1317 lines
106 KiB
XML
1317 lines
106 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.96.51962" ID-GBIF-Dataset="a134fec9-ee97-4d3e-ae3f-eb295c3267db" ID-GBIF-Taxon="164501238" ID-Pensoft-Pub="1860-0743-2-275" ID-Pensoft-UUID="FC312E66E3EA52F4A7706C2C191A8621" ID-ZooBank="4A3A5AE9D26340A08621430C7822CFF3" ModsDocID="1860-0743-2-275" checkinTime="1591887254079" checkinUser="pensoft" docAuthor="Viana, Sarah T. F. L. & Carvalho, Marcelo R. de" docDate="2020" docId="E4AF6EA097C05E95813FAAD0C44B8851" docLanguage="en" docName="ZoosystEvol 96(2): 275-311" docOrigin="Zoosystematics and Evolution 96 (2)" docSource="http://dx.doi.org/10.3897/zse.96.51962" docTitle="Squalus mitsukurii Jordan & Snyder 1903" docType="treatment" docVersion="4" id="FC312E66E3EA52F4A7706C2C191A8621" lastPageNumber="275" masterDocId="FC312E66E3EA52F4A7706C2C191A8621" masterDocTitle="Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited" masterLastPageNumber="311" masterPageNumber="275" pageId="0" pageNumber="275" updateTime="1634711231051" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Viana, Sarah T. F. L.</mods:namePart>
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<mods:affiliation>South African Institute for Aquatic Biodiversity, Private Bag 1015 Grahamstown, 6140, South Africa & Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo. Rua do Matao, Travessa 14, no 101, CEP 05508 - 090, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-6905-9292</mods:nameIdentifier>
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<mods:nameIdentifier type="email">stviana@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Carvalho, Marcelo R. de</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo. Rua do Matao, Travessa 14, no 101, CEP 05508 - 090, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>96</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>275</mods:start>
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<mods:end>311</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.96.51962</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.51962</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-275</mods:identifier>
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<mods:identifier type="ZooBank">4A3A5AE9D26340A08621430C7822CFF3</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FC312E66E3EA52F4A7706C2C191A8621</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164501238" LSID="urn:lsid:plazi:treatment:E4AF6EA097C05E95813FAAD0C44B8851" httpUri="http://treatment.plazi.org/id/E4AF6EA097C05E95813FAAD0C44B8851" lastPageNumber="275" pageId="0" pageNumber="275">
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<subSubSection pageId="0" pageNumber="275" type="nomenclature">
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName LSID="E4AF6EA0-97C0-5E95-813F-AAD0C44B8851" authority="Jordan & Snyder, 1903" authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">Squalus mitsukurii Jordan & Snyder, 1903</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Upper (A, C, E, G, I) and lower (B, D, F, H, J) teeth of S. shiraii sp. nov. (A, B; HUMZ 80329), S. mitsukurii (C, D; HUMZ 33680), S. japonicus (E, F; HUMZ 189737), S. brevirostris (G, H; HUMZ 189762) and S. suckleyi (I, J; NSMT-P 92640). Scale bars: 1 mm." figureDoi="10.3897/zse.96.51962.figure3" httpUri="https://binary.pensoft.net/fig/420111" pageId="0" pageNumber="275">, 3C, D</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron microscopy of the dermal denticles. Squalus shiraii sp. nov. (A, B): HUMZ 149389 (holotype), adult male, 590 mm TL. S. formosus (C, D): CSIRO H- 6816 - 01 (holotype), adult male, 720 mm TL. S. mitsukurii (E, F): HUMZ 102987, adult male, 970 mm TL. S. japonicus (G, H): NSMT-P 44380 (neotype), adult male, 645 mm TL (G); NSMT-P 91127, juvenile female, 410 mm TL (H). S. brevirostris (I, J): NSMT-P 47378, adult female, 600 mm TL (I); KAUM-I 187, adult female, 578 mm TL (J). S. suckleyi (K, L): NSMT P- 92640, adult female, 970 mm TL. Scale bars: 50 μm (B); 100 μm (D, F, L); 200 μm (A, C, E, G-K)." figureDoi="10.3897/zse.96.51962.figure7" httpUri="https://binary.pensoft.net/fig/420115" pageId="0" pageNumber="275">, 7E, F</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of the North-western Pacific Ocean, showing the geographical distribution of Squalus species. A: S. shiraii sp. nov. (red); B: S. mitsukurii (orange); C: S. japonicus (brown); D: S. brevirostris (yellow); E: S. acutirostris (green); F: S. suckleyi (pink). Star: holotype / neotype; Dotted circle: paratypes; Circle: other material." figureDoi="10.3897/zse.96.51962.figure8" httpUri="https://binary.pensoft.net/fig/420116" pageId="0" pageNumber="275">, 8B</figureCitation>
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;
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<figureCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. External measurements of S. mitsukurii expressed as percentage of total length (% TL). Values of the paratype of S. mitsukurii (SU 7184, identified as S. acanthias) and specimens of S. acutirostris are also provided. TL is expressed in millimeters. H: holotype; P: paratype; N: number of specimens; x: mean; SD: standard deviation." pageId="0" pageNumber="275">Tables 2</figureCitation>
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,
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<figureCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Meristic data for species of Squalus from Japan. H: holotype; P: paratypes; NT: neotype; N: number of specimens; x; mean; m: mode; SD: standard deviation." pageId="0" pageNumber="275">6 Longnose spurdog; Futo-tsunozame (Japanese)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="reference_group">
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">Squalus mitsukurii</taxonomicName>
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<bibRefCitation author="Jordan, DS" journalOrPublisher="Annotationes Zoologicae Japonenses" pageId="0" pageNumber="275" pagination="31 - 159" refId="B43" refString="Jordan, DS, Snyder, JO, 1901. A preliminary check list of the fishes of Japan. Annotationes Zoologicae Japonenses 3: 31 - 159" title="A preliminary check list of the fishes of Japan." volume="3" year="1901">Jordan and Snyder 1901</bibRefCitation>
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: 129 (name only);
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<bibRefCitation DOI="https://doi.org/10.5479/si.00963801.26-1324.593" author="Jordan, DS" editor="Jordan, DS" journalOrPublisher="Proceedings of the United States National Museum 26 (1324)" pageId="0" pageNumber="275" pagination="593 - 674" refId="B44" refString="Jordan, DS, Snyder, JO, 1903. Squalus. In: Jordan, DS, Fowler, HW, Eds., A review of the Elasmobranchiate fishes of Japan. Proceedings of the United States National Museum 26(1324): 593 - 674, DOI: https://doi.org/10.5479/si.00963801.26-1324.593" title="Squalus." url="https://doi.org/10.5479/si.00963801.26-1324.593" volumeTitle="A review of the Elasmobranchiate fishes of Japan." year="1903">Jordan and Snyder 1903</bibRefCitation>
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: 629; fig. 3 (original description, illustration; type by original designation; type locality: Misaki, Japan);
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<bibRefCitation author="Zhu, Y" journalOrPublisher="Nature of Kagoshima" pageId="0" pageNumber="275" refId="B101" refString="Zhu, Y, 1960. Classification of Chondrichthyes of China. Science Press, Beijing, China." title="Classification of Chondrichthyes of China. Science Press, Beijing, China." year="1960">Zhu 1960</bibRefCitation>
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: 107, 109-110 (description; Northwest Pacific Ocean);
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<bibRefCitation author="Zhu, Y" journalOrPublisher="Oceanologia et Limnologia Sinica" pageId="0" pageNumber="275" pagination="283 - 286" refId="B102" refString="Zhu, Y, Meng, QW, Li, S, 1984. Description of a new species of Squalidae of China. Oceanologia et Limnologia Sinica 15 (4): 283 - 286" title="Description of a new species of Squalidae of China." volume="15" year="1984">Zhu et al. 1984</bibRefCitation>
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: 284 (cited; Japan);
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<bibRefCitation author="Parin, N" journalOrPublisher="Journal of Ichthyology" pageId="0" pageNumber="275" pagination="43 - 50" refId="B70" refString="Parin, N, 1987. Species of spiny dogfish of genus Squalus, living on Southeastern Pacific Ocean Seamounts. Journal of Ichthyology 27 (5): 43 - 50" title="Species of spiny dogfish of genus Squalus, living on Southeastern Pacific Ocean Seamounts." volume="27" year="1987">Parin 1987</bibRefCitation>
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: 48 (cited; Japan);
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<bibRefCitation author="Siming, D" journalOrPublisher="Xue Lin Publishing House, Shangai" pageId="0" pageNumber="275" refId="B77" refString="Siming, D, Guoqiang, X, Hongxi, Z, 1988. Chondrichthyes. In: Jinao S, Yuanquan C (Eds) A general environmental feature of the deep water of East China Sea. Xue Lin Publishing House, Shangai" title="Chondrichthyes. In: Jinao S, Yuanquan C (Eds) A general environmental feature of the deep water of East China Sea." year="1988">Siming et al. 1988</bibRefCitation>
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: 1 (listed; Northwest Pacific Ocean);
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<bibRefCitation author="Compagno, LJV" editor="Carpenter, KE" journalOrPublisher="The living marine resources of the western central pacific. Cephalopods, crustaceans, holothurans and sharks 2. FAO, Senckenbergiana Biologica, Rome" pageId="0" pageNumber="275" pagination="687 - 1396" refId="B16" refString="Compagno, LJV, Niem, VH, 1998. Several family accounts. In: Carpenter, KE, Niem, VH, Eds., FAO species identification guide for fishery purposes. The living marine resources of the western central pacific. Cephalopods, crustaceans, holothurans and sharks 2. FAO, Senckenbergiana Biologica, Rome: 687 - 1396" title="Several family accounts." volumeTitle="FAO species identification guide for fishery purposes." year="1998">Compagno and Niem 1998</bibRefCitation>
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(in part): 1230 (cited; Northwest Pacific Ocean);
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<bibRefCitation author="Zhu, Y" journalOrPublisher="Oceanologia et Limnologia Sinica" pageId="0" pageNumber="275" refId="B103" refString="Zhu, Y, Meng, QW, 2001. Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." title="Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." year="2001">Zhu and Meng 2001</bibRefCitation>
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: 311, 319-321 (cited, description; Northwest Pacific Ocean);
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<bibRefCitation author="Compagno, LJV" editor="Carpenter, KE" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" pagination="357 - 505" refId="B13" refString="Compagno, LJV, 2002. Sharks. In: Carpenter, KE, Ed., The living marine resources of the Western Central Atlantic. Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome: 357 - 505" title="Sharks." volumeTitle="The living marine resources of the Western Central Atlantic." year="2002">Compagno 2002</bibRefCitation>
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(in part): 381, 385 (listed, description; Central Pacific Ocean);
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<bibRefCitation author="Nakabo, T" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" refId="B61" refString="Nakabo, T, 2002. Fishes of Japan with pictorial keys to the species, English edition. Tokai University Press, Tokyo." title="Fishes of Japan with pictorial keys to the species, English edition. Tokai University Press, Tokyo." year="2002">Nakabo 2002</bibRefCitation>
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: 156 (listed; Tohoku District, Ryukyu Islands, Okinawa Trough, South China Sea);
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<bibRefCitation author="Compagno, LJV" journalOrPublisher="Vol. 1. Introduction, molluscs, crustaceans, hagfishes, sharks, batoid fishes and chimaeras. FAO Species identification guide for fishery purposes and American Society of Ichthyologists Special Publication 5. FAO, Rome" pageId="0" pageNumber="275" refId="B14" refString="Compagno, LJV, Dando, M, Fowler, S, 2005a. Sharks of the World. Princeton University Press, New Jersey." title="Sharks of the World. Princeton University Press, New Jersey." year="2005 a">Compagno et al. 2005a</bibRefCitation>
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(in part): 78 (description; Japan to Taiwan);
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<bibRefCitation author="Nakabo, T" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" refId="B62" refString="Nakabo, T, 2013. Fishes of Japan with pictorial keys to the species. Tokai University Press, Tokyo." title="Fishes of Japan with pictorial keys to the species. Tokai University Press, Tokyo." year="2013">Nakabo 2013</bibRefCitation>
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: 196 (listed; Tohoku District, Ryukyu Islands, Okinawa Trough, South China Sea).
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</paragraph>
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<paragraph pageId="0" pageNumber="275">
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<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus blainvillei" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="blainvillei">Squalus blainvillei</taxonomicName>
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:
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<bibRefCitation author="Zhu, Y" journalOrPublisher="Oceanologia et Limnologia Sinica" pageId="0" pageNumber="275" refId="B103" refString="Zhu, Y, Meng, QW, 2001. Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." title="Faina Sinica - Cyclostomata, Chondrichthyes. Science Press, Beijing." year="2001">Zhu and Meng 2001</bibRefCitation>
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: 311, 315-316 (cited, description; Japan).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="275" type="type material">
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<paragraph pageId="0" pageNumber="275">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="275">
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AMNH 8822 (paratype), female, 248 mm TL; SU 7184 (paratype, identified as
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<taxonomicName genus="S." lsidName="S. acanthias" pageId="0" pageNumber="275" rank="species" species="acanthias">
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<emphasis italics="true" pageId="0" pageNumber="275">S. acanthias</emphasis>
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</taxonomicName>
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), male, 277 mm TL; SU 7748 (paratype), two females, 240-243 mm TL, male, 247 mm TL (All embryo paratypes taken from the holotype); SU 12793 (holotype), adult female, 710 mm TL, Honshu Island, Misaki, Japan, 35.159430°, 139.493865°, 14104 m depth (uncertain, according to CAS), collectors D. S. Jordan and J.O. Snyder in 1900; SU 12794 (paratype), adult male, 770 mm TL, data same as holotype.
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</paragraph>
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<caption ID-Table-UUID="0C0E1FD3168E91C13DD3E8840C980F39" httpUri="http://table.plazi.org/id/0C0E1FD3168E91C13DD3E8840C980F39" pageId="0" pageNumber="275" start="Table 2" startId="T2">
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<paragraph pageId="0" pageNumber="275">
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<emphasis bold="true" pageId="0" pageNumber="275">Table 2.</emphasis>
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External measurements of
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<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
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<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
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</taxonomicName>
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expressed as percentage of total length (% TL). Values of the paratype of
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<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
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<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
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</taxonomicName>
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(SU 7184, identified as
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<taxonomicName genus="S." lsidName="S. acanthias" pageId="0" pageNumber="275" rank="species" species="acanthias">
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<emphasis italics="true" pageId="0" pageNumber="275">S. acanthias</emphasis>
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</taxonomicName>
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) and specimens of
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<taxonomicName genus="S." lsidName="S. acutirostris" pageId="0" pageNumber="275" rank="species" species="acutirostris">
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<emphasis italics="true" pageId="0" pageNumber="275">S. acutirostris</emphasis>
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</taxonomicName>
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are also provided. TL is expressed in millimeters. H: holotype; P: paratype; N: number of specimens;
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<emphasis italics="true" pageId="0" pageNumber="275">x</emphasis>
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: mean; SD: standard deviation.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="275">
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<table pageId="0" pageNumber="275">
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<tr pageId="0" pageNumber="275">
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<th colspan="1" isEmpty="true" pageId="0" pageNumber="275" rowspan="2">-</th>
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<th colspan="5" pageId="0" pageNumber="275" rowspan="1">
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<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
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<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
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</taxonomicName>
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</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">
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<taxonomicName genus="S." lsidName="S. acanthias" pageId="0" pageNumber="275" rank="species" species="acanthias">
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<emphasis italics="true" pageId="0" pageNumber="275">S. acanthias</emphasis>
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</taxonomicName>
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</th>
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<th colspan="3" pageId="0" pageNumber="275" rowspan="1">
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<taxonomicName genus="S." lsidName="S. acutirostris" pageId="0" pageNumber="275" rank="species" species="acutirostris">
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<emphasis italics="true" pageId="0" pageNumber="275">S. acutirostris</emphasis>
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</taxonomicName>
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</th>
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</tr>
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<tr pageId="0" pageNumber="275">
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">H</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">N</th>
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<th colspan="1" isEmpty="true" pageId="0" pageNumber="275" rowspan="1">-</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">
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<emphasis italics="true" pageId="0" pageNumber="275">x</emphasis>
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</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">SD</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">SU 7184</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">H</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">P</th>
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<th colspan="1" pageId="0" pageNumber="275" rowspan="1">HUMZ 74990</th>
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</tr>
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<tr pageId="0" pageNumber="275">
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Total length (mm)</td>
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">710.0</td>
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">240.0-1120.0</td>
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">643.9</td>
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<td colspan="1" pageId="0" pageNumber="275" rowspan="1">344.0</td>
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|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">277.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">635.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">975.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">536.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Precaudal length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">77.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">76.9-80.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">78.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">79.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">81.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">79.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">78.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-second dorsal length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">61.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">58.4-63.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">60.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">60.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">64.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">63.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">59.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-first dorsal length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">32.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">28.3-31.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">30.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">36.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">32.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">31.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">28.4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pre-vent length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">50.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">46.4-52.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">49.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">50.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">52.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">46.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prepelvic length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">43.1-50.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">46.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">49.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">47.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">45.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prepectoral length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.0-24.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.4-24.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prebranchial length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">16.6-22.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">17.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">19.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prespiracular length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.4-21.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preorbital length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-10.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Prenarial length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1-6.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preoral length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.6-11.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Inner nostril-labial furrow space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.9-5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Mouth width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6-8.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Labial furrow length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1-2.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Internarial space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.6-5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Interorbital space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7-9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Eye length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.1-5.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Eye height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2-2.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Spiracle length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8-1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First gill-slit height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5-2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Fifth gill-slit height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.8-2.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Interdorsal space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.7-25.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">28.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Dorsal-caudal space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.1-11.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-pelvic space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.2-25.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic-caudal space</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">23.0-27.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">26.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.8-14.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin anterior margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.3-13.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin base length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.3-9.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9-9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin inner margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0-6.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.3</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin posterior margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.0-10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin spine length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1-5.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">First dorsal-fin spine base width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5-1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.2-13.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin anterior margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7-11.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin base length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.2-8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5-7.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin inner margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1-5.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin posterior margin</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.5-6.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin spine length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.3-5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Second dorsal-fin spine base width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7-1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin anterior margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.3-17.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">14.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin inner margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7-9.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin base length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.5-5.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pectoral-fin posterior margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.2-12.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.4-12.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Pelvic-fin inner margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.1-7.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Dorsal caudal margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">24.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.1-23.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">21.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">16.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">20.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Preventral caudal margin length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7-12.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.7</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Caudal fork width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.1-7.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width at nostrils</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.6-8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">6.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width at mouth</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.4-12.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">0.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.8</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">22.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.4-15.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Trunk width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">18.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.8-12.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Abdomen width</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">15.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.4-11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Head height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.3-12.5</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Trunk height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.7-14.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">11.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Abdomen height</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.7</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">12</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.9-14.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">10.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">13.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">8.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">9.6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Clasper outer length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.5-4.9</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">1.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4.3</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="275">
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">Clasper inner length</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.4-8.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">5.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">2.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">3.1</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">7.4</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="275" rowspan="1">-</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="275">Additional material.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
HUMZ 33680, adult female, 760 mm TL, East China Sea,
|
|
<geoCoordinate degrees="29" direction="north" minutes="38" orientation="latitude" precision="925" value="29.633333">29°38'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="134" direction="east" orientation="longitude" precision="55555" value="134.0">134°E</geoCoordinate>
|
|
; HUMZ 79798, adult female, 835 mm TL, Kyushu-Palau Ridge, 26°46.5'-26°46.6'N, 135°20.3'-135°20.8'E, 340-640 m depth; HUMZ 89858, juvenile female, 705 mm TL, off Hachijo-jima Island, Tokyo, Japan; HUMZ 97463, adult female, 800 mm TL, Ishikari Bay, Tokyo, Japan,
|
|
<geoCoordinate degrees="43" direction="north" minutes="12.4" orientation="latitude" precision="92" value="43.206665">43°12.4'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="141" direction="east" minutes="08.5" orientation="longitude" precision="92" value="141.14166">141°08.5'E</geoCoordinate>
|
|
, 22 m depth; HUMZ 102986, adult female, 1005 mm TL, Central Pacific Ocean, near Northern Mariana Islands,
|
|
<geoCoordinate degrees="17" direction="north" minutes="39.4" orientation="latitude" precision="92" value="17.656666">17°39.4'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="145" direction="east" minutes="50.3" orientation="longitude" precision="92" value="145.83833">145°50.3'E</geoCoordinate>
|
|
, 450 m depth; HUMZ 102987, adult female, 970 mm TL, Central Pacific Ocean, near Northern Mariana Islands,
|
|
<geoCoordinate degrees="19" direction="north" minutes="09" orientation="latitude" precision="925" value="19.15">19°09'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate degrees="142" direction="east" minutes="59" orientation="longitude" precision="925" value="142.98334">142°59'E</geoCoordinate>
|
|
, 520 m depth; HUMZ 102988, adult female, 1025 mm TL, same locality as HUMZ 102987; HUMZ 113586, adult female, 1120 mm TL, off Shirahama, Shimoda, Shizuoka Prefecture, Japan, 45 m depth; HUMZ 113587, adult female, 990 mm TL, same locality as HUMZ 113586; HUMZ 113588 adult female, 960 mm TL, same locality as HUMZ 113586; NSMT-P 44097, adult female, 740 mm TL, Izu, Honshu, Japan; NSMT-P 44381, adult male, 770 mm TL, off Koura, Suruga Bay, Honshu, Japan; NSMT-P 65518, adult female, 1119 mm TL, Suno-saki Point, Boso Peninsula, Chiba Prefecture, Honshu, Japan, 25 m depth; NSMT-P 77187, adult male, 1000 mm TL, unknown locality; NSMT-P 97762, neonate male, 278 mm TL, Tsushima, Nagasaki Prefecture, Kyusyu, Japan; SU 14245, adult female, 580 mm TL, Hong Kong, China; ZUMT 1360, female embryo, 178 mm TL, off Atami, Kanagawa Prefecture, Japan; ZUMT 21114, female embryo, 190 mm TL, Nagasaki, Japan.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="275">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Large-sized (710-1120 mm TL for adults)
|
|
<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus</emphasis>
|
|
</taxonomicName>
|
|
species that is distinguished from its regional congeners by: body dark grey to black in colour vs. body grey to light grey for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
vs. brownish-grey for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
vs. brown for
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.; dermal denticles tricuspidate and rhomboid (except for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
) vs. unicuspid and lanceolate.
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
is separated from
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
when adults by prenarial length 5.6%, 4.1%-5.6%TL (vs. 5.9%, 5.9%-6.8% TL for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
vs. 4.0%, 3.8%-4.3% TL for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
) and from
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by smaller eyes, its length 3.6%, 3.1%-4.0% TL (vs. 5.0%, 4.2%-5.1% TL for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
vs. 4.9%, 4.4%-4.9% TL for
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.) and length of second dorsal-fin spine 0.8-1.1 times length of first dorsal-fin spine (vs. 1.3-1.8 times for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
vs. 1.2-1.3 times for
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.). It is further distinct from
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
by smaller preoral length and distance nostril-upper labial furrow (10.3%, 7.6%-10.3% TL and 4.3%, 3.9%-4.5% TL vs. 10.7%, 10.7%-12.2% TL and 4.6%, 4.6%-5.1% TL for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
), more elongate first dorsal fin, its base length 8.2%, 7.6%-9.0% TL (vs. 6.7%, 6.6%-7.5% TL for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
), broader pectoral fins and head (pectoral-fin posterior margin length 11.7%, 9.9%-12.5% TL and head width at mouth 12.2%, 10.9%-12.2% TL vs. 8.6%, 7.8%-9.2% TL and 9.7%, 9.7%-10.6% TL for
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
differs from
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
by: pectoral fin with free rear tips rounded and posterior margin straight to weakly concave vs. free rear tips pointed and posterior margin conspicuously concave for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
; shorter pectoral-fin inner margin, its length 9.5%, 7.7%-9.5% TL vs. 10.3%, 9.8%-11.2% TL for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
; postventral caudal margins not uniformly white vs. uniformly white for
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="description">
|
|
<paragraph pageId="0" pageNumber="275">Description.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">External morphology.</emphasis>
|
|
Body fusiform, markedly humped anteriorly from posterior margin of the spiracle to pelvic fin origin, turning slender posteriorly; body conspicuously robust and stout with head height 1.2 (0.8-1.4) times trunk height and 1.6 (0.9-1.7) times abdomen height and head width 1.2 (1.0-1.6) times trunk width and 1.5 (1.1-2.6) times abdomen width (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2A-C</figureCitation>
|
|
). Head flattened anteriorly, conspicuously elongate, its length 24.2% (20.4%-24.4%) TL and broad, its width at mouth 12.2% (10.4%-12.8%) TL. Snout rounded at the tip and noticeably elongate, its preorbital length 7.3% (6.9%-10.7%) TL; prenarial length 1.3 (1.1-1.4) times distance between nostrils to upper labial furrow and 0.5 (0.5-0.6) times preoral length; anterior margin of nostrils wide and bi-lobed; internarial space 1.3 (0.8-1.4) times eye length (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2D, E</figureCitation>
|
|
). Eyes elliptical with anterior margin concave and posterior margin notched, placed laterally; eyes large, its length 4.1 (1.6-3.0) times eye height. Prespiracular length 0.5 (0.5-1.0) times prepectoral length and 1.8 (1.5-2.8) times preorbital length. Spiracles crescent, located laterally and behind the eyes; spiracles elongate, its length 0.4 (0.2-0.4) times eye length. Prebranchial length 1.6 (1.0-1.8) times prespiracular length. Gill slits vertical and concave, placed anterior to origin of pectoral fins; gill slits very tall with first gill slit height 1.7% (1.5%-2.8%) TL; fifth gill slit 1.3 (0.7-1.7) times taller than first gill slit.
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275" start="Figure 2" startId="F2">
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" pageId="0" pageNumber="275">Figure 2.</emphasis>
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
: lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A-C</emphasis>
|
|
) and ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">D, E</emphasis>
|
|
) views; first (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">F, J</emphasis>
|
|
) and second (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">G, K</emphasis>
|
|
) dorsal fins; pectoral (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">H, L</emphasis>
|
|
) and caudal (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">I, M</emphasis>
|
|
) fins. SU 12794 (paratype), adult male, 770 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A, D, F-I</emphasis>
|
|
); HUMZ 102988, adult female, 1025 mm TL (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">B, E, J-M</emphasis>
|
|
); NSMT-P 97762, neonate male, 278 mm TL (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">C</emphasis>
|
|
). Scale bars: 50 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A, B, D, E</emphasis>
|
|
); 25 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">C</emphasis>
|
|
); 20 mm (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">F-M</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Preoral length 1.2 (1.0-1.5) times greater than mouth width. Mouth somewhat straight and conspicuously broad (mouth width 1.8, 1.5-2.1 times internarial width and 1.5, 1.2-1.8 times prenarial length); upper labial furrow elongate, its length 2.5% (2.1%-2.7%) TL, bearing a thin fold; lower labial furrow markedly elongate, lacking fold. Teeth unicuspid, flattened labial-lingually, similar in both jaws; teeth somewhat rectangular, very broad at the crown; cusp short, thick, pointed and oblique; mesial cutting edge convex and oblique; distal heel conspicuously rounded; mesial heel sharp; apron thick and small, placed more laterally in the upper teeth and in the midline of crown base in the lower teeth, slightly more elongate in the lower teeth than in the upper teeth; median teeth present on upper jaw in adult paratype only and distinct from the subsequent teeth by: teeth hexagonal with cusp and apron placed more medially; both mesial and distal heels pointed (Fig.
|
|
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Upper (A, C, E, G, I) and lower (B, D, F, H, J) teeth of S. shiraii sp. nov. (A, B; HUMZ 80329), S. mitsukurii (C, D; HUMZ 33680), S. japonicus (E, F; HUMZ 189737), S. brevirostris (G, H; HUMZ 189762) and S. suckleyi (I, J; NSMT-P 92640). Scale bars: 1 mm." figureDoi="10.3897/zse.96.51962.figure3" httpUri="https://binary.pensoft.net/fig/420111" pageId="0" pageNumber="275">3C, D</figureCitation>
|
|
). One to two series of functional teeth on upper jaw and two series in lower jaw; 13-0-15 for holotype (13-1-13) teeth rows on upper jaw; 12-0-12 (11-0-11) teeth rows on lower jaw.
|
|
</paragraph>
|
|
<caption doi="10.3897/zse.96.51962.figure3" httpUri="https://binary.pensoft.net/fig/420111" pageId="0" pageNumber="275" start="Figure 3" startId="F3">
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" pageId="0" pageNumber="275">Figure 3.</emphasis>
|
|
Upper (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A, C, E, G, I</emphasis>
|
|
) and lower (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">B, D, F, H, J</emphasis>
|
|
) teeth of
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. (
|
|
<emphasis bold="true" pageId="0" pageNumber="275">A, B</emphasis>
|
|
; HUMZ 80329),
|
|
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="275">C, D</emphasis>
|
|
; HUMZ 33680),
|
|
<taxonomicName genus="S." lsidName="S. japonicus" pageId="0" pageNumber="275" rank="species" species="japonicus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. japonicus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="275">E, F</emphasis>
|
|
; HUMZ 189737),
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="275">G, H</emphasis>
|
|
; HUMZ 189762) and
|
|
<taxonomicName genus="S." lsidName="S. suckleyi" pageId="0" pageNumber="275" rank="species" species="suckleyi">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. suckleyi</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis bold="true" pageId="0" pageNumber="275">I, J</emphasis>
|
|
; NSMT-P 92640). Scale bars: 1 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Interdorsal space 2.0 (1.9-3.0) times larger than dorsal-caudal space and 0.9 (0.9-1.3) times prepectoral length. Origin of first dorsal fin located prior to vertical traced at pectoral-fin free rear tips. First dorsal fin elongate, its length 1.1 (1.0-1.3) times length of second dorsal fin; first dorsal-fin base length 0.8 (0.7-1.3) times height of first dorsal fin. First dorsal fin conspicuously broad at fin web; first dorsal-fin anterior margin markedly convex and elongate, its length 12.0% (9.3%-13.2%) TL; first dorsal-fin posterior margin concave, its length 9.3% (7.0%-10.6%) TL; first dorsal-fin inner margin elongate, its length 6.2% (5.0%-6.6%) TL; first dorsal-fin apex broadly rounded; first dorsal fin vertical and low, its height corresponding to 1.6 (1.1-1.6) times length of first dorsal-fin inner margin and 1.3 (0.8-1.4) times preorbital length (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2F, J</figureCitation>
|
|
). Origin of first dorsal-fin spine anterior to vertical traced at pectoral-fin free rear tips. First dorsal-fin spine thick, its width at base 1.0% (0.5%-1.0%) TL; first dorsal-fin spine elongate, its length 3.9% (2.1%-5.4%) TL) and corresponding to 0.4 (0.3-0.6) times first dorsal fin height, although not reaching first dorsal-fin apex. Pre-second dorsal length 2.5 (2.4-3.1) times prepectoral length. Origin of second dorsal fin behind a vertical traced at pelvic-fin free rear tips. Second dorsal fin broad at fin web with second dorsal-fin apex conspicuously rounded and lobe-like; second dorsal fin small, its length at base 0.9 (0.7-1.3) times first dorsal fin base length; second dorsal fin oblique and low with second dorsal-fin height 1.3 (0.8-1.4) times length of second dorsal-fin inner margin; second dorsal-fin anterior margin convex and elongate, its length 10.2% (7.7%-11.9%) TL; second dorsal-fin posterior margin concave and somewhat falcate, its length 6.3% (4.5%-6.3%) TL; second dorsal-fin inner margin length 5.3% (4.1%-5.9%) TL (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2G, K</figureCitation>
|
|
). Second dorsal-fin spine elongate, its length 0.6 (0.5-1.0) times second dorsal fin height and 1.1 (0.8-1.9) times length of first dorsal-fin spine, although never reaching second dorsal-fin apex; second dorsal-fin spine thick, its base width 0.9% (0.7%-1.1%) TL.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Pectoral fins markedly broad and elongate (pectoral-fin posterior margin length 11.7%, 9.2%-12.5% TL), although never extending up to trunk height when adpressed laterally on body; pectoral-fin anterior margin straight; pectoral-fin inner margin convex; pectoral-fin posterior margin weakly concave medially; pectoral-fin apex rounded and lobe-like; pectoral-fin free rear tips rounded, reaching the same length when a horizontal line is traced at pectoral-fin apex (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2H, L</figureCitation>
|
|
); pectoral-fin anterior margin large, its length 1.6 (1.4-2.3) times greater than pectoral-fin inner margin length and 1.3 (1.1-1.5) times pectoral-fin posterior margin length. Pectoral-pelvic space 0.9 (0.7-1.1) times pelvic-caudal space. Pelvic fins broad with pelvic-fin anterior and posterior margins convex; pelvic-fin free rear tips rounded and lobe-like; pelvic fin elongate, its length 11.5% (8.4%-12.6%) TL; pelvic fins nearer to second dorsal fin than to first dorsal fin. Adult males with clasper extremely robust and thick; clasper groove medial-dorsally and longitudinal, profound and short, not extending forward to inner margin of the clasper; clasper inner margin length 0.7-1.3 times length of pelvic-fin inner margin; apopyle broad, placed anteriorly in the clasper groove; hypopyle wide, located just above the rhipidion; rhipidion blade-like and evidently slender and elongate, placed at the medial-distal end of the clasper.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
Caudal keel prominent and situated laterally in the precaudal peduncle since insertion of second dorsal fin to behind origin of caudal fin. Caudal fin conspicuously slender in the upper caudal lobe; dorsal caudal margin somewhat convex proximally and conspicuously straight distally (Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2I, M</figureCitation>
|
|
); caudal fin elongate with dorsal caudal margin length 1.0 (0.9-1.1) times head length and 2.0 (1.7-2.0) times larger than length of preventral caudal margin; upper postventral caudal margin straight; lower postventral caudal margin somewhat convex; preventral caudal margin convex, its length 1.9 (1.5-3.8) times length of pelvic-fin inner margin; both posterior and ventral caudal tips rounded and thin; caudal fork discontinuous and conspicuously concave, its width 7.0% (6.1%-7.7%) TL.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<taxonomicName genus="Squamation" lsidName="Squamation" pageId="0" pageNumber="275" rank="genus">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">Squamation</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Scanning electron microscopy of the dermal denticles. Squalus shiraii sp. nov. (A, B): HUMZ 149389 (holotype), adult male, 590 mm TL. S. formosus (C, D): CSIRO H- 6816 - 01 (holotype), adult male, 720 mm TL. S. mitsukurii (E, F): HUMZ 102987, adult male, 970 mm TL. S. japonicus (G, H): NSMT-P 44380 (neotype), adult male, 645 mm TL (G); NSMT-P 91127, juvenile female, 410 mm TL (H). S. brevirostris (I, J): NSMT-P 47378, adult female, 600 mm TL (I); KAUM-I 187, adult female, 578 mm TL (J). S. suckleyi (K, L): NSMT P- 92640, adult female, 970 mm TL. Scale bars: 50 μm (B); 100 μm (D, F, L); 200 μm (A, C, E, G-K)." figureDoi="10.3897/zse.96.51962.figure7" httpUri="https://binary.pensoft.net/fig/420115" pageId="0" pageNumber="275">7E, F</figureCitation>
|
|
). Dermal denticles tricuspid, rhomboid to heart-shaped and imbricate, conspicuously broad at the crown, its length equal to its width; cusps pointed and posterior; lateral cusps often inconspicuous and much shorter than median cusp; median cusp conspicuous and wide; median ridge prominent, tall and thick with anterior furrow oval and profound; anterior margin of the crown broad, convex and arrow-shaped with one small ridge on each side; one to two lateral ridges, thin and smaller than median ridge. Dermal denticles slightly larger and more imbricate in adults than juveniles.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Colouration</emphasis>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Squalus mitsukurii: lateral (A-C) and ventral (D, E) views; first (F, J) and second (G, K) dorsal fins; pectoral (H, L) and caudal (I, M) fins. SU 12794 (paratype), adult male, 770 mm (A, D, F-I); HUMZ 102988, adult female, 1025 mm TL (B, E, J-M); NSMT-P 97762, neonate male, 278 mm TL (C). Scale bars: 50 mm (A, B, D, E); 25 mm (C); 20 mm (F-M)." figureDoi="10.3897/zse.96.51962.figure2" httpUri="https://binary.pensoft.net/fig/420110" pageId="0" pageNumber="275">2</figureCitation>
|
|
). Body black dorsally, dark grey ventrally and laterally since first dorsal fin insertion to origin of caudal fin. Dorsal fins blackish, dark grey at fin base near the dorsal-fin spines; lower half of anterior margin of dorsal fins whitish. Dorsal-fin spines brownish anteriorly and white posteriorly and at the tips. Pectoral fins black dorsally and ventrally, although dark grey near pectoral-fin inner margin; pectoral-fin posterior margin fairly white. Pelvic fins dark grey dorsally and ventrally, somewhat dark brown at the fin base; pelvic-fin posterior margins slightly white. Caudal fin black, dark grey near caudal fin origin and proximally over vertebral column; dorsal caudal margin dark grey proximally; postventral caudal margins slightly white, except near the caudal fork; preventral caudal margin blackish. Embryos, neonates and young juveniles differ from adults by having body light grey, dorsal fins light grey with dorsal-fin apex blackish and dorsal-fin free rear tips white, caudal fin light grey with postventral caudal margins broadly white, dark caudal bar conspicuous, black upper caudal blotch prominent and black caudal blotch at the base of lower caudal lobe, located horizontally.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="275">
|
|
<taxonomicName genus="Vertebral" lsidName="Vertebral" pageId="0" pageNumber="275" rank="genus">Vertebral</taxonomicName>
|
|
counts
|
|
</emphasis>
|
|
(Table
|
|
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Meristic data for species of Squalus from Japan. H: holotype; P: paratypes; NT: neotype; N: number of specimens; x; mean; m: mode; SD: standard deviation." httpUri="http://table.plazi.org/id/2DA11A1AF0C18FEB0512CBB43BD9057C" pageId="0" pageNumber="275" tableUuid="2DA11A1AF0C18FEB0512CBB43BD9057C">6</tableCitation>
|
|
). 68 (holotype) 70-72 (other material) diplospondylous vertebrae; 30 (26-31) caudal vertebrae; 116-117 total vertebrae.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="distribution">
|
|
<paragraph pageId="0" pageNumber="275">Geographical distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
is a regional endemic to the North-western Pacific Ocean with records along the coasts of China, Marianna Islands and Japan. It inhabits continental shelves and upper continental slopes within depth ranges of 22 to 640 m (Fig.
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Map of the North-western Pacific Ocean, showing the geographical distribution of Squalus species. A: S. shiraii sp. nov. (red); B: S. mitsukurii (orange); C: S. japonicus (brown); D: S. brevirostris (yellow); E: S. acutirostris (green); F: S. suckleyi (pink). Star: holotype / neotype; Dotted circle: paratypes; Circle: other material." figureDoi="10.3897/zse.96.51962.figure8" httpUri="https://binary.pensoft.net/fig/420116" pageId="0" pageNumber="275">8B</figureCitation>
|
|
). This species possibly reaches depths of up to 980 m in the Western Pacific Ocean, but it is often caught between 180-300 m depth (
|
|
<bibRefCitation author="Chen, CT" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" pagination="26 - 42" refId="B8" refString="Chen, CT, Taniuchi, T, Nose, Y, 1979. Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology 26 (1): 26 - 42" title="Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus." volume="26" year="1979">Chen et al. 1979</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="275" type="remarks">
|
|
<paragraph pageId="0" pageNumber="275">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="275">
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
has been misidentified with many nominal species in all oceans and, more particularly, with
|
|
<taxonomicName genus="S." lsidName="S. acutirostris" pageId="0" pageNumber="275" rank="species" species="acutirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. acutirostris</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. boretzi" pageId="0" pageNumber="275" rank="species" species="boretzi">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. boretzi</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. in the North Pacific Ocean.
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
exhibits smaller interdorsal space than
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
(21.1%, 21.1%-25.9% TL vs. 26.7%-30.0% TL), larger first and second dorsal fins (first dorsal-fin base length 8.2%, 7.6%-9.0% TL and second dorsal-fin base length 7.2%, 6.8%-7.9% TL vs. 6.2%-7.4% TL and 4.9%-5.5% TL in
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
), which corroborates the findings of
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.798.28375" author="Daly-Engel, TS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="275" pagination="135 - 157" refId="B18" refString="Daly-Engel, TS, Koch, A, Anderson, JM, Cotton, CF, Grubbs, RD, 2018. Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific. ZooKeys 798: 135 - 157, DOI: https://doi.org/10.3897/zookeys.798.28375" title="Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific." url="https://doi.org/10.3897/zookeys.798.28375" volume="798" year="2018">Daly-Engel et al. (2018)</bibRefCitation>
|
|
. Torso measurements that are suggested as being diagnostic to separate these species in the referred study show minor overlapping and they must be used with caution despite range values of the direct measurements (e.g. pre-first and pre-second dorsal lengths, precaudal length) are somewhat higher in
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
. The Japanese species further exhibit more elongated pectoral fins, its inner margin 9.5%, 7.7%-9.5% TL (vs. 6.4%-7.4% TL in
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
<taxonomicName genus="Vertebral" lsidName="Vertebral" pageId="0" pageNumber="275" rank="genus">Vertebral</taxonomicName>
|
|
counts are congruent between these species, which is contrary to
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.798.28375" author="Daly-Engel, TS" journalOrPublisher="ZooKeys" pageId="0" pageNumber="275" pagination="135 - 157" refId="B18" refString="Daly-Engel, TS, Koch, A, Anderson, JM, Cotton, CF, Grubbs, RD, 2018. Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific. ZooKeys 798: 135 - 157, DOI: https://doi.org/10.3897/zookeys.798.28375" title="Description of a new deep-water dogfish shark from Hawaii, with comments on the Squalus mitsukurii species complex in the West Pacific." url="https://doi.org/10.3897/zookeys.798.28375" volume="798" year="2018">Daly-Engel et al. (2018)</bibRefCitation>
|
|
when comparing data taken from
|
|
<bibRefCitation author="Chen, CT" journalOrPublisher="Japanese Journal of Ichthyology" pageId="0" pageNumber="275" pagination="26 - 42" refId="B8" refString="Chen, CT, Taniuchi, T, Nose, Y, 1979. Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus. Japanese Journal of Ichthyology 26 (1): 26 - 42" title="Blainville's dogfish, Squalus blainvillei, from Japan, with notes on S. mitsukurii and S. japonicus." volume="26" year="1979">Chen et al. (1979)</bibRefCitation>
|
|
. The latter work possibly included specimens of
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. which is the only Japanese species with the highest number of vertebral counts. Morphologically similar
|
|
<taxonomicName class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus</emphasis>
|
|
</taxonomicName>
|
|
species tend to exhibit overlapping of vertebral counts as seen here between
|
|
<taxonomicName genus="S." lsidName="S. mitsukurii" pageId="0" pageNumber="275" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. hawaiiensis" pageId="0" pageNumber="275" rank="species" species="hawaiiensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. hawaiiensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. boretzi" pageId="0" pageNumber="275" rank="species" species="boretzi">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. boretzi</emphasis>
|
|
</taxonomicName>
|
|
, with a few exceptions.
|
|
<taxonomicName authorityName="Jordan & Snyder" authorityYear="1903" class="Elasmobranchii" family="Squalidae" genus="Squalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squalus mitsukurii" order="Squaliformes" pageId="0" pageNumber="275" phylum="Chordata" rank="species" species="mitsukurii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">Squalus mitsukurii</emphasis>
|
|
</taxonomicName>
|
|
may be separated from
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<emphasis italics="true" pageId="0" pageNumber="275">
|
|
<taxonomicName genus="S." lsidName="S. melanurus" pageId="0" pageNumber="275" rank="species" species="melanurus">S. melanurus</taxonomicName>
|
|
<taxonomicName genus="S." lsidName="S. blainvillei" pageId="0" pageNumber="275" rank="species" species="blainvillei">S. blainvillei</taxonomicName>
|
|
</emphasis>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. grahami" pageId="0" pageNumber="275" rank="species" species="grahami">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. grahami</emphasis>
|
|
</taxonomicName>
|
|
by monospondylous vertebrae and from
|
|
<taxonomicName genus="S." lsidName="S. blainvillei" pageId="0" pageNumber="275" rank="species" species="blainvillei">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. blainvillei</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. melanurus" pageId="0" pageNumber="275" rank="species" species="melanurus">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. melanurus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. montalbani" pageId="0" pageNumber="275" rank="species" species="montalbani">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. montalbani</emphasis>
|
|
</taxonomicName>
|
|
by diplospondylous vertebrae and
|
|
<taxonomicName genus="S." lsidName="S. brevirostris" pageId="0" pageNumber="275" rank="species" species="brevirostris">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. brevirostris</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.,
|
|
<taxonomicName genus="S." lsidName="S. blainvillei" pageId="0" pageNumber="275" rank="species" species="blainvillei">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. blainvillei</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. lalannei" pageId="0" pageNumber="275" rank="species" species="lalannei">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. lalannei</emphasis>
|
|
</taxonomicName>
|
|
by total vertebrae. Precaudal vertebrae separates it from
|
|
<taxonomicName genus="S." lsidName="S. shiraii" pageId="0" pageNumber="275" rank="species" species="shiraii">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. shiraii</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.,
|
|
<taxonomicName genus="S." lsidName="S. blainvillei" pageId="0" pageNumber="275" rank="species" species="blainvillei">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. blainvillei</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="S." lsidName="S. lalannei" pageId="0" pageNumber="275" rank="species" species="lalannei">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. lalannei</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="S." lsidName="S. montalbani" pageId="0" pageNumber="275" rank="species" species="montalbani">
|
|
<emphasis italics="true" pageId="0" pageNumber="275">S. montalbani</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |