368 lines
38 KiB
XML
368 lines
38 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.889.39090" ID-GBIF-Dataset="f50f6008-500c-492f-84b8-295ba991ec7b" ID-PMC="PMC6872603" ID-Pensoft-Pub="1313-2970-889-81" ID-Pensoft-UUID="0C546DE0BA425539879C1421D8C25A63" ID-PubMed="31777439" ID-ZooBank="7E7A3D196D7040398C087B248A27EB33" ModsDocID="1313-2970-889-81" checkinTime="1574358895821" checkinUser="pensoft" docAuthor="Hendrich, Lars, Shaverdo, Helena, Hajek, Jiri & Balke, Michael" docDate="2019" docId="8B4B21BB58A65727A8F1B41547F93D7B" docLanguage="en" docName="ZooKeys 889: 81-152" docOrigin="ZooKeys 889" docSource="http://dx.doi.org/10.3897/zookeys.889.39090" docTitle="Copelatus marginatus Sharp 1882" docType="treatment" docVersion="4" id="0C546DE0BA425539879C1421D8C25A63" lastPageNumber="81" masterDocId="0C546DE0BA425539879C1421D8C25A63" masterDocTitle="Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="152" masterPageNumber="81" pageNumber="81" updateTime="1668168070667" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of Australian Copelatus Erichson, 1832 (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hendrich, Lars</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hajek, Jiri</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>889</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>81</mods:start>
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<mods:end>152</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.889.39090</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.889.39090</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-889-81</mods:identifier>
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<mods:identifier type="ZooBank">7E7A3D196D7040398C087B248A27EB33</mods:identifier>
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<mods:identifier type="Pensoft-UUID">0C546DE0BA425539879C1421D8C25A63</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="160511456" LSID="urn:lsid:plazi:treatment:8B4B21BB58A65727A8F1B41547F93D7B" httpUri="http://treatment.plazi.org/id/8B4B21BB58A65727A8F1B41547F93D7B" lastPageNumber="81" pageId="0" pageNumber="81">
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<subSubSection pageId="0" pageNumber="81" type="nomenclature">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName LSID="8B4B21BB-58A6-5727-A8F1-B41547F93D7B" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marginatus">Copelatus marginatus Sharp, 1882</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Copelatus marginatus, female. Total length 5.8 mm." figureDoi="10.3897/zookeys.889.39090.figure6" httpUri="https://binary.pensoft.net/fig/356475" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure6">Figures 6</figureCitation>
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,
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Copelatus marginatus A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure15" httpUri="https://binary.pensoft.net/fig/356484" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure15">15</figureCitation>
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,
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<figureCitation captionStart="Figure 25" captionStartId="F24" captionText="Figure 25. Distribution of Copelatus marginatus." figureDoi="10.3897/zookeys.889.39090.figure25" httpUri="https://binary.pensoft.net/fig/356493" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure25">25</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="reference_group">
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marginatus">Copelatus marginatus</taxonomicName>
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<bibRefCitation author="Sharp, D" journalOrPublisher="Scientific Transactions of the Royal Dublin Society Series" pageId="0" pageNumber="81" pagination="179 - 1003" refId="B33" refString="Sharp, D, 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society Series 2: 179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae." volume="2" year="1882">Sharp 1882</bibRefCitation>
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: 579 (original description);
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<bibRefCitation DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts 1978</bibRefCitation>
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: 124 (description);
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<bibRefCitation author="Watts, CHS" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="81" pagination="22 - 28" refId="B37" refString="Watts, CHS, 1985. A faunal assessment of Australian Hydradephaga. Proceedings of the Academy of Natural Sciences of Philadelphia 137: 22 - 28" title="A faunal assessment of Australian Hydradephaga." volume="137" year="1985">Watts 1985</bibRefCitation>
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: 26 (general distribution);
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<bibRefCitation author="Larson, DJ" journalOrPublisher="Proceedings of the Royal Society of Queensland" pageId="0" pageNumber="81" pagination="47 - 63" refId="B20" refString="Larson, DJ, 1993. Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species. Proceedings of the Royal Society of Queensland 103: 47 - 63" title="Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species." volume="103" year="1993">Larson 1993</bibRefCitation>
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: 53 (habitat description);
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<bibRefCitation author="Watts, CHS" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" pagination="1 - 110" refId="B38" refString="Watts, CHS, 2002. Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia). Identification Ecology Guide 43: 1 - 110" title="Checklist and guides to the identification, to genus, of adults and larval Australian water beetles of the families Dytiscidae, Noteridae, Hygrobiidae, Haliplidae, Gyrinidae, Hydraenidae and the superfamily Hydrophiloidea (Insecta-Coleoptera). Cooperative Research Centre for Freshwater Ecology (Australia)." volume="43" year="2002">Watts 2002</bibRefCitation>
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: 42 (checklist);
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<bibRefCitation author="Wewalka, G" editor="Jaech, MA" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" refId="B39" refString="Wewalka, G, Balke, M, Hendrich, L, Jaech, MA, Balke, M, 2010. Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1).Monographs on Coleoptera 3, 45-128." title="Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1). Monographs on Coleoptera 3, 45 - 128." year="2010">Wewalka et al. 2010</bibRefCitation>
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: 50-51 (description, new records);
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<bibRefCitation author="Nilsson, AN" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B28" refString="Nilsson, AN, Hajek, J, 2019. .http://www.waterbeetles.eu" url="http://www.waterbeetles.eu" year="2019">
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Nilsson and
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<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
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2019
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</bibRefCitation>
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: 58 (catalogue).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type locality">
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<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="81">"Australia, [Queensland], Rockhampton".</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="type material">
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<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<taxonomicName genus="Lectotype" lsidName="Lectotype" pageId="0" pageNumber="81" rank="genus">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Lectotype</emphasis>
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</taxonomicName>
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(designated by
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<bibRefCitation DOI="https://doi.org/10.1071/AJZS057" author="Watts, CHS" journalOrPublisher="Australian Journal of Zoology Supplement Series" pageId="0" pageNumber="81" pagination="1 - 166" refId="B36" refString="Watts, CHS, 1978. A revision of the Australian Dytiscidae (Coleoptera). Australian Journal of Zoology Supplement Series 57: 1 - 166" title="A revision of the Australian Dytiscidae (Coleoptera)." url="https://doi.org/10.1071/AJZS057" volume="57" year="1978">Watts: 1978</bibRefCitation>
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: 124): Male, "
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<taxonomicName genus="Lectotype" lsidName="Lectotype" pageId="0" pageNumber="81" rank="genus">Lectotype</taxonomicName>
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" [printed label with violet frame]; "Rockhampton 675." [printed label]; "Sharp Coll 1905-313." [printed label]; "Type 675
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marginatus">Copelatus marginatus</taxonomicName>
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Rockhampton" [hw label] (NHMUK).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Paralectotypes</emphasis>
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: 1 male, 5 females,
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<normalizedToken originalValue="“Paralectotype”">"Paralectotype"</normalizedToken>
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[printed label with blue frame]; "Rockhampton 675." [printed label]; "Sharp Coll 1905-313." [printed label]; "Type 675
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marginatus">Copelatus marginatus</taxonomicName>
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Co-type" [hw label] (NHMUK).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="81" type="materials_examined">
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<paragraph pageId="0" pageNumber="81">Additional material studied (244 specimens).</paragraph>
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<paragraph pageId="0" pageNumber="81">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Western Australia:</emphasis>
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1 ex., "Australia07, WA 50, Parrys Lagoon, Marigu Billabong, 25 km SE Wyndham, 77 m, 15.32.98S 128.15.59E, M. Baehr" (ZSM); 1 ex., "AUSTRALIA/WA, Koolan Island [
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<geoCoordinate degrees="16" direction="south" minutes="09" orientation="latitude" precision="925" value="-16.15">16°09'S</geoCoordinate>
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,
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<geoCoordinate degrees="123" direction="east" minutes="45" orientation="longitude" precision="925" value="123.75">123°45'E</geoCoordinate>
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],
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<normalizedToken originalValue="VIII–IX">VIII-IX</normalizedToken>
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1967, Milton O. leg." (WAM); 1 ex., "AUSTRALIA/WA, Derby [
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<geoCoordinate degrees="17.3166" direction="south" orientation="latitude" precision="5" value="-17.3166">-17.3166</geoCoordinate>
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,
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<geoCoordinate degrees="123.6333" direction="east" orientation="longitude" precision="5" value="123.6333">123.6333</geoCoordinate>
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], 2.1962, Beamish, G. leg." (WAM); 1 ex., idem but
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<normalizedToken originalValue="„10.4.1964”">"10.4.1964"</normalizedToken>
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(WAM).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Northern Territory</emphasis>
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: 1 ex., "Australia, N.T. Binis Track 10.IV.2011 LF; H = 427 m
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<geoCoordinate degrees="20" direction="south" minutes="51" orientation="latitude" precision="1" seconds="51.4" value="-20.864279">20°51'51.4"S</geoCoordinate>
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,
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<geoCoordinate degrees="135" direction="east" minutes="12" orientation="longitude" precision="1" seconds="01.6" value="135.20044">135°12'01.6"E</geoCoordinate>
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leg. Michael Langer" (ZSM); 1 ex., "Australia, NT, 80 km W Roper Bar,
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<geoCoordinate degrees="14" direction="south" minutes="54" orientation="latitude" precision="15" seconds="19" value="-14.905277">14°54'19"S</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="57" orientation="longitude" precision="15" seconds="28" value="133.95778">133°57'28"E</geoCoordinate>
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, 78 m, at light, 14.V.2006, leg. Berger & Dostal (7/06)" (CGW); 1 ex., "Australia, NT, Kakadu NP, Muirella, 40 m, at light, 10.V.2006, leg. Berger & Dostal (3/06)" (CGW); 1 ex., "Katherine (T) Vestjens, 1.12.1967 [
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<geoCoordinate degrees="14.51667" direction="south" orientation="latitude" precision="1" value="-14.51667">-14.51667</geoCoordinate>
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,
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<geoCoordinate degrees="132.3667" direction="east" orientation="longitude" precision="5" value="132.3667">132.3667</geoCoordinate>
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], W.J.M. Tindal" (ANIC); 1 ex., "Katherine [
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<geoCoordinate degrees="14.46667" direction="south" orientation="latitude" precision="1" value="-14.46667">-14.46667</geoCoordinate>
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,
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<geoCoordinate degrees="132.2667" direction="east" orientation="longitude" precision="5" value="132.2667">132.2667</geoCoordinate>
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], 9.2.1968, Watson, J.A.L." (ANIC); 1 ex., "Howard Springs 27.01.1968 [-12.45, 131.05] Watson, J.A.L." (ANIC); 1 ex., "Northern Territory Katherine [
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<geoCoordinate degrees="14.46667" direction="south" orientation="latitude" precision="1" value="-14.46667">-14.46667</geoCoordinate>
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,
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<geoCoordinate degrees="132.2667" direction="east" orientation="longitude" precision="5" value="132.2667">132.2667</geoCoordinate>
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], 6.2.1968, Matthews, E." (ANIC); 1 ex., "Northern Territory, Lee Point, Darwin [-12.33333333, 130.9], 7.3.1967, Upton, M.S." (ANIC); 1 ex., "Northern Territory, Roper Gulf (S), Mataranka [
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<geoCoordinate degrees="14.93333333" direction="south" orientation="latitude" precision="1" value="-14.933333">-14.93333333</geoCoordinate>
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,
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<geoCoordinate degrees="133.0666667" direction="east" orientation="longitude" precision="1" value="133.06667">133.0666667</geoCoordinate>
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], 1.3.1967, Upton, M.S." (ANIC); 1 ex., "NT Davenport Murchison Ranges, Barkly (S), 15 miles N of Tennant Ck. [-19.4, 134.1833333], 27.2.1967, Upton, M.S." (ANIC); 1 ex., "NT Brock Creek, Burnside [Brocks Creek] [
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<geoCoordinate degrees="13.46667" direction="south" orientation="latitude" precision="1" value="-13.46667">-13.46667</geoCoordinate>
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,
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<geoCoordinate degrees="131.4167" direction="east" orientation="longitude" precision="5" value="131.4167">131.4167</geoCoordinate>
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], 27.3.1929, Campbell, T.G." (ANIC); 1 ex., "Howard Springs, Litchfield (M) [-12.45, 131.05], 27.1.1968, Matthews, E." (ANIC); 1 ex., "Australia N.T. Kununurra m 50 23.VIII.1996 P.M. Giachino legit" (CLH); 1 ex., "Australia N.T. Umbrawarra Gorge Campground, ca. 40 km S Pine Creek, 27.IV.-18.V.2009, S
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<geoCoordinate degrees="13" direction="south" minutes="58" orientation="latitude" precision="1" seconds="01.4" value="-13.967055">13°58'01.4"S</geoCoordinate>
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,
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<geoCoordinate degrees="131" direction="east" minutes="41" orientation="longitude" precision="1" seconds="57.2" value="131.69922">131°41'57.2"E</geoCoordinate>
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, 163 m, NF, leg. M. Langer" (CLH); 2 exs.,"Australia: N.T./ ca. 30 km S Daly Waters und ca. 90 km N Elliott am Stuart Hwy 04.IV.2011 LF H = 269 m
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<geoCoordinate degrees="16" direction="south" minutes="46" orientation="latitude" precision="1" seconds="38.8" value="-16.777445">16°46'38.8"S</geoCoordinate>
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,
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<geoCoordinate degrees="133" direction="east" minutes="25" orientation="longitude" precision="1" seconds="53.8" value="133.43161">133°25'53.8"E</geoCoordinate>
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leg. Michael Langer" (CLH); 11 exs., "Australia: NT, Litchfield NP, TJAYNERA FALLS,
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<geoCoordinate degrees="13" direction="south" minutes="15" orientation="latitude" precision="925" value="-13.25">13°15'S</geoCoordinate>
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,
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<geoCoordinate degrees="130" direction="east" minutes="44" orientation="longitude" precision="925" value="130.73334">130°44'E</geoCoordinate>
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, 63 m, S.
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<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
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leg., 20
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<normalizedToken originalValue="–27.XI.2008”">-27.XI.2008"</normalizedToken>
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(NMPC); 5 exs., "AUSTRALIA: NT, 25 km S of KATHERINE, rd to Kununurra,
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<geoCoordinate degrees="14" direction="south" minutes="44" orientation="latitude" precision="925" value="-14.733334">14°44'S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="01" orientation="longitude" precision="925" value="132.01666">132°01'E</geoCoordinate>
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, 100 m, S.
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<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
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leg., 17.
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<normalizedToken originalValue="–31.XII.2008”">-31.XII.2008"</normalizedToken>
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(NMPC); 12 exs., "AUSTRALIA: NT, 25 km S of KATHERINE, nr Cutta Cutta caves,
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<geoCoordinate degrees="14" direction="south" minutes="31" orientation="latitude" precision="925" value="-14.516666">14°31'S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="25" orientation="longitude" precision="925" value="132.41667">132°25'E</geoCoordinate>
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, 168 m, S.
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<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
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leg., 23.
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<normalizedToken originalValue="–31.XII.2008”">-31.XII.2008"</normalizedToken>
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(NMPC); 1 ex., "Australia: NT, Nitmiluk NP, Edit Falls,
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<geoCoordinate degrees="14" direction="south" minutes="10" orientation="latitude" precision="925" value="-14.166667">14°10'S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="06" orientation="longitude" precision="925" value="132.1">132°06'E</geoCoordinate>
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, 37 m, S.
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<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
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leg., 3.XII.2008" (NMPC); 2 exs., "AUSTR. NT, Mataranka,
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<geoCoordinate degrees="14" direction="south" minutes="53" orientation="latitude" precision="925" value="-14.883333">14°53'S</geoCoordinate>
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,
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<geoCoordinate degrees="132" direction="east" minutes="01" orientation="longitude" precision="925" value="132.01666">132°01'E</geoCoordinate>
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, 18.12.08, 148 m, Sv.
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<normalizedToken originalValue="Bílý">Bily</normalizedToken>
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leg." (NMPC); 70 exs., "Australia: NT, 70 km SW of Mataranka,
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<geoCoordinate degrees="15" direction="south" minutes="19" orientation="latitude" precision="925" value="-15.316667">15°19'S</geoCoordinate>
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,
|
||
<geoCoordinate degrees="132" direction="east" minutes="50" orientation="longitude" precision="925" value="132.83333">132°50'E</geoCoordinate>
|
||
, 190 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 22.
|
||
<normalizedToken originalValue="–23.XII.2008”">-23.XII.2008"</normalizedToken>
|
||
(NMPC); 1 ex., "AUSTRALIA: NT, West Macdonnell Range NP, SIMPSON GAP,
|
||
<geoCoordinate degrees="23" direction="south" minutes="40" orientation="latitude" precision="925" value="-23.666666">23°40'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="133" direction="east" minutes="43" orientation="longitude" precision="925" value="133.71666">133°43'E</geoCoordinate>
|
||
, 600 m, S.
|
||
<normalizedToken originalValue="Jákl">Jakl</normalizedToken>
|
||
leg., 3.
|
||
<normalizedToken originalValue="–5.I.2009”">-5.I.2009"</normalizedToken>
|
||
(NMPC).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Queensland</emphasis>
|
||
: 1 ex., "Cardstone [
|
||
<geoCoordinate degrees="17.76666667" direction="south" orientation="latitude" precision="1" value="-17.766666">-17.76666667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="145.5833333" direction="east" orientation="longitude" precision="1" value="145.58333">145.5833333</geoCoordinate>
|
||
], 16.12.1965, Hyde, K." (ANIC); 1 ex., "Mary Creek [-16.55, 145.2], 4.12.1968, Britton, E.B. & Misko, S." (ANIC); 1 ex., "Cairns [
|
||
<geoCoordinate degrees="16.91667" direction="south" orientation="latitude" precision="1" value="-16.91667">-16.91667</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="145.7667" direction="east" orientation="longitude" precision="5" value="145.7667">145.7667</geoCoordinate>
|
||
] Taylor, F.H." (ANIC); 1 ex., "Queensland, Whitsunday, Mimosa Motel, 3 m. S Bowen [-20.01667, 148.25], 23.3.1967, Brooks, J.G." (ANIC); 1 ex., "Townsville [
|
||
<geoCoordinate degrees="19.26667023" direction="south" orientation="latitude" precision="1" value="-19.26667">-19.26667023</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="146.8166656" direction="east" orientation="longitude" precision="1" value="146.81667">146.8166656</geoCoordinate>
|
||
], Taylor, F.H." (ANIC); 36 exs., "Australia/Queensland 30 km E Normanton riverside at light, 1.1996, S. Lamond leg. Coll. Hendrich" (CLH); 31 exs., "Australia/ Queensland 20 km E of Normanton February 1996, at light, Steven Lamond leg. Coll. L.Hendrich" (CLH); 4 exs., "Bowen N.Q. 23/III/1962 A. & M. Walford-Huggins" (CLH); 10 exs., "Australia, QL, Rollingstone, 20 km NW Townsville, 20 m, 17.I.1993, leg. G. Wewalka (7)" (CGW); 3 exs., "Australia, QL, Dalrymple, 30 km N Charters Towers, 300 m, 18.I.1993, leg. G. Wewalka (10, 11)" (CGW); 1 ex., "QL, 15 km S Ogmore, 24.I.2012, leg. S. Prepsl" (CGW); 3 exs., "Australia, Queensland, Horn Islet, Sir Edward Pellew Group, 15. February 1968-21. February 1964, B. Cantrell" (QM); 1 ex., "Australia, Queensland, Iron Range, Cape York 23. April 1966, G. Monteith" (QM); 2 exs., "Australia, Queensland, CYP Batavia Downs,
|
||
<geoCoordinate degrees="12.40" direction="south" orientation="latitude" precision="555" value="-12.4">12.40S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="142.40" direction="east" orientation="longitude" precision="555" value="142.4">142.40E</geoCoordinate>
|
||
, 3. March 1993-10. March 1993, at light, I. Cunningham" (QDPIB); 6 exs., "Australia, Queensland, Tolga, 26. Jan.-31. Jan. 1981, at light, N. Gough" (QDPIB); 1 ex., "Australia, Queensland, Burketown 31. Jan. 1992, at light, B.M. Waterhouse, J.F. Grimshaw" (QDPIB); 1 ex., "Australia, Queensland Danbulla S.F. via Yungaburra 13. February 1992 at light, Storey, De Faveri & Huwer" (QDPIB); 1 ex., "Australia, Queensland, Normanton, 4. September 1982, at light, R.I. Storey" (QDPIB); 1 ex., "Australia Queensland Kauri Creek, Para Grass, 30. August 1995, B. Hebert" (QDPIB); 1 ex., "Australia, Queensland, Ingham, 6. March 1984 at light, K.H. Halfpapp" (QDPIB); 4 exs., "AUSTRALIA Qld. Windsor Tableland Feb 6-8/91 Larson & Storey" (ZSM); 2 exs., "N Queensland, 14.1.2000, Kuranda, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 1 ex., "N Queensland, 9.2.2000, Kuranda, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 1 ex., "N Queensland, 7.2.2000, Mt. Carbine, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC); 1 ex., "N Queensland, 13.2.2000, Undara, Sv.
|
||
<normalizedToken originalValue="Bílý">Bily</normalizedToken>
|
||
leg." (NMPC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="description">
|
||
<paragraph pageId="0" pageNumber="81">Description of male.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Body shape:</emphasis>
|
||
In dorsal view oval, almost ovoid, broadest in basal third of elytra, moderately convex. Body outline without discontinuity between pronotum and elytra. Head relatively broad; anterior margin of clypeus truncate. Pronotum broadest between posterior angles, lateral margins moderately curved. Base of elytra as broad as pronotal base; lateral margins of elytra moderately curved (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Copelatus marginatus, female. Total length 5.8 mm." figureDoi="10.3897/zookeys.889.39090.figure6" httpUri="https://binary.pensoft.net/fig/356475" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure6">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.889.39090.figure6" httpUri="https://binary.pensoft.net/fig/356475" pageId="0" pageNumber="81" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">Figure 6.</emphasis>
|
||
Habitus and colouration of
|
||
<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus marginatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="marginatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Copelatus marginatus</emphasis>
|
||
</taxonomicName>
|
||
, female. Total length 5.8 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" pageId="0" pageNumber="81">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">Colouration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body dark red-brown to black, clypeus anteriorly, sides of pronotum broadly and elytra laterally (including epipleura) paler, ventral side and appendages testaceous (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Copelatus marginatus, female. Total length 5.8 mm." figureDoi="10.3897/zookeys.889.39090.figure6" httpUri="https://binary.pensoft.net/fig/356475" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure6">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Dorsal surface sculpture:</emphasis>
|
||
Whole surface almost matt (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus and colouration of Copelatus marginatus, female. Total length 5.8 mm." figureDoi="10.3897/zookeys.889.39090.figure6" httpUri="https://binary.pensoft.net/fig/356475" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure6">Fig. 6</figureCitation>
|
||
). Head uniformly microreticulated, reticulation composed of moderately deeply impressed meshes. Punctation composed of coarse setigerous punctures, and very small punctures spreading sparsely on surface; rows of coarse punctures present around inner margin of eyes and in small depression anterolaterally of eyes. Pronotum with lateral beading very thin and indistinct. Microreticulation and punctation similar to that of head; row of coarse setigerous punctures present along anterior margin, basal margin (except for basomedially), and laterally close to sides. Elytra with microreticulation similar to that of head and pronotum, but less impressed. Serial punctures on elytra rather weak. On each elytron six moderately impressed thin discal and one submarginal longitudinal striae, progressively closer towards sides; stria 1 (sutural stria) reduced to apical fourth, striae 2 and 4 complete, striae 3 and 5 a little shorter basally. Submarginal stria short, expanding from behind middle of elytron to end of stria 6.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Antennae and legs:</emphasis>
|
||
Antenna with antennomeres long and slender. Protibia modified, distinctly broadened anteriorly. Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive discs on their ventral side; claws simple.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Ventral part:</emphasis>
|
||
Finely microreticulated, with intermixed, sparsely distributed, very small punctures. Prosternal process shortly lanceolate, apex obtuse; distinctly bordered laterally. Lateral parts of metaventrite tongue-shaped, slender. Metacoxal lines close, evenly and moderately diverging anteriorly, ending short of metasternum. Metacoxae and abdominal ventrites 1-3 with numerous longitudinal or oblique striae.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Male genitalia:</emphasis>
|
||
Median lobe narrow, simple, tapering and slightly curved downwards apically in lateral view (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Copelatus marginatus A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure15" httpUri="https://binary.pensoft.net/fig/356484" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure15">Fig. 15A, B, C</figureCitation>
|
||
). Shape of paramere broadly triangular, with weak, relatively short setae, mainly along dorsal margin of subdistal part (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Copelatus marginatus A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.889.39090.figure15" httpUri="https://binary.pensoft.net/fig/356484" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure15">Fig. 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="female">
|
||
<paragraph pageId="0" pageNumber="81">Female.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">Similar to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae. Pronotum basolaterally with numerous short longitudinal strioles. Elytra between striae and side with numerous short longitudinal strioles.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="description">
|
||
<paragraph pageId="0" pageNumber="81">Measurements.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">TL = 5.7-6.0 mm; TL-H = 5.3-5.6 mm; MW = 2.7-2.8 mm.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="variability">
|
||
<paragraph pageId="0" pageNumber="81">Variability.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">All specimens studied are rather uniform and vary only in body length. In some specimens, the submarginal stria is reduced to few elongate grooves.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
The species is close to
|
||
<taxonomicName lsidName="C. tenebrosus" pageId="0" pageNumber="81" rank="species" species="tenebrosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. tenebrosus</emphasis>
|
||
</taxonomicName>
|
||
but can be easily separated by the larger size, much shorter inner striae of elytra, and the form of the median lobe.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="distribution">
|
||
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
The species occurs in tropical and subtropical northern and central Australia, along the east coast south to Brisbane (WA, NT, QLD) (
|
||
<figureCitation captionStart="Figure 25" captionStartId="F24" captionText="Figure 25. Distribution of Copelatus marginatus." figureDoi="10.3897/zookeys.889.39090.figure25" httpUri="https://binary.pensoft.net/fig/356493" pageId="0" pageNumber="81" tableDoi="10.3897/zookeys.889.39090.figure25">Fig. 25</figureCitation>
|
||
). Additionally,
|
||
<taxonomicName lsidName="C. marginatus" pageId="0" pageNumber="81" rank="species" species="marginatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marginatus</emphasis>
|
||
</taxonomicName>
|
||
was recorded from New Caledonia (South Province) and West Samoa by
|
||
<bibRefCitation author="Wewalka, G" editor="Jaech, MA" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" refId="B39" refString="Wewalka, G, Balke, M, Hendrich, L, Jaech, MA, Balke, M, 2010. Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1).Monographs on Coleoptera 3, 45-128." title="Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1). Monographs on Coleoptera 3, 45 - 128." year="2010">Wewalka et al. (2010)</bibRefCitation>
|
||
.
|
||
<bibRefCitation author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="0" pageNumber="81" refId="B41" refString="Zimmermann, A, 1927. Dytiscidae. Insects of Samoa and Other Samoan Terrestrial Arthropoda, part IV, fasc. 1, 15-19." title="Dytiscidae. Insects of Samoa and Other Samoan Terrestrial Arthropoda, part IV, fasc. 1, 15 - 19." year="1927">Zimmermann (1927</bibRefCitation>
|
||
: 17) reported the species from Tonga (two specimens from Tonga: Nukualofa, 22.xi.1925). He wrote also that the presence of the species in Samoa is confirmed by material of Dr K. Friederichs (
|
||
<bibRefCitation author="Friederichs, K" journalOrPublisher="Zootaxa" pageId="0" pageNumber="81" refId="B8" refString="Friederichs, K, 1922. Die von Dr. K. Friederichs in Samoa and Indochina gesammelten Kaefer. Archiv fuer Naturgeschichte 88A(10): 147-159." title="Die von Dr. K. Friederichs in Samoa and Indochina gesammelten Kaefer. Archiv fuer Naturgeschichte 88 A (10): 147 - 159." year="1922">Friederichs 1922</bibRefCitation>
|
||
: 148: Samoa, five specimens). Later,
|
||
<taxonomicName lsidName="C. marginatus" pageId="0" pageNumber="81" rank="species" species="marginatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marginatus</emphasis>
|
||
</taxonomicName>
|
||
was reported from Samoa (Upolu), Fiji (Viti Levu and Vanua Levu) and New Guinea (Dorey, now Manokwari, Indonesia: West Papua Province) by Balfour-Browne (1945: 106 and 113) which was repeated by some later authors (
|
||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Izvestija na Zoologitjeskija Institut s Musei Sofia" pageId="0" pageNumber="81" pagination="5 - 45" refId="B9" refString="Gueorguiev, VB, 1968. Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae). Izvestija na Zoologitjeskija Institut s Musei Sofia 28: 5 - 45" title="Essai de classification des coleopteres Dytiscidae. I. Tribus Copelatini (Colymbetinae)." volume="28" year="1968">
|
||
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||
1968
|
||
</bibRefCitation>
|
||
: 21;
|
||
<bibRefCitation author="Gueorguiev, VB" journalOrPublisher="Frustula Entomologica (Nuova Serie)" pageId="0" pageNumber="81" pagination="147 - 166" refId="B10" refString="Gueorguiev, VB, Rocchi, S, 1993. Contributo alla conoscenza dei Dytiscidae della Nuova Guinea (Coleoptera). Frustula Entomologica (Nuova Serie) 15: 147 - 166" title="Contributo alla conoscenza dei Dytiscidae della Nuova Guinea (Coleoptera)." volume="15" year="1993">
|
||
<normalizedToken originalValue="Guéorguiev">Gueorguiev</normalizedToken>
|
||
and Rocchi 1993
|
||
</bibRefCitation>
|
||
: 159;
|
||
<bibRefCitation author="Wewalka, G" editor="Jaech, MA" journalOrPublisher="Identification Ecology Guide" pageId="0" pageNumber="81" refId="B39" refString="Wewalka, G, Balke, M, Hendrich, L, Jaech, MA, Balke, M, 2010. Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1).Monographs on Coleoptera 3, 45-128." title="Dytiscidae: Copelatinae (Coleoptera). In: JaechMABalkeM (Eds) Water beetles of New Caledonia (part 1). Monographs on Coleoptera 3, 45 - 128." year="2010">Wewalka et al. 2010</bibRefCitation>
|
||
: 51). The records from Fiji and New Guinea are in need of confirmation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="81" type="habitat">
|
||
<paragraph pageId="0" pageNumber="81">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="81">
|
||
Most specimens were collected during or just after the rainy season when the beetles can be collected in seasonal flood meadows along rivers and creeks, shallow roadside ditches and swampy areas. At that time of the year,
|
||
<taxonomicName lsidName="C. marginatus" pageId="0" pageNumber="81" rank="species" species="marginatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="81">C. marginatus</emphasis>
|
||
</taxonomicName>
|
||
is often attracted in larger numbers to light (e.g., Normanton). In north-eastern Queensland, the species was collected in a small roadside pool and in isolated pools in seasonal fingertip tributary streams, all in closed forest (
|
||
<bibRefCitation author="Larson, DJ" journalOrPublisher="Proceedings of the Royal Society of Queensland" pageId="0" pageNumber="81" pagination="47 - 63" refId="B20" refString="Larson, DJ, 1993. Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species. Proceedings of the Royal Society of Queensland 103: 47 - 63" title="Ecology of tropical Australian Hydradephaga (Insecta: Coleoptera). Part 1. Natural history and distribution of northern Queensland species." volume="103" year="1993">Larson 1993</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |