250 lines
42 KiB
XML
250 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e82644" ID-GBIF-Dataset="1d579114-3554-4e38-bd81-ec33a8c44b9f" ID-Pensoft-Pub="1864-8312-80-279" ID-Pensoft-UUID="42DA6B89695151758680B79C0CF21997" ID-ZooBank="1D57CB0C56C44DA591E1CF7E9BB40E74" ModsDocID="1864-8312-80-279" checkinTime="1658318833579" checkinUser="pensoft" docAuthor="Yin, Zi-Wei, Tihelka, Erik, Lozano-Fernandez, Jesus & Cai, Chen-Yang" docDate="2022" docId="CA25E907AEFA593D8DC198430F341A6F" docLanguage="en" docName="ArthropodSystPhyl 80: 279-294" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-07-19" docSource="http://dx.doi.org/10.3897/asp.80.e82644" docTitle="Europharinodes schaufussi Yin & Cai 2022, sp. nov." docType="treatment" docUuid="5BB9EDF8-BE3F-43D5-854B-89585BF7E453" docUuidSource="ZooBank" docVersion="2" id="42DA6B89695151758680B79C0CF21997" lastPageNumber="279" masterDocId="42DA6B89695151758680B79C0CF21997" masterDocTitle="The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications" masterLastPageNumber="294" masterPageNumber="279" pageNumber="279" updateTime="1658319438294" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Yin, Zi-Wei</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6659-9448</mods:nameIdentifier>
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<mods:affiliation>Laboratory of Systematic Entomology, College of Life and Environmental Sciences, Shanghai Normal University, Shanghai, 200234, China</mods:affiliation>
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<mods:nameIdentifier type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Tihelka, Erik</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5048-5355</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lozano-Fernandez, Jesus</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3597-1221</mods:nameIdentifier>
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<mods:affiliation>Department of Genetics, Microbiology and Statistics, Biodiversity Research Institute (IRBio), University of Barcelona, Avd. Diagonal 643, 08028</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cai, Chen-Yang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9283-8323</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; cycai @ nigpas. ac. cn & School of Earth Sciences, University of Bristol, Life Sciences Building, Tyndall Avenue, Bristol, BS 8 1 TQ, UK</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-19</mods:number>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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<mods:extent unit="page">
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<mods:start>279</mods:start>
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<mods:end>294</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e82644</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e82644</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-279</mods:identifier>
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<mods:identifier type="ZooBank">1D57CB0C56C44DA591E1CF7E9BB40E74</mods:identifier>
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<mods:identifier type="Pensoft-UUID">42DA6B89695151758680B79C0CF21997</mods:identifier>
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<subSection pageId="0" pageNumber="279" type="3. results">
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<treatment LSID="urn:lsid:zoobank.org:act:5BB9EDF8-BE3F-43D5-854B-89585BF7E453" httpUri="http://treatment.plazi.org/id/CA25E907AEFA593D8DC198430F341A6F" lastPageNumber="279" pageId="0" pageNumber="279">
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<subSubSection pageId="0" pageNumber="279" type="nomenclature">
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<paragraph pageId="0" pageNumber="279">
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<taxonomicName LSID="http://zoobank.org/5BB9EDF8-BE3F-43D5-854B-89585BF7E453" authority="Yin & Cai" authorityName="Yin & Cai" authorityYear="2022" family="Staphylinidae" genus="Europharinodes" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Europharinodes schaufussi" order="Coleoptera" pageId="0" pageNumber="279" rank="species" species="schaufussi" status="sp. nov.">Europharinodes schaufussi Yin & Cai</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="279">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="279" type="description">
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<paragraph pageId="0" pageNumber="279">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">Figs 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Hybocephalini. A: Location of Baltic amber forest (yellow star) on a paleogeographic map of the middle Eocene. B: Distribution and species density of modern Hybocephalini (red dots) and location of Baltic amber deposit in the present-day Yantarny, Russia (yellow star)." figureDoi="10.3897/asp.80.e82644.figure6" httpUri="https://binary.pensoft.net/fig/717128" pageId="0" pageNumber="279">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="279" type="type material">
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<paragraph pageId="0" pageNumber="279">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="279">
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<emphasis bold="true" pageId="0" pageNumber="279">Holotype</emphasis>
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(#SNUC-Paleo-0102), deposited in SNUC; a complete, well-preserved male in a 11.6 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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7.1 mm
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<normalizedToken originalValue="×">x</normalizedToken>
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7.0 mm transparent yellow amber piece, without syninclusions; two surfaces regarding
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<normalizedToken originalValue="beetle’s">beetle's</normalizedToken>
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lateral and dorsal aspect were cut and polished for observation and photography.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="279" type="locality and horizon">
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<paragraph pageId="0" pageNumber="279">Locality and horizon.</paragraph>
