirangensis sp. nov., female holotype unless stated otherwise: 21. Frontal area of carapace of male paratype (MZB. Ambl. 150); 22. External view of left chelicerae; 23. Sternal area of opisthosoma, ventral view; 24. Antero-dorsal view of left pedipalp; 25. Fourth basitibial segment and distibiae of right leg IV, male paratype (MZB. Ambl. 150)." httpUri="https://zenodo.org/record/197815/files/figure.png" pageId="17" pageNumber="18">Fig. 25</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |