treatments-xml/data/1A/0E/87/1A0E87F8BF45FFAFB0D9E70B674CFEDB.xml

<document id="7F2590015108265FE42A65DC40122CFC" ID-DOI="10.11646/zootaxa.5453.1.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="11233458" ID-ZooBank="6543A9F4-67A2-475A-AD10-863378CB6364" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1716281923545" checkinUser="plazi" docAuthor="Shrubovych, Julia &amp; D’Haese, Cyrille" docDate="2024" docId="1A0E87F8BF45FFAFB0D9E70B674CFEDB" docLanguage="en" docName="zootaxa.5453.1.4.pdf" docOrigin="Zootaxa 5453 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5453.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphientulus markstivensi Shrubovych &amp; D’Haese 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="96" masterDocId="E637FF80BF40FFA5B04EE335662AFFBF" masterDocTitle="Phylogeny of Australian Acerentomidae species (Protura: Acerentomidae), with a key to identification" masterLastPageNumber="104" masterPageNumber="86" pageNumber="91" updateTime="1716284322880" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BAB042ED1D8E1E98259BFE78F8E35070">Phylogeny of Australian Acerentomidae species (Protura: Acerentomidae), with a key to identification</mods:title>
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<mods:namePart id="F3ECE9E1FA9CB18E8B379B9E6AF55EE9">Shrubovych, Julia</mods:namePart>
<mods:affiliation id="FC8D01959BB24533AFC262C5FE3F5195">Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, ul. Sławkowska 17, 31 - 016 Kraków, Poland &amp; State Museum of Natural History, Ukrainian National Academy of Sciences, Teatral’na St. 18, UA 79008 L’viv, Ukraine &amp; Institute of Soil Biology and Biogeochemistry, Biology Centre CAS, Na Sádkách 7, CZ- 370 05, Česke Budĕjovice, Czech Republic</mods:affiliation>
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<mods:namePart id="C26BD7A2D9C987F317E0F72EE25FD507">D’Haese, Cyrille</mods:namePart>
<mods:affiliation id="DD9BD6C13149E53C1EFDF6A982C1E61B">MECADEV, UMR 7179 CNRS MNHN, Muséum National d’Histoire Naturelle, CP 50 Entomology, 45 rue Buffon, 75231 Paris cedex 05, France</mods:affiliation>
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<treatment id="1A0E87F8BF45FFAFB0D9E70B674CFEDB" ID-DOI="http://doi.org/10.5281/zenodo.11233383" ID-Zenodo-Dep="11233383" LSID="urn:lsid:plazi:treatment:1A0E87F8BF45FFAFB0D9E70B674CFEDB" httpUri="http://treatment.plazi.org/id/1A0E87F8BF45FFAFB0D9E70B674CFEDB" lastPageId="10" lastPageNumber="96" pageId="5" pageNumber="91">
<subSubSection id="DABD6565BF45FFA0B0D9E70B6462FBE6" box="[151,584,1086,1113]" pageId="5" pageNumber="91" type="nomenclature">
<paragraph id="921836EEBF45FFA0B0D9E70B6462FBE6" blockId="5.[151,584,1086,1149]" box="[151,584,1086,1113]" pageId="5" pageNumber="91">
<heading id="C9508182BF45FFA0B0D9E70B6462FBE6" bold="true" box="[151,584,1086,1113]" fontSize="11" level="1" pageId="5" pageNumber="91" reason="1">
<emphasis id="A0D3EAFCBF45FFA0B0D9E70B6462FBE6" bold="true" box="[151,584,1086,1113]" pageId="5" pageNumber="91">
<taxonomicName id="55A74D6DBF45FFA0B0D9E70B67C9FBE6" authority="Shrubovych &amp; D’Haese, 2024" authorityName="Shrubovych &amp; D’Haese" authorityYear="2024" box="[151,483,1086,1113]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="5" pageNumber="91" phylum="Arthropoda" rank="species" species="markstivensi" status="sp. nov.">
<emphasis id="A0D3EAFCBF45FFA0B0D9E70B67C9FBE6" bold="true" box="[151,483,1086,1113]" italics="true" pageId="5" pageNumber="91">Amphientulus markstivensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBE05787BF45FFA0B1A4E70A6462FBE6" box="[490,584,1087,1113]" pageId="5" pageNumber="91" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DABD6565BF45FFA0B0D9E75667CEFBC2" box="[151,484,1122,1149]" pageId="5" pageNumber="91" type="description">
<paragraph id="921836EEBF45FFA0B0D9E75667CEFBC2" blockId="5.[151,584,1086,1149]" box="[151,484,1122,1149]" pageId="5" pageNumber="91">
(
<figureCitation id="0A9C2A6BBF45FFA0B0D1E756670DFBC2" box="[159,295,1123,1149]" captionStart-0="FIGURES 21–30" captionStart-1="FIGURES 31–44" captionStartId-0="6.[151,264,1285,1309]" captionStartId-1="7.[151,264,1769,1793]" captionTargetBox-0="[151,1436,181,1261]" captionTargetBox-1="[294,1292,181,1746]" captionTargetId-0="figure-17@6.[151,1436,181,1261]" captionTargetId-1="figure-17@7.[294,1292,181,1746]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." captionText-1="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi-0="http://doi.org/10.5281/zenodo.11233466" figureDoi-1="http://doi.org/10.5281/zenodo.11233468" httpUri-0="https://zenodo.org/record/11233466/files/figure.png" httpUri-1="https://zenodo.org/record/11233468/files/figure.png" pageId="5" pageNumber="91">Figs. 21–44</figureCitation>
, 
<figureCitation id="0A9C2A6BBF45FFA0B17DE7576756FBC2" box="[307,380,1122,1149]" captionStart-0="FIGURES 45–50" captionStart-1="FIGURES 51–56" captionStart-2="FIGURES 57–62" captionStartId-0="9.[151,264,1771,1795]" captionStartId-1="10.[151,264,1906,1930]" captionStartId-2="11.[151,264,1693,1717]" captionTargetBox-0="[151,1436,237,1748]" captionTargetBox-1="[151,1436,381,1882]" captionTargetBox-2="[151,1436,181,1669]" captionTargetId-0="figure-41@9.[151,1436,237,1748]" captionTargetId-1="figure-109@10.[151,1436,381,1882]" captionTargetId-2="figure-17@11.[151,1436,181,1669]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." captionText-1="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." captionText-2="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi-0="http://doi.org/10.5281/zenodo.11233470" figureDoi-1="http://doi.org/10.5281/zenodo.11233474" figureDoi-2="http://doi.org/10.5281/zenodo.11233488" httpUri-0="https://zenodo.org/record/11233470/files/figure.png" httpUri-1="https://zenodo.org/record/11233474/files/figure.png" httpUri-2="https://zenodo.org/record/11233488/files/figure.png" pageId="5" pageNumber="91">45–62</figureCitation>
; 
<tableCitation id="DF250355BF45FFA0B1C9E75667F1FBC2" box="[391,475,1123,1149]" captionStart="TABLE 2" captionStartId="8.[152,244,152,176]" captionText="TABLE 2. Body chaetotaxy of Amphientulus markstivensi sp. nov." pageId="5" pageNumber="91">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DABD6565BF45FFA0B0D9E79F65B4FAEA" pageId="5" pageNumber="91" type="materials_examined">
<paragraph id="921836EEBF45FFA0B0D9E79F6753FB7B" blockId="5.[151,1437,1194,1905]" box="[151,377,1194,1220]" pageId="5" pageNumber="91">
<emphasis id="A0D3EAFCBF45FFA0B0D9E79F6753FB7B" bold="true" box="[151,377,1194,1220]" pageId="5" pageNumber="91">Material examined</emphasis>
</paragraph>
<paragraph id="921836EEBF45FFA0B0D9E7FA65B4FAEA" blockId="5.[151,1437,1194,1905]" pageId="5" pageNumber="91">
<materialsCitation id="22CF3CB3BF45FFA0B0D9E7FA646FFA8E" collectingDate="2015-03-31" collectionCode="ISEA" collectorName="C. D'Haese." country="Australia" county="Derwent Bridge Cradle Mountain" elevation="946" latitude="42.10393" location="Shadow Lake" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" pageId="5" pageNumber="91" specimenCode="ISEA 6652" specimenCount="1" stateProvince="Tasmania" typeStatus="holotype">
<typeStatus id="4D1C884CBF45FFA0B0D9E7FA672EFB56" box="[151,260,1231,1257]" pageId="5" pageNumber="91" type="holotype">Holotype</typeStatus>
: female (
<specimenCode id="C2019E95BF45FFA0B128E7FA67CDFB56" box="[358,487,1230,1257]" collectionCode="ISEA" pageId="5" pageNumber="91">ISEA 6652</specimenCode>
): 
<collectingCountry id="EAB0767EBF45FFA0B1B6E7FA6448FB56" box="[504,610,1231,1257]" name="Australia" pageId="5" pageNumber="91">Australia</collectingCountry>
, 
<collectingRegion id="5063F80CBF45FFA0B224E7FA64F2FB56" box="[618,728,1231,1257]" country="Australia" name="Tasmania" pageId="5" pageNumber="91">Tasmania</collectingRegion>
, 
<collectingCounty id="7B794E62BF45FFA0B2ACE7FA6270FB56" box="[738,1114,1231,1257]" pageId="5" pageNumber="91">Derwent Bridge Cradle Mountain</collectingCounty>
, 
<collectingMunicipality id="727CAC94BF45FFA0B42AE7FA63B2FB56" box="[1124,1432,1231,1257]" pageId="5" pageNumber="91">Lake St Clair National Park</collectingMunicipality>
, 
<location id="97786035BF45FFA0B0D9E7C66718FAB2" LSID="urn:lsid:plazi:treatment:1A0E87F8BF45FFAFB0D9E70B674CFEDB:97786035BF45FFA0B0D9E7C66718FAB2" box="[151,306,1267,1293]" country="Australia" county="Derwent Bridge Cradle Mountain" latitude="42.10393" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" name="Shadow Lake" pageId="5" pageNumber="91" stateProvince="Tasmania">Shadow Lake</location>
track to 
<location id="97786035BF45FFA0B1D5E7C6641BFAB2" LSID="urn:lsid:plazi:treatment:1A0E87F8BF45FFAFB0D9E70B674CFEDB:97786035BF45FFA0B1D5E7C6641BFAB2" box="[411,561,1267,1293]" country="Australia" county="Derwent Bridge Cradle Mountain" latitude="42.10393" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" name="Mount Rufus" pageId="5" pageNumber="91" stateProvince="Tasmania">Mount Rufus</location>
track, 
<location id="97786035BF45FFA0B2CCE7C664FCFAB2" LSID="urn:lsid:plazi:treatment:1A0E87F8BF45FFAFB0D9E70B674CFEDB:97786035BF45FFA0B2CCE7C664FCFAB2" box="[642,726,1267,1293]" country="Australia" county="Derwent Bridge Cradle Mountain" latitude="42.10393" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" name="Berlese" pageId="5" pageNumber="91" stateProvince="Tasmania">Berlese</location>
of buton grass litter, 
<geoCoordinate id="F7935029BF45FFA0B382E7C6627DFAB2" box="[972,1111,1267,1293]" degrees="42.10393" direction="north" orientation="latitude" pageId="5" pageNumber="91" precision="1" value="42.10393">N42.10393°</geoCoordinate>
, 
<geoCoordinate id="F7935029BF45FFA0B42AE7C66329FAB2" box="[1124,1283,1266,1293]" degrees="146.12159" direction="east" orientation="longitude" pageId="5" pageNumber="91" precision="1" value="146.1216">E 146.12159°</geoCoordinate>
, 
<quantity id="555F9B0BBF45FFA0B55EE7C66373FAB2" box="[1296,1369,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.459999999999999" pageId="5" pageNumber="91" unit="m" value="946.0">
<elevation id="198AD1DDBF45FFA0B55EE7C66373FAB2" box="[1296,1369,1267,1293]" metricMagnitude="2" metricUnit="m" metricValue="9.459999999999999" pageId="5" pageNumber="91" unit="m" value="946.0">946 m</elevation>
</quantity>
elev., 
<date id="E619102EBF45FFA0B0D9E623673DFA8E" box="[151,279,1302,1329]" pageId="5" pageNumber="91" value="2015-03-31">
<collectingDate id="F65DE9C6BF45FFA0B0D9E623673DFA8E" box="[151,279,1302,1329]" pageId="5" pageNumber="91" value="2015-03-31">31.III.2015</collectingDate>
</date>
, collected by 
<collectorName id="3F525338BF45FFA0B1F7E622646FFA8E" box="[441,581,1303,1329]" pageId="5" pageNumber="91">C. D’Haese.</collectorName>
</materialsCitation>
<materialsCitation id="22CF3CB3BF45FFA0B200E622637AFA8E" box="[590,1360,1302,1329]" collectingDate="2015-03-31" collectionCode="ISEA" collectorName="C. D'Haese." country="Australia" county="Derwent Bridge Cradle Mountain" elevation="946" latitude="42.10393" location="Shadow Lake" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" pageId="5" pageNumber="91" specimenCode="ISEA 6653, 6654" specimenCount="2" specimenCount-female="2" stateProvince="Tasmania" typeStatus="paratype">
<typeStatus id="4D1C884CBF45FFA0B200E6226491FA8E" box="[590,699,1303,1329]" pageId="5" pageNumber="91" type="paratype">Paratypes</typeStatus>
<specimenCount id="84A1FD67BF45FFA0B28AE6226518FA8E" box="[708,818,1303,1329]" count="2" pageId="5" pageNumber="91" type="female">2 females</specimenCount>
(
<specimenCode id="C2019E95BF45FFA0B30CE62265EFFA8E" box="[834,965,1302,1329]" collectionCode="ISEA" pageId="5" pageNumber="91">ISEA 6653</specimenCode>
, 
<specimenCode id="C2019E95BF45FFA0B39CE6236224FA8E" box="[978,1038,1302,1329]" collectionCode="ISEA" pageId="5" pageNumber="91">6654</specimenCode>
), the same data as holotype
</materialsCitation>
. 
<materialsCitation id="22CF3CB3BF45FFA0B512E62265B1FAEA" collectingDate="2015-03-31" collectionCode="ISEA" collectorName="C. D'Haese." country="Australia" county="Derwent Bridge Cradle Mountain" elevation="946" latitude="42.10393" location="Shadow Lake" longLatPrecision="1" longitude="146.1216" municipality="Lake St Clair National Park" pageId="5" pageNumber="91" specimenCount="1" specimenCount-larva="1" stateProvince="Tasmania">
<typeStatus id="4D1C884CBF45FFA0B512E6226728FAEA" isOtherTypeStatus="true" pageId="5" pageNumber="91">Other materials</typeStatus>
: 2 maturi juniores and 
<specimenCount id="84A1FD67BF45FFA0B24CE60E6472FAEA" box="[514,600,1339,1365]" count="1" pageId="5" pageNumber="91" type="larva">1 Larva</specimenCount>
I, the same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DABD6565BF45FFADB089E66B6211F83B" lastPageId="8" lastPageNumber="94" pageId="5" pageNumber="91" type="description">
<paragraph id="921836EEBF45FFA0B089E66B650DF992" blockId="5.[151,1437,1194,1905]" pageId="5" pageNumber="91">
<emphasis id="A0D3EAFCBF45FFA0B089E66B677BFAC7" bold="true" box="[199,337,1374,1400]" pageId="5" pageNumber="91">Description</emphasis>
. Body long and slender (
<figureCitation id="0A9C2A6BBF45FFA0B23EE66A64EFFAC6" box="[624,709,1374,1401]" captionStart="FIGURES 45–50" captionStartId="9.[151,264,1771,1795]" captionTargetBox="[151,1436,237,1748]" captionTargetId="figure-41@9.[151,1436,237,1748]" captionTargetPageId="9" captionText="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." figureDoi="http://doi.org/10.5281/zenodo.11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="5" pageNumber="91">Fig. 45</figureCitation>
). Head with short setae, cephalic setae 
<emphasis id="A0D3EAFCBF45FFA0B4CCE66A62BDFAC6" box="[1154,1175,1375,1401]" italics="true" pageId="5" pageNumber="91">l3</emphasis>
, 
<emphasis id="A0D3EAFCBF45FFA0B4E8E66A62E7FAC6" box="[1190,1229,1375,1401]" italics="true" pageId="5" pageNumber="91">sd4</emphasis>
and 
<emphasis id="A0D3EAFCBF45FFA0B54BE66A6306FAC6" box="[1285,1324,1375,1401]" italics="true" pageId="5" pageNumber="91">sd5</emphasis>
short and setiform (
<figureCitation id="0A9C2A6BBF45FFA0B14CE6B667A2FA22" box="[258,392,1411,1437]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Figs. 21, 22</figureCitation>
, 
<figureCitation id="0A9C2A6BBF45FFA0B1DCE6B667F6FA22" box="[402,476,1411,1437]" captionStart="FIGURES 45–50" captionStartId="9.[151,264,1771,1795]" captionTargetBox="[151,1436,237,1748]" captionTargetId="figure-41@9.[151,1436,237,1748]" captionTargetPageId="9" captionText="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." figureDoi="http://doi.org/10.5281/zenodo.11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="5" pageNumber="91">46–48</figureCitation>
), seta 
<emphasis id="A0D3EAFCBF45FFA0B251E6B76411FA23" box="[543,571,1410,1436]" italics="true" pageId="5" pageNumber="91">d6</emphasis>
absent (
<figureCitation id="0A9C2A6BBF45FFA0B2D8E6B664C2FA22" box="[662,744,1411,1437]" captionStart="FIGURES 45–50" captionStartId="9.[151,264,1771,1795]" captionTargetBox="[151,1436,237,1748]" captionTargetId="figure-41@9.[151,1436,237,1748]" captionTargetPageId="9" captionText="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." figureDoi="http://doi.org/10.5281/zenodo.11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="5" pageNumber="91">Fig. 46</figureCitation>
), length ratio of posterior setae 
<emphasis id="A0D3EAFCBF45FFA0B40DE6B66275FA22" box="[1091,1119,1411,1437]" italics="true" pageId="5" pageNumber="91">d7</emphasis>
: 
<emphasis id="A0D3EAFCBF45FFA0B422E6B662B9FA22" box="[1132,1171,1411,1437]" italics="true" pageId="5" pageNumber="91">sd7</emphasis>
as 1.0:1.5. Pseudoculus circular, with short posterior extension, 
<collectingRegion id="5063F80CBF45FFA0B219E6926453FA7E" box="[599,633,1447,1473]" country="Brazil" name="Para" pageId="5" pageNumber="91">PR</collectingRegion>
= 14 (
<figureCitation id="0A9C2A6BBF45FFA0B28DE692653DFA7E" box="[707,791,1447,1473]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Fig. 21</figureCitation>
). Maxillary palps apically with a tuft of setae, four single setae and two basal sensilla slender and equal in length (
<figureCitation id="0A9C2A6BBF45FFA0B341E6FE6546FA5A" box="[783,876,1483,1509]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Figs. 23</figureCitation>
, 
<figureCitation id="0A9C2A6BBF45FFA0B336E6FE65B2FA5A" box="[888,920,1483,1509]" captionStart="FIGURES 45–50" captionStartId="9.[151,264,1771,1795]" captionTargetBox="[151,1436,237,1748]" captionTargetId="figure-41@9.[151,1436,237,1748]" captionTargetPageId="9" captionText="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." figureDoi="http://doi.org/10.5281/zenodo.11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="5" pageNumber="91">49</figureCitation>
). Labial palps reduced to four setae and broad basal sensillum (
<figureCitation id="0A9C2A6BBF45FFA0B11BE6DA679EF9B6" box="[341,436,1519,1545]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Figs. 24</figureCitation>
, 
<figureCitation id="0A9C2A6BBF45FFA0B18CE6DB67C8F9B6" box="[450,482,1518,1545]" captionStart="FIGURES 45–50" captionStartId="9.[151,264,1771,1795]" captionTargetBox="[151,1436,237,1748]" captionTargetId="figure-41@9.[151,1436,237,1748]" captionTargetPageId="9" captionText="FIGURES 45–50. Amphientulus markstivensi sp. nov. 45) Habitus. 46) Head. 47) Pseudoculus with setae l3 and sd4 (fp = frontal pore). 48) Cephalic seta sd5. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum)." figureDoi="http://doi.org/10.5281/zenodo.11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="5" pageNumber="91">50</figureCitation>
). Maxillary gland with small elongated and weakly granulated calyx, long posterior filament and bilobed posterior dilation, CF = 6.4 (
<figureCitation id="0A9C2A6BBF45FFA0B288E5266531F992" box="[710,795,1554,1581]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Fig. 25</figureCitation>
).
</paragraph>
<paragraph id="921836EEBF45FFA0B089E502672FF8CE" blockId="5.[151,1437,1194,1905]" pageId="5" pageNumber="91">
Foretarsus with sensillum 
<emphasis id="A0D3EAFCBF45FFA0B1B8E5026427F9EE" box="[502,525,1591,1617]" italics="true" pageId="5" pageNumber="91">b’</emphasis>
, 
<emphasis id="A0D3EAFCBF45FFA0B251E502641FF9EE" box="[543,565,1591,1617]" italics="true" pageId="5" pageNumber="91">t1</emphasis>
claviform, 
<emphasis id="A0D3EAFCBF45FFA0B2F2E50264F8F9EE" box="[700,722,1591,1617]" italics="true" pageId="5" pageNumber="91">t3</emphasis>
small and jar-shaped (
<figureCitation id="0A9C2A6BBF45FFA0B394E502624FF9EE" box="[986,1125,1591,1617]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="5" pageNumber="91">Figs. 27–29</figureCitation>
, 
<figureCitation id="0A9C2A6BBF45FFA0B43DE5036294F9EE" box="[1139,1214,1590,1617]" captionStart="FIGURES 51–56" captionStartId="10.[151,264,1906,1930]" captionTargetBox="[151,1436,381,1882]" captionTargetId="figure-109@10.[151,1436,381,1882]" captionTargetPageId="10" captionText="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." figureDoi="http://doi.org/10.5281/zenodo.11233474" httpUri="https://zenodo.org/record/11233474/files/figure.png" pageId="5" pageNumber="91">51–53</figureCitation>
); 
<emphasis id="A0D3EAFCBF45FFA0B49AE50262C1F9EE" box="[1236,1259,1591,1617]" italics="true" pageId="5" pageNumber="91">a’</emphasis>
broadened and sword-shaped, other sensilla slender and parallel-sided. Sensillum 
<emphasis id="A0D3EAFCBF45FFA0B3CBE56E65B9F9CB" box="[901,915,1627,1652]" italics="true" pageId="5" pageNumber="91">a</emphasis>
extending past base of 
<emphasis id="A0D3EAFCBF45FFA0B4D3E56E6299F9CA" box="[1181,1203,1627,1653]" italics="true" pageId="5" pageNumber="91">t2</emphasis>
. Sensillum 
<emphasis id="A0D3EAFCBF45FFA0B577E56E636DF9CA" box="[1337,1351,1627,1653]" italics="true" pageId="5" pageNumber="91">b</emphasis>
shorter than 
<emphasis id="A0D3EAFCBF45FFA0B081E54A66F1F927" box="[207,219,1663,1688]" italics="true" pageId="5" pageNumber="91">c</emphasis>
, reaching base of seta 
<emphasis id="A0D3EAFCBF45FFA0B193E54A67DDF926" box="[477,503,1663,1689]" italics="true" pageId="5" pageNumber="91">γ3</emphasis>
. Sensillum 
<emphasis id="A0D3EAFCBF45FFA0B233E54A64A3F927" box="[637,649,1663,1688]" italics="true" pageId="5" pageNumber="91">c</emphasis>
almost reaching base of sensillum 
<emphasis id="A0D3EAFCBF45FFA0B458E54A620DF926" box="[1046,1063,1663,1689]" italics="true" pageId="5" pageNumber="91">f.</emphasis>
Sensilla 
<emphasis id="A0D3EAFCBF45FFA0B4C3E54A62B1F926" box="[1165,1179,1663,1689]" italics="true" pageId="5" pageNumber="91">b</emphasis>
and 
<emphasis id="A0D3EAFCBF45FFA0B49DE54A62F5F927" box="[1235,1247,1663,1688]" italics="true" pageId="5" pageNumber="91">c</emphasis>
inserted at same level, sensillum 
<emphasis id="A0D3EAFCBF45FFA0B100E5966776F902" box="[334,348,1699,1725]" italics="true" pageId="5" pageNumber="91">d</emphasis>
inserted between sensilla 
<emphasis id="A0D3EAFCBF45FFA0B2CCE59664A4F903" box="[642,654,1699,1724]" italics="true" pageId="5" pageNumber="91">c</emphasis>
and 
<emphasis id="A0D3EAFCBF45FFA0B288E59664F6F902" box="[710,732,1699,1725]" italics="true" pageId="5" pageNumber="91">t2</emphasis>
, close to base of sensillum 
<emphasis id="A0D3EAFCBF45FFA0B45FE5966237F903" box="[1041,1053,1699,1724]" italics="true" pageId="5" pageNumber="91">c</emphasis>
. Sensillum 
<emphasis id="A0D3EAFCBF45FFA0B4ECE5966293F902" box="[1186,1209,1699,1725]" italics="true" pageId="5" pageNumber="91">a’</emphasis>
extending past base of 
<emphasis id="A0D3EAFCBF45FFA0B0FBE5F266E6F95E" box="[181,204,1735,1761]" italics="true" pageId="5" pageNumber="91">b’</emphasis>
. Sensillum 
<emphasis id="A0D3EAFCBF45FFA0B11EE5F2674DF95E" box="[336,359,1735,1761]" italics="true" pageId="5" pageNumber="91">b’</emphasis>
extending past base of sensillum 
<emphasis id="A0D3EAFCBF45FFA0B293E5F264D9F95E" box="[733,755,1735,1761]" italics="true" pageId="5" pageNumber="91">c’</emphasis>
, apex of sensillum 
<emphasis id="A0D3EAFCBF45FFA0B384E5F265CAF95E" box="[970,992,1735,1761]" italics="true" pageId="5" pageNumber="91">c’</emphasis>
extending past the base of claw. Length formula of sensilla: 
<emphasis id="A0D3EAFCBF45FFA0B13AE5DE67A0F8BA" box="[372,394,1771,1797]" italics="true" pageId="5" pageNumber="91">t3</emphasis>
&lt;
<emphasis id="A0D3EAFCBF45FFA0B1E8E5DE6796F8BA" box="[422,444,1771,1797]" italics="true" pageId="5" pageNumber="91">t1</emphasis>
&lt;
<emphasis id="A0D3EAFCBF45FFA0B199E5DE67CFF8BB" box="[471,485,1771,1796]" italics="true" pageId="5" pageNumber="91">g</emphasis>
&lt;(
<emphasis id="A0D3EAFCBF45FFA0B244E5DE6432F8BA" box="[522,536,1771,1797]" italics="true" pageId="5" pageNumber="91">b</emphasis>
= 
<emphasis id="A0D3EAFCBF45FFA0B27DE5DE6460F8BA" box="[563,586,1771,1797]" italics="true" pageId="5" pageNumber="91">a’</emphasis>
) &lt;(
<emphasis id="A0D3EAFCBF45FFA0B237E5DE64ADF8BB" box="[633,647,1771,1796]" italics="true" pageId="5" pageNumber="91">a</emphasis>
= 
<emphasis id="A0D3EAFCBF45FFA0B2ECE5DE649AF8BA" box="[674,688,1771,1797]" italics="true" pageId="5" pageNumber="91">d</emphasis>
) &lt;(
<emphasis id="A0D3EAFCBF45FFA0B290E5DE64CCF8BA" box="[734,742,1771,1797]" italics="true" pageId="5" pageNumber="91">f</emphasis>
= 
<emphasis id="A0D3EAFCBF45FFA0B34CE5DE6533F8BA" box="[770,793,1771,1797]" italics="true" pageId="5" pageNumber="91">b’</emphasis>
) &lt;(
<emphasis id="A0D3EAFCBF45FFA0B309E5DE65A2F8BA" box="[839,904,1771,1797]" italics="true" pageId="5" pageNumber="91">e = t2</emphasis>
) &lt;
<emphasis id="A0D3EAFCBF45FFA0B3E3E5DE65E9F8BA" box="[941,963,1771,1797]" italics="true" pageId="5" pageNumber="91">c’</emphasis>
. Setae 
<emphasis id="A0D3EAFCBF45FFA0B45CE5DE6207F8BA" box="[1042,1069,1771,1797]" italics="true" pageId="5" pageNumber="91">β1</emphasis>
and 
<emphasis id="A0D3EAFCBF45FFA0B42CE5DE6257F8BA" box="[1122,1149,1771,1797]" italics="true" pageId="5" pageNumber="91">δ4</emphasis>
setiform, slightly broader than but same length as other 
<emphasis id="A0D3EAFCBF45FFA0B1ABE43A67D1F896" box="[485,507,1807,1833]" italics="true" pageId="5" pageNumber="91">δ-</emphasis>
setae (8 μm). Pores on foretarsus near base of sensillum 
<emphasis id="A0D3EAFCBF45FFA0B43DE43A62A3F896" box="[1139,1161,1807,1833]" italics="true" pageId="5" pageNumber="91">t3</emphasis>
and near base of seta 
<emphasis id="A0D3EAFCBF45FFA0B5CEE43A63B6F896" box="[1408,1436,1807,1833]" italics="true" pageId="5" pageNumber="91">α3</emphasis>
between bases of sensilla 
<emphasis id="A0D3EAFCBF45FFA0B1F4E40667E2F8F3" box="[442,456,1843,1868]" italics="true" pageId="5" pageNumber="91">a</emphasis>
and 
<emphasis id="A0D3EAFCBF45FFA0B1B1E4066421F8F3" box="[511,523,1843,1868]" italics="true" pageId="5" pageNumber="91">c</emphasis>
. Claw short, without inner tooth, empodial appendage short. BS = 0.3, TR = 3.1, EU = 0.1.
</paragraph>
<caption id="C6D86666BF46FFA3B0D9E63063A1FAD9" ID-DOI="http://doi.org/10.5281/zenodo.11233466" ID-Zenodo-Dep="11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="6" pageNumber="92" startId="6.[151,264,1285,1309]" targetBox="[151,1436,181,1261]" targetPageId="6" targetType="figure">
<paragraph id="921836EEBF46FFA3B0D9E63063A1FAD9" blockId="6.[151,1437,1285,1382]" pageId="6" pageNumber="92">
<emphasis id="A0D3EAFCBF46FFA3B0D9E630677DFAA2" bold="true" box="[151,343,1285,1309]" pageId="6" pageNumber="92">FIGURES 21–30.</emphasis>
<taxonomicName id="55A74D6DBF46FFA3B12EE630645FFAA2" authority="Shrubovych &amp; D’Haese, 2024" authorityName="Shrubovych &amp; D’Haese" authorityYear="2024" box="[352,629,1285,1309]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="6" pageNumber="92" phylum="Arthropoda" rank="species" species="markstivensi" status="sp. nov.">
<emphasis id="A0D3EAFCBF46FFA3B12EE630645FFAA2" box="[352,629,1285,1309]" italics="true" pageId="6" pageNumber="92">Amphientulus markstivensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBE05787BF46FFA3B230E63064F9FAA2" box="[638,723,1285,1309]" pageId="6" pageNumber="92" rank="species">
<emphasis id="A0D3EAFCBF46FFA3B230E63064E6FAA2" bold="true" box="[638,716,1285,1309]" pageId="6" pageNumber="92">sp. nov</emphasis>
.
</taxonomicNameLabel>
, holotype (ISEA 6652). 21) Pseudoculus with setae 
<emphasis id="A0D3EAFCBF46FFA3B4B3E630633BFAA2" box="[1277,1297,1285,1309]" italics="true" pageId="6" pageNumber="92">l3</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B506E6306358FAA2" box="[1352,1394,1285,1309]" italics="true" pageId="6" pageNumber="92">sd4.</emphasis>
22) Cephalic seta 
<emphasis id="A0D3EAFCBF46FFA3B162E61C6765FAFE" box="[300,335,1321,1345]" italics="true" pageId="6" pageNumber="92">sd5</emphasis>
. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum 
<emphasis id="A0D3EAFCBF46FFA3B301E6786549FADA" box="[847,867,1357,1381]" italics="true" pageId="6" pageNumber="92">t3</emphasis>
. 30) Comb. Arrows indicate pores. Scale bars: 20 µm.
</paragraph>
</caption>
<paragraph id="921836EEBF46FFA3B089E682675CF971" blockId="6.[151,1437,1463,1742]" pageId="6" pageNumber="92">
Formula of the chaetotaxy given in 
<tableCitation id="DF250355BF46FFA3B22DE6826491FA6D" box="[611,699,1463,1490]" captionStart="TABLE 2" captionStartId="8.[152,244,152,176]" captionText="TABLE 2. Body chaetotaxy of Amphientulus markstivensi sp. nov." pageId="6" pageNumber="92">Table 2</tableCitation>
. Setae on nota strongly differing in length. Pronotal setae 
<emphasis id="A0D3EAFCBF46FFA3B515E68D63B6FA6E" box="[1371,1436,1464,1490]" italics="true" pageId="6" pageNumber="92">1:2 =</emphasis>
1.0:2.0 (
<figureCitation id="0A9C2A6BBF46FFA3B0BBE6EE6779FA49" box="[245,339,1499,1526]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Figs. 31</figureCitation>
, 
<figureCitation id="0A9C2A6BBF46FFA3B111E6EE6755FA49" box="[351,383,1499,1526]" captionStart="FIGURES 51–56" captionStartId="10.[151,264,1906,1930]" captionTargetBox="[151,1436,381,1882]" captionTargetId="figure-109@10.[151,1436,381,1882]" captionTargetPageId="10" captionText="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." figureDoi="http://doi.org/10.5281/zenodo.11233474" httpUri="https://zenodo.org/record/11233474/files/figure.png" pageId="6" pageNumber="92">54</figureCitation>
). Mesonotal setae 
<emphasis id="A0D3EAFCBF46FFA3B21EE6E96454FA49" box="[592,638,1500,1526]" italics="true" pageId="6" pageNumber="92">P1a</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B2FBE6E964C8FA49" box="[693,738,1500,1526]" italics="true" pageId="6" pageNumber="92">P2a</emphasis>
very short (2 μm), oblong (
<figureCitation id="0A9C2A6BBF46FFA3B455E6EE6253FA49" box="[1051,1145,1499,1526]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Figs. 31</figureCitation>
, 
<figureCitation id="0A9C2A6BBF46FFA3B4C8E6EE628FFA49" box="[1158,1189,1499,1526]" captionStart="FIGURES 51–56" captionStartId="10.[151,264,1906,1930]" captionTargetBox="[151,1436,381,1882]" captionTargetId="figure-109@10.[151,1436,381,1882]" captionTargetPageId="10" captionText="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." figureDoi="http://doi.org/10.5281/zenodo.11233474" httpUri="https://zenodo.org/record/11233474/files/figure.png" pageId="6" pageNumber="92">55</figureCitation>
), metanotal setae 
<emphasis id="A0D3EAFCBF46FFA3B521E6E963B7FA49" box="[1391,1437,1500,1526]" italics="true" pageId="6" pageNumber="92">P1a</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B086E53566DCF9A5" box="[200,246,1536,1562]" italics="true" pageId="6" pageNumber="92">P2a</emphasis>
twice as long (4 μm), oblong (
<figureCitation id="0A9C2A6BBF46FFA3B215E6CA649BF9A5" box="[603,689,1535,1562]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Fig. 32</figureCitation>
). Seta 
<emphasis id="A0D3EAFCBF46FFA3B2B1E5356506F9A5" box="[767,812,1536,1562]" italics="true" pageId="6" pageNumber="92">P2a</emphasis>
on meso- and metanota situated very close to 
<emphasis id="A0D3EAFCBF46FFA3B50CE535634BF9A5" box="[1346,1377,1536,1562]" italics="true" pageId="6" pageNumber="92">P3</emphasis>
, seta 
<emphasis id="A0D3EAFCBF46FFA3B0D9E51166EFF981" box="[151,197,1572,1598]" italics="true" pageId="6" pageNumber="92">P3a</emphasis>
setiform and four times longer than 
<emphasis id="A0D3EAFCBF46FFA3B211E51164A7F981" box="[607,653,1572,1598]" italics="true" pageId="6" pageNumber="92">P1a</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B28DE51164DBF981" box="[707,753,1572,1598]" italics="true" pageId="6" pageNumber="92">P2a</emphasis>
(
<figureCitation id="0A9C2A6BBF46FFA3B34EE5166579F981" box="[768,851,1571,1598]" captionStart="FIGURES 51–56" captionStartId="10.[151,264,1906,1930]" captionTargetBox="[151,1436,381,1882]" captionTargetId="figure-109@10.[151,1436,381,1882]" captionTargetPageId="10" captionText="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." figureDoi="http://doi.org/10.5281/zenodo.11233474" httpUri="https://zenodo.org/record/11233474/files/figure.png" pageId="6" pageNumber="92">Fig. 56</figureCitation>
), seta 
<emphasis id="A0D3EAFCBF46FFA3B3D5E5116590F981" box="[923,954,1572,1598]" italics="true" pageId="6" pageNumber="92">P4</emphasis>
setiform and long, seta 
<emphasis id="A0D3EAFCBF46FFA3B486E51162CDF981" box="[1224,1255,1572,1598]" italics="true" pageId="6" pageNumber="92">P5</emphasis>
gemmate (
<figureCitation id="0A9C2A6BBF46FFA3B52AE51666C8F9DD" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Figs. 31, 32</figureCitation>
). Length ratio of mesonotal setae 
<emphasis id="A0D3EAFCBF46FFA3B22EE57D64AAF9DD" box="[608,640,1608,1634]" italics="true" pageId="6" pageNumber="92">P1</emphasis>
: 
<emphasis id="A0D3EAFCBF46FFA3B2C9E57D648CF9DD" box="[647,678,1608,1634]" italics="true" pageId="6" pageNumber="92">P2</emphasis>
= 1.0:1.5 (
<figureCitation id="0A9C2A6BBF46FFA3B36CE572655CF9DD" box="[802,886,1607,1634]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Fig. 31</figureCitation>
). Mesonotum with 
<emphasis id="A0D3EAFCBF46FFA3B41CE572624FF9DE" box="[1106,1125,1607,1633]" italics="true" pageId="6" pageNumber="92">sl</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B4D2E5726298F9DE" box="[1180,1202,1607,1633]" italics="true" pageId="6" pageNumber="92">al</emphasis>
pores (
<figureCitation id="0A9C2A6BBF46FFA3B548E5726349F9DD" box="[1286,1379,1607,1634]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Figs. 31</figureCitation>
, 
<figureCitation id="0A9C2A6BBF46FFA3B53EE57263A5F9DD" box="[1392,1423,1607,1634]" captionStart="FIGURES 51–56" captionStartId="10.[151,264,1906,1930]" captionTargetBox="[151,1436,381,1882]" captionTargetId="figure-109@10.[151,1436,381,1882]" captionTargetPageId="10" captionText="FIGURES 51–56. Amphientulus markstivensi sp. nov. 51) Foretarsal sensilla a, b, c, t2 and t3. 52) Foretarsal sensilla a’, b’,c’ and t3. 53) Foretarsal sensillum t1. 54) Pronotal setae 1 and 2, mesonotal al and sl pores. 55) Mesonotal seta P1a. 56) Setae P2a and P3a on metanotum." figureDoi="http://doi.org/10.5281/zenodo.11233474" httpUri="https://zenodo.org/record/11233474/files/figure.png" pageId="6" pageNumber="92">55</figureCitation>
), metanotum with 
<emphasis id="A0D3EAFCBF46FFA3B116E55E6741F93A" box="[344,363,1643,1669]" italics="true" pageId="6" pageNumber="92">sl</emphasis>
pore only (
<figureCitation id="0A9C2A6BBF46FFA3B1BEE55E646CF939" box="[496,582,1643,1670]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Fig. 32</figureCitation>
). Pro- and mesosterna without pores; metasternum with one or two closely placed 
<emphasis id="A0D3EAFCBF46FFA3B0A8E5A566D7F916" box="[230,253,1680,1705]" italics="true" pageId="6" pageNumber="92">sc</emphasis>
pores (
<figureCitation id="0A9C2A6BBF46FFA3B101E5BA6788F915" box="[335,418,1679,1706]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Fig. 39</figureCitation>
). Setae 
<emphasis id="A0D3EAFCBF46FFA3B1B9E5A5643CF915" box="[503,534,1680,1706]" italics="true" pageId="6" pageNumber="92">A2</emphasis>
and 
<emphasis id="A0D3EAFCBF46FFA3B204E5A56445F915" box="[586,623,1680,1706]" italics="true" pageId="6" pageNumber="92">M2</emphasis>
on prosternum and seta 
<emphasis id="A0D3EAFCBF46FFA3B333E5A565B7F915" box="[893,925,1680,1706]" italics="true" pageId="6" pageNumber="92">A2</emphasis>
on thoracic sterna short, setiform, their length 6 μm (
<figureCitation id="0A9C2A6BBF46FFA3B0AEE5866740F971" box="[224,362,1715,1742]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="6" pageNumber="92">Figs. 37–39</figureCitation>
).
</paragraph>
<caption id="C6D86666BF47FFA2B0D9E5DC6562F865" ID-DOI="http://doi.org/10.5281/zenodo.11233468" ID-Zenodo-Dep="11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="7" pageNumber="93" startId="7.[151,264,1769,1793]" targetBox="[294,1292,181,1746]" targetPageId="7" targetType="figure">
<paragraph id="921836EEBF47FFA2B0D9E5DC6562F865" blockId="7.[151,1437,1769,2010]" pageId="7" pageNumber="93">
<emphasis id="A0D3EAFCBF47FFA2B0D9E5DC6773F8BD" bold="true" box="[151,345,1769,1794]" pageId="7" pageNumber="93">FIGURES 31–44.</emphasis>
<taxonomicName id="55A74D6DBF47FFA2B12DE5DF6450F8BD" authority="Shrubovych &amp; D’Haese, 2024" authorityName="Shrubovych &amp; D’Haese" authorityYear="2024" box="[355,634,1770,1794]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="markstivensi" status="sp. nov.">
<emphasis id="A0D3EAFCBF47FFA2B12DE5DF6450F8BD" box="[355,634,1770,1794]" italics="true" pageId="7" pageNumber="93">Amphientulus markstivensi</emphasis>
</taxonomicName>
<emphasis id="A0D3EAFCBF47FFA2B2CBE5DF64F0F8BD" bold="true" box="[645,730,1770,1794]" pageId="7" pageNumber="93">
<taxonomicNameLabel id="BBE05787BF47FFA2B2CBE5DF64F0F8BD" box="[645,730,1770,1794]" pageId="7" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm.
</paragraph>
</caption>
<caption id="C6D86666BF48FFADB0D6E3AD6543FF0E" box="[152,873,152,177]" pageId="8" pageNumber="94" startId="8.[152,244,152,176]" targetType="table">
<paragraph id="921836EEBF48FFADB0D6E3AD6543FF0E" blockId="8.[152,873,152,177]" box="[152,873,152,177]" pageId="8" pageNumber="94">
<emphasis id="A0D3EAFCBF48FFADB0D6E3AD6725FF0F" bold="true" box="[152,271,152,176]" pageId="8" pageNumber="94">TABLE 2.</emphasis>
Body chaetotaxy of 
<taxonomicName id="55A74D6DBF48FFADB1A3E3AD6527FF0F" authority="Shrubovych &amp; D’Haese, 2024" authorityName="Shrubovych &amp; D’Haese" authorityYear="2024" box="[493,781,152,176]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="markstivensi" status="sp. nov.">
<emphasis id="A0D3EAFCBF48FFADB1A3E3AD6527FF0F" box="[493,781,152,176]" italics="true" pageId="8" pageNumber="94">Amphientulus markstivensi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBE05787BF48FFADB35AE3AC6543FF0E" box="[788,873,153,177]" pageId="8" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="921836EEBF48FFADB0D1E3DC65E8FC44" pageId="8" pageNumber="94">
<table id="E0A7C44EBF48005AB0D1E3F7633DFC44" box="[159,1303,194,1019]" colsContinueFrom="3.[159,1326,590,1498]" colsContinueIn="13.[159,1308,914,1739]" gridcols="5" gridrows="15" pageId="8" pageNumber="94">
<tr id="2C9734ACBF48005AB0D1E3F7633DFF40" box="[159,1303,194,255]" gridrow="0" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E3F76739FF40" box="[159,275,194,255]" gridcol="0" gridrow="0" pageId="8" pageNumber="94">Segment</th>
<th id="6F465DD0BF48005AB1DFE3F767C8FF40" box="[401,482,194,255]" gridcol="1" gridrow="0" pageId="8" pageNumber="94">Dorsal Formula</th>
<th id="6F465DD0BF48005AB206E3F76503FF40" box="[584,809,194,255]" gridcol="2" gridrow="0" pageId="8" pageNumber="94">Setal composition</th>
<th id="6F465DD0BF48005AB3F8E3F7622DFF40" box="[950,1031,194,255]" gridcol="3" gridrow="0" pageId="8" pageNumber="94">Ventral Formula</th>
<th id="6F465DD0BF48005AB424E3F7633DFF40" box="[1130,1303,194,255]" gridcol="4" gridrow="0" pageId="8" pageNumber="94">Setal composition</th>
</tr>
<tr id="2C9734ACBF48005AB0D1E221633DFEF1" box="[159,1303,276,334]" gridrow="1" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E2216739FEF1" box="[159,275,276,334]" gridcol="0" gridrow="1" pageId="8" pageNumber="94">Th. I</th>
<td id="6F465DD0BF48005AB1DFE22167C8FEF1" box="[401,482,276,334]" gridcol="1" gridrow="1" pageId="8" pageNumber="94">4</td>
<td id="6F465DD0BF48005AB206E2216503FEF1" box="[584,809,276,334]" gridcol="2" gridrow="1" pageId="8" pageNumber="94">1, 2</td>
<td id="6F465DD0BF48005AB3F8E221622DFEF1" box="[950,1031,276,334]" gridcol="3" gridrow="1" pageId="8" pageNumber="94">4+4 6</td>
<td id="6F465DD0BF48005AB424E221633DFEF1" box="[1130,1303,276,334]" gridcol="4" gridrow="1" pageId="8" pageNumber="94">A1, 2, M1, 2 P1, 2, 3</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E252633DFEC2" box="[159,1303,359,381]" gridrow="2" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E2526739FEC2" box="[159,275,359,381]" gridcol="0" gridrow="2" pageId="8" pageNumber="94">Th. II–III</th>
<td id="6F465DD0BF48005AB1DFE25267C8FEC2" box="[401,482,359,381]" gridcol="1" gridrow="2" pageId="8" pageNumber="94">6</td>
<td id="6F465DD0BF48005AB206E2526503FEC2" box="[584,809,359,381]" gridcol="2" gridrow="2" pageId="8" pageNumber="94">A2, 4, M</td>
<td id="6F465DD0BF48005AB3F8E252622DFEC2" box="[950,1031,359,381]" gridcol="3" gridrow="2" pageId="8" pageNumber="94">7+2</td>
<td id="6F465DD0BF48005AB424E252633DFEC2" box="[1130,1303,359,381]" gridcol="4" gridrow="2" pageId="8" pageNumber="94">Ac, 2, 3, 4, M</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E2BE633DFE1E" box="[159,1303,395,417]" gridrow="3" pageId="8" pageNumber="94" rowspan-0="1">
<td id="6F465DD0BF48005AB1DFE2BE67C8FE1E" box="[401,482,395,417]" gridcol="1" gridrow="3" pageId="8" pageNumber="94">16</td>
<td id="6F465DD0BF48005AB206E2BE6503FE1E" box="[584,809,395,417]" gridcol="2" gridrow="3" pageId="8" pageNumber="94">P1, 1a, 2, 2a, 3, 3a, 4, 5</td>
<td id="6F465DD0BF48005AB3F8E2BE622DFE1E" box="[950,1031,395,417]" gridcol="3" gridrow="3" pageId="8" pageNumber="94">4</td>
<td id="6F465DD0BF48005AB424E2BE633DFE1E" box="[1130,1303,395,417]" gridcol="4" gridrow="3" pageId="8" pageNumber="94">P1, 3</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E28F633DFE4C" box="[159,1303,442,499]" gridrow="4" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E28F6739FE4C" box="[159,275,442,499]" gridcol="0" gridrow="4" pageId="8" pageNumber="94">Abd. I</th>
<td id="6F465DD0BF48005AB1DFE28F67C8FE4C" box="[401,482,442,499]" gridcol="1" gridrow="4" pageId="8" pageNumber="94">6 10</td>
<td id="6F465DD0BF48005AB206E28F6503FE4C" box="[584,809,442,499]" gridcol="2" gridrow="4" pageId="8" pageNumber="94">A1, 2, 5 P1, 2, 2a, 3, 4</td>
<td id="6F465DD0BF48005AB3F8E28F622DFE4C" box="[950,1031,442,499]" gridcol="3" gridrow="4" pageId="8" pageNumber="94">3 2</td>
<td id="6F465DD0BF48005AB424E28F633DFE4C" box="[1130,1303,442,499]" gridcol="4" gridrow="4" pageId="8" pageNumber="94">Ac, 2 P1</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E139633DFDF9" box="[159,1303,524,582]" gridrow="5" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E1396739FDF9" box="[159,275,524,582]" gridcol="0" gridrow="5" pageId="8" pageNumber="94">Abd. II–III</th>
<td id="6F465DD0BF48005AB1DFE13967C8FDF9" box="[401,482,524,582]" gridcol="1" gridrow="5" pageId="8" pageNumber="94">6 14</td>
<td id="6F465DD0BF48005AB206E1396503FDF9" box="[584,809,524,582]" gridcol="2" gridrow="5" pageId="8" pageNumber="94">A1, 2, 5 P1, 2, 2a, 3, 4, 4a, 5</td>
<td id="6F465DD0BF48005AB3F8E139622DFDF9" box="[950,1031,524,582]" gridcol="3" gridrow="5" pageId="8" pageNumber="94">3 5</td>
<td id="6F465DD0BF48005AB424E139633DFDF9" box="[1130,1303,524,582]" gridcol="4" gridrow="5" pageId="8" pageNumber="94">Ac, 2 Pc, 1a, 2</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E16A633DFD26" box="[159,1303,607,665]" gridrow="6" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E16A6739FD26" box="[159,275,607,665]" gridcol="0" gridrow="6" pageId="8" pageNumber="94">Abd. IV–VI</th>
<td id="6F465DD0BF48005AB1DFE16A67C8FD26" box="[401,482,607,665]" gridcol="1" gridrow="6" pageId="8" pageNumber="94">6 14</td>
<td id="6F465DD0BF48005AB206E16A6503FD26" box="[584,809,607,665]" gridcol="2" gridrow="6" pageId="8" pageNumber="94">A1, 2, 5 P1, 2, 2a, 3, 4, 4a, 5</td>
<td id="6F465DD0BF48005AB3F8E16A622DFD26" box="[950,1031,607,665]" gridcol="3" gridrow="6" pageId="8" pageNumber="94">3 8</td>
<td id="6F465DD0BF48005AB424E16A633DFD26" box="[1130,1303,607,665]" gridcol="4" gridrow="6" pageId="8" pageNumber="94">Ac, 2 P1, 1a, 2, 3</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E187633DFD77" box="[159,1303,690,712]" gridrow="7" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E1876739FD77" box="[159,275,690,712]" gridcol="0" gridrow="7" pageId="8" pageNumber="94">Abd. VII</th>
<td id="6F465DD0BF48005AB1DFE18767C8FD77" box="[401,482,690,712]" gridcol="1" gridrow="7" pageId="8" pageNumber="94">8 (7)</td>
<td id="6F465DD0BF48005AB206E1876503FD77" box="[584,809,690,712]" gridcol="2" gridrow="7" pageId="8" pageNumber="94">A1*, 2, 4, 5</td>
<td id="6F465DD0BF48005AB3F8E187622DFD77" box="[950,1031,690,712]" gridcol="3" gridrow="7" pageId="8" pageNumber="94">3</td>
<td id="6F465DD0BF48005AB424E187633DFD77" box="[1130,1303,690,712]" gridcol="4" gridrow="7" pageId="8" pageNumber="94">Ac, 2</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E1E3633DFD53" box="[159,1303,726,748]" gridrow="8" pageId="8" pageNumber="94" rowspan-0="1">
<td id="6F465DD0BF48005AB1DFE1E367C8FD53" box="[401,482,726,748]" gridcol="1" gridrow="8" pageId="8" pageNumber="94">16</td>
<td id="6F465DD0BF48005AB206E1E36503FD53" box="[584,809,726,748]" gridcol="2" gridrow="8" pageId="8" pageNumber="94">P1, 1a, 2, 2a, 3, 4, 4a, 5</td>
<td id="6F465DD0BF48005AB3F8E1E3622DFD53" box="[950,1031,726,748]" gridcol="3" gridrow="8" pageId="8" pageNumber="94">8</td>
<td id="6F465DD0BF48005AB424E1E3633DFD53" box="[1130,1303,726,748]" gridcol="4" gridrow="8" pageId="8" pageNumber="94">P1, 1a, 2, 3</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E030633DFCA4" box="[159,1303,773,795]" gridrow="9" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E0306739FCA4" box="[159,275,773,795]" gridcol="0" gridrow="9" pageId="8" pageNumber="94">Abd. VIII</th>
<td id="6F465DD0BF48005AB1DFE03067C8FCA4" box="[401,482,773,795]" gridcol="1" gridrow="9" pageId="8" pageNumber="94">6</td>
<td id="6F465DD0BF48005AB206E0306503FCA4" box="[584,809,773,795]" gridcol="2" gridrow="9" pageId="8" pageNumber="94">A1, 4, 5</td>
<td id="6F465DD0BF48005AB3F8E030622DFCA4" box="[950,1031,773,795]" gridcol="3" gridrow="9" pageId="8" pageNumber="94">4</td>
<td id="6F465DD0BF48005AB424E030633DFCA4" box="[1130,1303,773,795]" gridcol="4" gridrow="9" pageId="8" pageNumber="94">1, 2</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E01C633DFC80" box="[159,1303,809,831]" gridrow="10" pageId="8" pageNumber="94" rowspan-0="1" rowspan-3="1" rowspan-4="1">
<td id="6F465DD0BF48005AB1DFE01C67C8FC80" box="[401,482,809,831]" gridcol="1" gridrow="10" pageId="8" pageNumber="94">16</td>
<td id="6F465DD0BF48005AB206E01C6503FC80" box="[584,809,809,831]" gridcol="2" gridrow="10" pageId="8" pageNumber="94">P1, 2, 2a, 3, 3a, 4, 4a, 5</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E06D633DFCD1" box="[159,1303,856,878]" gridrow="11" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E06D6739FCD1" box="[159,275,856,878]" gridcol="0" gridrow="11" pageId="8" pageNumber="94">Abd. IX</th>
<td id="6F465DD0BF48005AB1DFE06D67C8FCD1" box="[401,482,856,878]" gridcol="1" gridrow="11" pageId="8" pageNumber="94">14</td>
<td id="6F465DD0BF48005AB206E06D6503FCD1" box="[584,809,856,878]" gridcol="2" gridrow="11" pageId="8" pageNumber="94">1, 1a, 2, 2a, 3, 3a, 4</td>
<td id="6F465DD0BF48005AB3F8E06D622DFCD1" box="[950,1031,856,878]" gridcol="3" gridrow="11" pageId="8" pageNumber="94">4</td>
<td id="6F465DD0BF48005AB424E06D633DFCD1" box="[1130,1303,856,878]" gridcol="4" gridrow="11" pageId="8" pageNumber="94">1, 2</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E0B2633DFC22" box="[159,1303,903,925]" gridrow="12" pageId="8" pageNumber="94">
<th id="6F465DD0BF48005AB0D1E0B26739FC22" box="[159,275,903,925]" gridcol="0" gridrow="12" pageId="8" pageNumber="94">Abd. X</th>
<td id="6F465DD0BF48005AB1DFE0B267C8FC22" box="[401,482,903,925]" gridcol="1" gridrow="12" pageId="8" pageNumber="94">12</td>
<td id="6F465DD0BF48005AB206E0B26503FC22" box="[584,809,903,925]" gridcol="2" gridrow="12" pageId="8" pageNumber="94">1, 1a, 2, 2a, 3, 4</td>
<td id="6F465DD0BF48005AB3F8E0B2622DFC22" box="[950,1031,903,925]" gridcol="3" gridrow="12" pageId="8" pageNumber="94">4</td>
<td id="6F465DD0BF48005AB424E0B2633DFC22" box="[1130,1303,903,925]" gridcol="4" gridrow="12" pageId="8" pageNumber="94">1, 2</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E083633DFC73" box="[159,1303,950,972]" gridrow="13" pageId="8" pageNumber="94" rowspan-4="1">
<th id="6F465DD0BF48005AB0D1E0836739FC73" box="[159,275,950,972]" gridcol="0" gridrow="13" pageId="8" pageNumber="94">Abd. XI</th>
<td id="6F465DD0BF48005AB1DFE08367C8FC73" box="[401,482,950,972]" gridcol="1" gridrow="13" pageId="8" pageNumber="94">6</td>
<td id="6F465DD0BF48005AB206E0836503FC73" box="[584,809,950,972]" gridcol="2" gridrow="13" pageId="8" pageNumber="94">1, 3, 4</td>
<td id="6F465DD0BF48005AB3F8E083622DFC73" box="[950,1031,950,972]" gridcol="3" gridrow="13" pageId="8" pageNumber="94">6</td>
</tr>
<tr id="2C9734ACBF48005AB0D1E0D0633DFC44" box="[159,1303,997,1019]" gridrow="14" pageId="8" pageNumber="94" rowspan-2="1" rowspan-4="1">
<th id="6F465DD0BF48005AB0D1E0D06739FC44" box="[159,275,997,1019]" gridcol="0" gridrow="14" pageId="8" pageNumber="94">Abd. XII</th>
<td id="6F465DD0BF48005AB1DFE0D067C8FC44" box="[401,482,997,1019]" gridcol="1" gridrow="14" pageId="8" pageNumber="94">9</td>
<td id="6F465DD0BF48005AB3F8E0D0622DFC44" box="[950,1031,997,1019]" gridcol="3" gridrow="14" pageId="8" pageNumber="94">6</td>
</tr>
</table>
</paragraph>
<paragraph id="921836EEBF48FFADB0D9E73B6441FB9B" blockId="8.[151,619,1038,1060]" box="[151,619,1038,1060]" pageId="8" pageNumber="94">
<tableNote id="6F413760BF48FFADB0D9E73B6441FB9B" box="[151,619,1038,1060]" pageId="8" pageNumber="94" targetBox="[159,1303,194,1019]" targetPageId="8">* Asymmetrical presence of setae in a specimen.</tableNote>
</paragraph>
<paragraph id="921836EEBF48FFADB089E743625AFAFA" blockId="8.[151,1437,1142,2033]" pageId="8" pageNumber="94">
Accessory setae 
<emphasis id="A0D3EAFCBF48FFADB1CCE7436785FB2F" box="[386,431,1142,1168]" italics="true" pageId="8" pageNumber="94">P2a</emphasis>
and 
<emphasis id="A0D3EAFCBF48FFADB1A8E7436439FB2F" box="[486,531,1142,1168]" italics="true" pageId="8" pageNumber="94">P4a</emphasis>
on abdominal tergites I–VI setiform and short, their lengths 4 μm (
<figureCitation id="0A9C2A6BBF48FFADB548E74363BAFB2F" box="[1286,1424,1142,1168]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 33, 34</figureCitation>
). Accessory setae 
<emphasis id="A0D3EAFCBF48FFADB11CE7AF6755FB0B" box="[338,383,1178,1204]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
, 
<emphasis id="A0D3EAFCBF48FFADB1C3E7AF6790FB0B" box="[397,442,1178,1204]" italics="true" pageId="8" pageNumber="94">P2a</emphasis>
and 
<emphasis id="A0D3EAFCBF48FFADB1BEE7AF6437FB0B" box="[496,541,1178,1204]" italics="true" pageId="8" pageNumber="94">P4a</emphasis>
on tergite VII setiform, their lengths 7 μm (
<figureCitation id="0A9C2A6BBF48FFADB447E7AF624CFB0A" box="[1033,1126,1178,1205]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 35</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB43CE7AF62B8FB0A" box="[1138,1170,1178,1205]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">57</figureCitation>
). Seta 
<emphasis id="A0D3EAFCBF48FFADB493E7AF62D6FB0B" box="[1245,1276,1178,1204]" italics="true" pageId="8" pageNumber="94">P3</emphasis>
on abdominal tergites II–VI inserted anteriorly to other setae of 
<emphasis id="A0D3EAFCBF48FFADB285E78A64F6FB67" box="[715,732,1215,1240]" italics="true" pageId="8" pageNumber="94">P</emphasis>
-row (
<figureCitation id="0A9C2A6BBF48FFADB36FE78B65AAFB67" box="[801,896,1214,1240]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 34</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB3C3E78B6586FB66" box="[909,940,1214,1241]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">58</figureCitation>
), 
<emphasis id="A0D3EAFCBF48FFADB38FE78B65CAFB67" box="[961,992,1214,1240]" italics="true" pageId="8" pageNumber="94">P3</emphasis>
on tergites I and VII on the same level with other setae of 
<emphasis id="A0D3EAFCBF48FFADB122E7D66757FB43" box="[364,381,1251,1276]" italics="true" pageId="8" pageNumber="94">P</emphasis>
-row (
<figureCitation id="0A9C2A6BBF48FFADB18FE7D76462FB42" box="[449,584,1250,1277]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 33, 35</figureCitation>
). Abdominal tergite VII anteriorly with two cuticular lines crossed in medial part of tergite (
<figureCitation id="0A9C2A6BBF48FFADB10EE63367B9FA9E" box="[320,403,1286,1313]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 35</figureCitation>
). Pores 
<emphasis id="A0D3EAFCBF48FFADB1A0E6326431FA9F" box="[494,539,1287,1312]" italics="true" pageId="8" pageNumber="94">psm</emphasis>
present on abdominal tergites I–VII between setae 
<emphasis id="A0D3EAFCBF48FFADB411E6336254FA9F" box="[1119,1150,1286,1312]" italics="true" pageId="8" pageNumber="94">P1</emphasis>
and 
<emphasis id="A0D3EAFCBF48FFADB4FBE63362FEFA9F" box="[1205,1236,1286,1312]" italics="true" pageId="8" pageNumber="94">P2</emphasis>
, and close to seta 
<emphasis id="A0D3EAFCBF48FFADB0D9E61F66EFFAFB" box="[151,197,1322,1348]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
on tergite VII (
<figureCitation id="0A9C2A6BBF48FFADB13CE61F67EFFAFA" box="[370,453,1322,1349]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">Fig. 57</figureCitation>
), 
<emphasis id="A0D3EAFCBF48FFADB194E61F67DAFAFB" box="[474,496,1322,1348]" italics="true" pageId="8" pageNumber="94">al</emphasis>
pores on tergites II–VII, 
<emphasis id="A0D3EAFCBF48FFADB342E61F6507FAFB" box="[780,813,1322,1348]" italics="true" pageId="8" pageNumber="94">psl</emphasis>
on tergite VII (
<figureCitation id="0A9C2A6BBF48FFADB394E61F6249FAFA" box="[986,1123,1322,1349]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 33–35</figureCitation>
).
</paragraph>
<paragraph id="921836EEBF48FFADB089E67B6206F9A3" blockId="8.[151,1437,1142,2033]" pageId="8" pageNumber="94">
Abdominal legs with 4, 2, 2 setae. Subapical seta on second and third pairs of abdominal legs longer than apical lateral seta, the length of subapical seta 23 μm and apical lateral 14 μm (
<figureCitation id="0A9C2A6BBF48FFADB458E647625EFA33" box="[1046,1140,1394,1420]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 41</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB4CFE647628BFA32" box="[1153,1185,1394,1421]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">59</figureCitation>
). Accessory setae 
<emphasis id="A0D3EAFCBF48FFADB521E64763B7FA33" box="[1391,1437,1394,1420]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
on abdominal sternites II–VII setiform and longer, than on abdominal tergites I–VII (9 μm) (
<figureCitation id="0A9C2A6BBF48FFADB49BE6A3634BFA0F" box="[1237,1377,1430,1456]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 41–43</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB53EE6A363BAFA0F" box="[1392,1424,1430,1456]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">60</figureCitation>
). Abdominal sternite VI with a connecting cuticular line (
<figureCitation id="0A9C2A6BBF48FFADB350E68F655EFA6B" box="[798,884,1466,1492]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 42</figureCitation>
), sternite VII with two parallel connecting lines (
<figureCitation id="0A9C2A6BBF48FFADB0D1E6EB66DEFA47" box="[159,244,1502,1528]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 43</figureCitation>
). Sternites I–IV without pores (
<figureCitation id="0A9C2A6BBF48FFADB21AE6EB64F6FA47" box="[596,732,1502,1528]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 40, 41</figureCitation>
). Abdominal sternites V–VI with a pair of 
<emphasis id="A0D3EAFCBF48FFADB4F5E6EA62D9FA47" box="[1211,1267,1503,1528]" italics="true" pageId="8" pageNumber="94">spsm</emphasis>
pores, situated between bases of setae 
<emphasis id="A0D3EAFCBF48FFADB1D2E53767E0F9A3" box="[412,458,1538,1564]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
and 
<emphasis id="A0D3EAFCBF48FFADB24EE5376435F9A3" box="[512,543,1538,1564]" italics="true" pageId="8" pageNumber="94">P2</emphasis>
(
<figureCitation id="0A9C2A6BBF48FFADB260E53764A8F9A3" box="[558,642,1538,1564]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 42</figureCitation>
), sternite VII with 
<emphasis id="A0D3EAFCBF48FFADB318E53665A9F9A3" box="[854,899,1539,1564]" italics="true" pageId="8" pageNumber="94">spm</emphasis>
pore (
<figureCitation id="0A9C2A6BBF48FFADB385E5376235F9A3" box="[971,1055,1538,1564]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 43</figureCitation>
).
</paragraph>
<paragraph id="921836EEBF48FFADB089E5136744F94A" blockId="8.[151,1437,1142,2033]" pageId="8" pageNumber="94">
Abdominal segment VIII with striate band reduced to two lines and without parallel-sided striae, but with hook-shaped design (a few oblique, sinuate striae) (
<figureCitation id="0A9C2A6BBF48FFADB2DBE57F6536F9DB" box="[661,796,1610,1636]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Figs. 36, 44</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB369E57F655AF9DB" box="[807,880,1610,1637]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">57, 61</figureCitation>
). Pore 
<emphasis id="A0D3EAFCBF48FFADB3F0E57E65C1F9DB" box="[958,1003,1611,1636]" italics="true" pageId="8" pageNumber="94">psm</emphasis>
with several accompanying teeth (
<figureCitation id="0A9C2A6BBF48FFADB521E57F669DF937" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 36</figureCitation>
). Posterior margin of abdominal sternite VIII and laterotergites smooth (
<figureCitation id="0A9C2A6BBF48FFADB396E55B6201F937" box="[984,1067,1646,1672]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 44</figureCitation>
). Comb on tergite VIII with 9–10 short teeth (
<figureCitation id="0A9C2A6BBF48FFADB152E5A76753F913" box="[284,377,1682,1708]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="8" pageNumber="94">Figs. 30</figureCitation>
, 
<figureCitation id="0A9C2A6BBF48FFADB1CBE5A7678FF913" box="[389,421,1682,1708]" captionStart="FIGURES 57–62" captionStartId="11.[151,264,1693,1717]" captionTargetBox="[151,1436,181,1669]" captionTargetId="figure-17@11.[151,1436,181,1669]" captionTargetPageId="11" captionText="FIGURES 57–62. Amphientulus markstivensi sp. nov. 57) Setae P1a and P2a on abdominal tergite VII (arrow indicates pore psm) and striate band on abdominal tergite VIII. 58) Position of seta P3 on abdominal tergites VI and VII. 59) Second and third pairs of abdominal legs. 60) Seta P1a on abdominal sternite VII. 61) Striate band on abdominal sternite VIII. 62) Comb on abdominal tergite VIII." figureDoi="http://doi.org/10.5281/zenodo.11233488" httpUri="https://zenodo.org/record/11233488/files/figure.png" pageId="8" pageNumber="94">62</figureCitation>
). Abdominal sternites IX–XII with smooth hind margin and with setae of different lengths, medial setae half the length of lateral ones (
<figureCitation id="0A9C2A6BBF48FFADB231E58364F9F96F" box="[639,723,1718,1744]" captionStart="FIGURES 31–44" captionStartId="7.[151,264,1769,1793]" captionTargetBox="[294,1292,181,1746]" captionTargetId="figure-17@7.[294,1292,181,1746]" captionTargetPageId="7" captionText="FIGURES 31–44. Amphientulus markstivensi sp. nov. holotype (ISEA 6652). 31) Pronotum and mesonotum, right side. 32) Lateral part of metanotum. 33) Tergite I, right side. 34) Tergite VI, right part. 35) Abdominal tergite VII, right part. 36) Abdominal tergite VIII, right part. 37) Anterior part of prosternum. 38) Antero-lateral part of mesosternum. 39) Anterolateral part of metasternum. 40) Part of abdominal sternite I. 41) Abdominal sternite III with abdominal leg. 42) Abdominal sternite VI. 43) Abdominal sternite VII. 44) Abdominal sternites VIII–X. Arrows indicate pores on abdominal tergites and sternites (al = anterolateral, sl = sublateral, psm = posterosubmedial, psl = posterosublateral, sc = sternal central, spsm = sternal posterosubmedial, spm = sternal posteromedial). Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233468" httpUri="https://zenodo.org/record/11233468/files/figure.png" pageId="8" pageNumber="94">Fig. 44</figureCitation>
). Medial pore on dorsal lobe of segment XII, one pair of sternal anterolateral pores.
</paragraph>
<paragraph id="921836EEBF48FFADB089E5CB65A9F8A7" blockId="8.[151,1437,1142,2033]" box="[199,899,1790,1817]" pageId="8" pageNumber="94">
Female squama genitalis with short, forked acrostyli (
<figureCitation id="0A9C2A6BBF48FFADB36CE5CB655CF8A7" box="[802,886,1790,1816]" captionStart="FIGURES 21–30" captionStartId="6.[151,264,1285,1309]" captionTargetBox="[151,1436,181,1261]" captionTargetId="figure-17@6.[151,1436,181,1261]" captionTargetPageId="6" captionText="FIGURES 21–30. Amphientulus markstivensi sp. nov., holotype (ISEA 6652). 21) Pseudoculus with setae l3 and sd4. 22) Cephalic seta sd5. 23) Maxillary palp. 24) Labial palp. 25) Maxillary gland. 26) Female squama genitalis. 27) Foretarsus, exterior view. 28) Foretarsus, interior view. 29) Foretarsal sensillum t3. 30) Comb. Arrows indicate pores. Scale bars: 20 µm." figureDoi="http://doi.org/10.5281/zenodo.11233466" httpUri="https://zenodo.org/record/11233466/files/figure.png" pageId="8" pageNumber="94">Fig. 26</figureCitation>
).
</paragraph>
<paragraph id="921836EEBF48FFADB089E4176211F83B" blockId="8.[151,1437,1142,2033]" pageId="8" pageNumber="94">
Body measurements (
<specimenCount id="84A1FD67BF48FFADB1F4E4176402F883" box="[442,552,1826,1852]" count="3" pageId="8" pageNumber="94" type="female">3 females</specimenCount>
) (in μm): body length 1150; head 135–140; pseudoculus 6; lever 2–3; posterior part of maxillary gland 22; head setae 
<emphasis id="A0D3EAFCBF48FFADB20CE4736474F8DF" box="[578,606,1862,1888]" italics="true" pageId="8" pageNumber="94">d7</emphasis>
15, 
<emphasis id="A0D3EAFCBF48FFADB2C1E473649FF8DF" box="[655,693,1862,1888]" italics="true" pageId="8" pageNumber="94">sd7</emphasis>
10, 
<emphasis id="A0D3EAFCBF48FFADB2A8E47364D6F8DF" box="[742,764,1862,1888]" italics="true" pageId="8" pageNumber="94">l5</emphasis>
6; pronotal setae 
<emphasis id="A0D3EAFCBF48FFADB38FE47365E5F8DF" box="[961,975,1862,1888]" italics="true" pageId="8" pageNumber="94">1</emphasis>
28, 
<emphasis id="A0D3EAFCBF48FFADB3B1E4736227F8DF" box="[1023,1037,1862,1888]" italics="true" pageId="8" pageNumber="94">2</emphasis>
13–14, mesonotal setae 
<emphasis id="A0D3EAFCBF48FFADB56EE4736315F8DF" box="[1312,1343,1862,1888]" italics="true" pageId="8" pageNumber="94">P1</emphasis>
21, 
<emphasis id="A0D3EAFCBF48FFADB521E47363B7F8DF" box="[1391,1437,1862,1888]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
2, 
<emphasis id="A0D3EAFCBF48FFADB0FDE45F66F8F83B" box="[179,210,1898,1924]" italics="true" pageId="8" pageNumber="94">P2</emphasis>
30–31; metanotal setae 
<emphasis id="A0D3EAFCBF48FFADB1ACE45F643AF83B" box="[482,528,1898,1924]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
4; foretarsus 80, claw 26, empodial appendage 2.
</paragraph>
</subSubSection>
<subSubSection id="DABD6565BF48FFAFB089E4BB674CFEDB" lastPageId="10" lastPageNumber="96" pageId="8" pageNumber="94" type="discussion">
<paragraph id="921836EEBF48FFACB089E4BB64BBFF6A" blockId="8.[151,1437,1142,2033]" lastBlockId="9.[151,1437,151,213]" lastPageId="9" lastPageNumber="95" pageId="8" pageNumber="94">
<emphasis id="A0D3EAFCBF48FFADB089E4BB6711F817" bold="true" box="[199,315,1934,1960]" pageId="8" pageNumber="94">Remarks.</emphasis>
The species is characterized by the absence of 
<emphasis id="A0D3EAFCBF48FFADB32FE4BB65A5F817" box="[865,911,1934,1960]" italics="true" pageId="8" pageNumber="94">P1a</emphasis>
setae on abdominal tergites I–VI and 
<emphasis id="A0D3EAFCBF48FFADB504E4BB635DF817" box="[1354,1399,1934,1960]" italics="true" pageId="8" pageNumber="94">P3a</emphasis>
on tergite VII, presence of 
<emphasis id="A0D3EAFCBF48FFADB1E8E48767F8F873" box="[422,466,1970,1996]" italics="true" pageId="8" pageNumber="94">P4a</emphasis>
setae on tergites II–VI, sensilliform shape of accessory setae on abdominal segments and two 
<emphasis id="A0D3EAFCBF48FFADB0B6E4E26723F84F" box="[248,265,2007,2032]" italics="true" pageId="8" pageNumber="94">P</emphasis>
setae on abdominal sternite I. 
<specimenCount id="84A1FD67BF48FFADB211E4E26529F84E" box="[607,771,2007,2033]" count="2" pageId="8" pageNumber="94" type="generic" typeStatus="paratype">Two paratypes</specimenCount>
(ISEA 6653, 6654) have 
<emphasis id="A0D3EAFCBF48FFADB46CE4E36268F84F" box="[1058,1090,2006,2032]" italics="true" pageId="8" pageNumber="94">A1</emphasis>
setae on abdominal tergite VII and posterior setae on abdominal sternite VIII, but the asymmetrical presence of these setae suggests a direction toward reduction of these setae in speciation.
</paragraph>
<caption id="C6D86666BF49FFACB0D9E5DE63A9F898" ID-DOI="http://doi.org/10.5281/zenodo.11233470" ID-Zenodo-Dep="11233470" httpUri="https://zenodo.org/record/11233470/files/figure.png" pageId="9" pageNumber="95" startId="9.[151,264,1771,1795]" targetBox="[151,1436,237,1748]" targetPageId="9" targetType="figure">
<paragraph id="921836EEBF49FFACB0D9E5DE63A9F898" blockId="9.[151,1437,1771,1831]" pageId="9" pageNumber="95">
<emphasis id="A0D3EAFCBF49FFACB0D9E5DE6779F8BC" bold="true" box="[151,339,1771,1795]" pageId="9" pageNumber="95">FIGURES 45–50.</emphasis>
<taxonomicName id="55A74D6DBF49FFACB119E5DE644DF8BC" authority="Shrubovych &amp; D’Haese, 2024" authorityName="Shrubovych &amp; D’Haese" authorityYear="2024" box="[343,615,1771,1795]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="9" pageNumber="95" phylum="Arthropoda" rank="species" species="markstivensi" status="sp. nov.">
<emphasis id="A0D3EAFCBF49FFACB119E5DE644DF8BC" box="[343,615,1771,1795]" italics="true" pageId="9" pageNumber="95">Amphientulus markstivensi</emphasis>
</taxonomicName>
<emphasis id="A0D3EAFCBF49FFACB222E5DE6491F8BC" bold="true" box="[620,699,1771,1795]" pageId="9" pageNumber="95">
<taxonomicNameLabel id="BBE05787BF49FFACB222E5DE6491F8BC" box="[620,699,1771,1795]" pageId="9" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
45) Habitus. 46) Head. 47) Pseudoculus with setae 
<emphasis id="A0D3EAFCBF49FFACB4F4E5DE62E4F8BC" box="[1210,1230,1771,1795]" italics="true" pageId="9" pageNumber="95">l3</emphasis>
and 
<emphasis id="A0D3EAFCBF49FFACB4B2E5DE6334F8BC" box="[1276,1310,1771,1795]" italics="true" pageId="9" pageNumber="95">sd4</emphasis>
(fp = frontal pore). 48) Cephalic seta 
<emphasis id="A0D3EAFCBF49FFACB1C1E43A6798F898" box="[399,434,1807,1831]" italics="true" pageId="9" pageNumber="95">sd5</emphasis>
. 49) Maxillary palp (d = dorsal sensillum, l = lateral sensillum). 50) Labial palp (s = sensillum).
</paragraph>
</caption>
<paragraph id="921836EEBF4AFFAFB089E3A2674CFEDB" blockId="10.[151,1437,151,357]" pageId="10" pageNumber="96">
All 
<taxonomicName id="55A74D6DBF4AFFAFB0BDE3A267DEFF0E" box="[243,500,151,177]" pageId="10" pageNumber="96">
<emphasis id="A0D3EAFCBF4AFFAFB0BDE3A267B0FF0E" box="[243,410,151,177]" italics="true" pageId="10" pageNumber="96">Tasmanentulus</emphasis>
species
</taxonomicName>
(
<taxonomicName id="55A74D6DBF4AFFAFB24BE3A2648AFF0F" baseAuthorityName="Tuxen" baseAuthorityYear="1967" box="[517,672,151,176]" class="Protura" family="Acerentomidae" genus="Tasmanentulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="tasmanicus">
<emphasis id="A0D3EAFCBF4AFFAFB24BE3A2648AFF0F" box="[517,672,151,176]" italics="true" pageId="10" pageNumber="96">T. tasmanicus</emphasis>
</taxonomicName>
, 
<taxonomicName id="55A74D6DBF4AFFAFB2FEE3A2653FFF0E" baseAuthorityName="Tuxen" baseAuthorityYear="1967" box="[688,789,151,177]" class="Protura" family="Acerentomidae" genus="Tasmanentulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="A0D3EAFCBF4AFFAFB2FEE3A2653FFF0E" box="[688,789,151,177]" italics="true" pageId="10" pageNumber="96">T. similis</emphasis>
</taxonomicName>
, 
<taxonomicName id="55A74D6DBF4AFFAFB36BE3A265ECFF0E" authorityName="Tuxen" authorityYear="1967" box="[805,966,151,177]" class="Protura" family="Acerentomidae" genus="Australentulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="10" pageNumber="96" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="A0D3EAFCBF4AFFAFB36BE3A265ECFF0E" box="[805,966,151,177]" italics="true" pageId="10" pageNumber="96">T. intermedius</emphasis>
</taxonomicName>
) are characterized by possession of setae 
<emphasis id="A0D3EAFCBF4AFFAFB0D9E38E66EFFF6A" box="[151,197,187,213]" italics="true" pageId="10" pageNumber="96">P1a</emphasis>
on abdominal tergites I–VI, setiform shape of accessory setae on the abdominal segments, 6 setae on abdominal sternite VIII and long foretarsal sensillum 
<emphasis id="A0D3EAFCBF4AFFAFB233E3EA64B9FF46" box="[637,659,223,249]" italics="true" pageId="10" pageNumber="96">t3</emphasis>
with cylindrical shape (
<bibRefCitation id="F6364B1FBF4AFFAFB3E8E3EA6218FF46" author="Tuxen, S. L." box="[934,1074,223,249]" pageId="10" pageNumber="96" pagination="285 - 295" refId="ref13722" refString="Tuxen, S. L. (1984) Brasilidia and a new genus and species of Protura (Insecta) from Argentina. Together with an improved generic key to Acerentomidae. Revue d'Ecologie et de Biologie du Sol, 21, 285 - 295." type="journal article" year="1984">Tuxen 1984</bibRefCitation>
, 
<bibRefCitation id="F6364B1FBF4AFFAFB470E3EA6250FF46" author="Tuxen, S. L." box="[1086,1146,223,249]" pageId="10" pageNumber="96" refId="ref13769" refString="Tuxen, S. L. (1986) Protura (Insecta). Fauna of New Zealand. 9 (1985). Department of Scientific and Industrial Research, Wellington, 50 pp." type="book" year="1986">1986</bibRefCitation>
). Therefore, we place the species in 
<taxonomicName id="55A74D6DBF4AFFAFB140E2366782FEA2" authorityName="Tuxen" authorityYear="1981" box="[270,424,259,285]" class="Protura" family="Acerentomidae" genus="Amphientulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Protura" pageId="10" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="A0D3EAFCBF4AFFAFB140E2366782FEA2" box="[270,424,259,285]" italics="true" pageId="10" pageNumber="96">Amphientulus</emphasis>
</taxonomicName>
due to the absence of 
<emphasis id="A0D3EAFCBF4AFFAFB2E4E23664F2FEA2" box="[682,728,259,285]" italics="true" pageId="10" pageNumber="96">P1a</emphasis>
setae on abdominal tergites I–VI, sensilliform accessory setae on abdominal segments, jar-shaped foretarsal sensillum 
<emphasis id="A0D3EAFCBF4AFFAFB347E2126535FEFE" box="[777,799,295,321]" italics="true" pageId="10" pageNumber="96">t3</emphasis>
and completely reduced striate band to two lines without any trace of striae.
</paragraph>
</subSubSection>
</treatment>
</document>