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<paragraph pageId="0" pageNumber="279">
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Amber mined from the open-pit mine in Yantarny (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Distribution of Hybocephalini. A: Location of Baltic amber forest (yellow star) on a paleogeographic map of the middle Eocene. B: Distribution and species density of modern Hybocephalini (red dots) and location of Baltic amber deposit in the present-day Yantarny, Russia (yellow star)." figureDoi="10.3897/asp.80.e82644.figure6" httpUri="https://binary.pensoft.net/fig/717128" pageId="0" pageNumber="279">6</figureCitation>
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), Kaliningrad region, Russia; layers between the Bartonian-Priabonian "Wilde Erde" plus "Blaue Erde" and the lower Lutetian "Untere Blaue Erde"; mid-Eocene, 45.0-38.0 Ma (
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4664.2.8" author="Bukejs, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="279" pagination="261 - 273" refId="B7" refString="Bukejs, A, Alekseev, VI, Pollock, DA, 2019. Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa 4664: 261 - 273, DOI: https://doi.org/10.11646/zootaxa.4664.2.8" title="Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location." url="https://doi.org/10.11646/zootaxa.4664.2.8" volume="4664" year="2019">Bukejs et al. 2019</bibRefCitation>
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).
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</paragraph>
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<caption doi="10.3897/asp.80.e82644.figure6" httpUri="https://binary.pensoft.net/fig/717128" pageId="0" pageNumber="279" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="279">
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<emphasis bold="true" pageId="0" pageNumber="279">Figure 6.</emphasis>
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Distribution of
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<taxonomicName authorityName="Raffray" authorityYear="1890" lsidName="" pageId="0" pageNumber="279" rank="tribe" tribe="Hybocephalini">Hybocephalini</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="279">A</emphasis>
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: Location of Baltic amber forest (yellow star) on a paleogeographic map of the middle Eocene.
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<emphasis bold="true" pageId="0" pageNumber="279">B</emphasis>
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: Distribution and species density of modern
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<taxonomicName authorityName="Raffray" authorityYear="1890" lsidName="" pageId="0" pageNumber="279" rank="tribe" tribe="Hybocephalini">Hybocephalini</taxonomicName>
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(red dots) and location of Baltic amber deposit in the present-day Yantarny, Russia (yellow star).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="279" type="diagnosis">
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<paragraph pageId="0" pageNumber="279">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="279">
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As for the genus (
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<emphasis italics="true" pageId="0" pageNumber="279">vide supra</emphasis>
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), plus the following: body length approximately 2.1 mm; male antennae lacking modifications; sternites 4 and 5 each with a nodule at middle; aedeagus relatively stout, with short basoventral projection, median lobe with large basal capsule and dorsal diaphragm, sclerotized endophallus present, parameres elongate, each with two long setae at the apex.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="279" type="description">
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<paragraph pageId="0" pageNumber="279">Description.</paragraph>
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<paragraph pageId="0" pageNumber="279">
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<emphasis italics="true" pageId="0" pageNumber="279">Male</emphasis>
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. Body (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2B-D</figureCitation>
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;
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3A, B</figureCitation>
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) length approximately 2.1 mm. Most surface of body covered with squamous setae. Head (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3E-G</figureCitation>
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) roundly triangular, sub-rectangular at base, slightly longer than wide, length 0.47 mm, width across eyes 0.45 mm, length/width 1.04; vertex roughly punctate, almost flat, distinct vertexal foveae (dorsal tentorial pits) located above level of posterior margin of eyes; frons with narrow and short rostrum, anteriorly confluent with sharply declining clypeus, impressed between slightly raised antennal tubercles; clypeus with smooth surface, anterior margin broadly rounded, carinate and moderately raised; ocular-mandibular carina absent; postgenal region moderately projected. Venter with small, broadly separated gular foveae (posterior tentorial pits) in transverse impression, longitudinally ridged along middle, weakly impressed admesally. Compound eyes large and prominent, each composed of approximately 30 ommatidia, vertical length of eye/temple 2.1. Antenna moderately elongate, length 0.90 mm, lacking modification; antennomeres with rough surfaces; distinct club (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2B-D, F</figureCitation>
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) formed by enlarged apical three antennomeres; antennomere 1 thick, subcylindrical, 2 and 3 each slightly elongate, 2 slightly longer and wider than 3, 4-7 subequal in width, approximately as long as wide, 8 shortest, slightly transverse, 9 enlarged, slightly longer than wide (including basal stalk), 10 as long as and slightly wider than 9, 11 largest, approximately as long as 9 and 10 combined, 1.24 times as broad as 10.
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</paragraph>
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<paragraph pageId="0" pageNumber="279">
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Prothorax with dense squamous setae (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A, C, D, I: under incident light; B, E-H: X-ray microtomographic reconstruction). A, B Dorsal habitus. C Left half of pronotum, showing squamous setae. D Lateral portion of the abdominal base, showing squamous setae. E Head dorsum. F Head venter. G Head, in antero-lateral view. H, I Left maxillary palpus. Abbreviations: ce, compound eye; cl, clypeus; ff, frontal fovea; gf, gular fovea; la, labrum; ma, mandible; p 2 - 4, maxillary palpomere 2 - 4; vf, vertexal fovea. Scale bars: 0.5 mm in A, B, 0.2 mm in E-I." figureDoi="10.3897/asp.80.e82644.figure3" httpUri="https://binary.pensoft.net/fig/717125" pageId="0" pageNumber="279">3C</figureCitation>
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). Pronotum (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4A</figureCitation>
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) approximately as long as wide, length 0.42 mm, width 0.41 mm, length/width 1.02, widest at middle; sides rounded, convergent apically and basally; disc moderately convex; with distinct median and lateral antebasal foveae. Hypomeron confluent with prosternum, with antero-hypomeral fovea (Fig.
|
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4B, C</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">ahf</emphasis>
|
||
), lacking hypomeral ridge. Prosternum with anterior part much shorter than coxal part, with small, broadly separated lateral procoxal foveae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4C</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">lpcf</emphasis>
|
||
); margin of coxal cavity moderately carinate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
Elytra (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4D</figureCitation>
|
||
) with squamous setae finer than those of pronotum and abdomen, much wider than long, length 0.71 mm, width 0.90 mm, length/width 0.79 mm; each elytron with two large basal foveae and two distinct discal striae; humerus weakly prominent, lacking subhumeral fovea or marginal stria. Hindwing fully developed, membranous.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
Mesoventrite short, well-demarcated from metaventrite; median mesoventral foveae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4E</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">mmsf</emphasis>
|
||
) moderately separated, with large lateral mesoventral fovea, mesoventral process short. Metaventrite distinctly raised admesally, inclined towards middle, with well-developed lateral mesocoxal (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">lmcf</emphasis>
|
||
) and single median metaventral fovea (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">mmtf</emphasis>
|
||
), posterior margin broadly emarginate, convex at middle, with small, short median split (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4F</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">ms</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
Legs moderately elongate, lacking modification; mesotrochanter elongate; all femora coarsely punctate; tarsi with short tarsomeres 1 and long tarsomeres 2 and 3, with 3 slightly longer than 2; each tarsus with two sub-equal pretarsal claws (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Amber piece (A) containing Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102, and its morphological details (B-G) of (A, B, G: under incident light; C-F: X-ray microtomographic reconstruction). A Original amber piece containing the holotype before cutting and polishing. B, C Left lateral habitus. D Right lateral habitus. E Head in lateral view, show tentorial arms and pharynx. F Right antennal club. G Pro- and metatarsus and respective pretarsal claws. Abbreviations: aed, aedeagus; hw, hind wing; ph, pharynx; ta, tentorial arm. Scale bars: 0.5 mm in B-D, 0.2 mm in E, F." figureDoi="10.3897/asp.80.e82644.figure2" httpUri="https://binary.pensoft.net/fig/717124" pageId="0" pageNumber="279">2G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
Abdomen widest at lateral margins of paratergite 1 (IV), length 0.64 mm, width 0.66 mm; whole surface covered with broad squamous setae. Tergite 1 (IV) slightly longer than 2 (V), deeply and broadly sulcate at base, at least with one pair of basolateral foveae, lacking discal carina; tergite 2-4 (IV-VII) subequal in length along midline, each distinctly sulcate at base and with one pair of basolateral foveae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">blf</emphasis>
|
||
), tergite 5 (VIII) semicircular, transverse, posterior margin roundly emarginate at middle; accompanying paratergites 1-3 (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">pt1-3</emphasis>
|
||
) broad, 4 triangular. Sternites successively shorter; sternites 2-5 (IV-VII) medially slightly impressed, each with broad sulcus at base and one pair of basolateral foveae (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4G</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">blf</emphasis>
|
||
), lacking carina, 4 and 5 each with single nodule at middle (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. X-ray microtomographic reconstruction of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102. A - C: prothorax, dorsal (A), lateral (B) and ventral (C). D Right elytron. E, F Meso- and metathorax, ventral (E) and lateral (F). G - I Abdomen, dorsal (G), lateral (H) and ventral (I). Abbreviations: ahf, antero-hypomeral fovea; bef, basal elytral fovea; blf, basal lateral fovea; ds, discal stria; laf, lateral antebasal fovea; lmcf, lateral metaventral fovea; lpcf, lateral procoxal fovea; maf, median antebasal fovea; mmsf, median mesoventral fovea; mc, marginal carina; mmtf, median metaventral fovea; mn, median nodule; ms, median split; pt 1 - 3, paratergite 1 - 3; s 2 - 6, sternite 2 - 6 (IV-VIII); t 1 - 5, tergite 1 - 5 (IV-VIII). Scale bars: 0.2 mm." figureDoi="10.3897/asp.80.e82644.figure4" httpUri="https://binary.pensoft.net/fig/717126" pageId="0" pageNumber="279">4I</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">mn</emphasis>
|
||
), sternite 6 (VIII) transverse, posterior margin broadly emarginate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
Aedeagus (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5D-F, I-K</figureCitation>
|
||
) 0.28 mm long, dorso-ventrally symmetrical, median lobe (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5E, J</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">ml</emphasis>
|
||
) with large basal capsule (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5E, J</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">bc</emphasis>
|
||
) and dorsal diaphragm (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5D, I</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">dd</emphasis>
|
||
), narrowed towards apex and moderately bent ventrally, basoventral projection (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5E, J</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">bp</emphasis>
|
||
) short; endophallus (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5G-J</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">en</emphasis>
|
||
) well-sclerotized; parameres (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Morphology of Europharinodes schaufussi gen. et sp. nov., holotype, SNUC-Paleo- 0102 (A-H: X-ray microtomographic reconstruction; I-K: schematic). A Right foreleg. B Right middle leg. C Left hindleg. D - F, I - K Aedeagus, dorsal (D, I), lateral (E, J) and ventral (F, K). G, H endophallus, dorsal (G) and lateral (H). Abbreviations: bc, basal capsule; bp, basoventral projection; dd, dorsal diaphragm; en, endophallus; ml, median lobe; pa, paramere. Scale bars: 0.3 mm in A-C; 0.1 mm in D-K." figureDoi="10.3897/asp.80.e82644.figure5" httpUri="https://binary.pensoft.net/fig/717127" pageId="0" pageNumber="279">5E, J</figureCitation>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="279">pa</emphasis>
|
||
) paired, elongate, rounded at apices, each with two long apical setae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
<emphasis italics="true" pageId="0" pageNumber="279">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="279" type="etymology">
|
||
<paragraph pageId="0" pageNumber="279">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="279">
|
||
The new species is named after the German naturalist Ludwig W. Schaufuss (1833-
|
||
<bibRefCitation author="Schaufuss, LW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="0" pageNumber="279" pagination="101 - 162" refId="B53" refString="Schaufuss, LW, 1890. System-schema der Pselaphiden, ein Blick in die Vorzeit, in die Gegenwart und in die Zukunft. Tijdschrift voor Entomologie 33: 101 - 162" title="System-schema der Pselaphiden, ein Blick in die Vorzeit, in die Gegenwart und in die Zukunft." volume="33" year="1890">1890</bibRefCitation>
|
||
), who described most of the known fossil pselaphines from Baltic amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |