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<mods:title id="8D659B647651ABF19F985D12B39E15AA">A revision of the genus Phellopsis LeConte (Coleoptera: Zopheridae)</mods:title>
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<heading id="A66881A2FFADFFBF1DC17C14FD89FBA4" bold="true" box="[151,530,1071,1097]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="3A9F4D4DFFADFFBF1DC17C14FD89FBA4" authority="Heyden" authorityName="Heyden" box="[151,530,1071,1097]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="CFEBEADCFFADFFBF1DC17C14FD89FBA4" bold="true" box="[151,530,1071,1097]" pageId="19" pageNumber="20">
<emphasis id="CFEBEADCFFADFFBF1DC17C14FE0CFBA4" bold="true" box="[151,407,1071,1097]" italics="true" pageId="19" pageNumber="20">Phellopsis amurensis</emphasis>
(Heyden)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FD2036CEFFADFFBF1DC17C69FE3FFB81" blockId="19.[151,530,1071,1132]" box="[151,420,1106,1132]" pageId="19" pageNumber="20">
(
<figureCitation id="65A42A4BFFADFFBF1DC97C69FF76FB81" box="[159,237,1106,1132]" captionStart="FIGURE 1. A" captionStartId="5.[151,255,1929,1953]" captionTargetBox="[186,1365,204,1928]" captionTargetId="figure@5.[151,1436,152,1956]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of specimen records for North American Phellopsis spp. P. p o rc a t a, green circles; P. obcordata red diamonds. B. Map of specimen records for Asian Phellopsis spp. P. yulongensis red diamonds; P. chinensis green circles; P. a m u re n s i s red cross; P. suberea green triangles." httpUri="https://zenodo.org/record/180606/files/figure.png" pageId="19" pageNumber="20">Figs. 1</figureCitation>
B, 3A, 4A, 6A)
</paragraph>
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<taxonomicName id="3A9F4D4DFFADFFBF1DC17C99FD35FB54" authority="Heyden, 1885: 306" authorityName="Heyden" authorityPageNumber="306" authorityYear="1885" box="[151,686,1186,1209]" class="Insecta" family="Zopheridae" genus="Pseudonosoderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="amurense">
<emphasis id="CFEBEADCFFADFFBF1DC17C99FE56FB54" box="[151,461,1186,1209]" italics="true" pageId="19" pageNumber="20">Pseudonosoderma amurense</emphasis>
<treatmentCitation id="7C3E10DFFFADFFBF1C8E7C99FD35FB54" author="Heyden" box="[472,686,1186,1209]" page="306" pageId="19" pageNumber="20" year="1885">
<bibRefCitation id="990E4B3FFFADFFBF1C8E7C99FDEEFB54" author="Heyden" box="[472,629,1186,1209]" pageId="19" pageNumber="26" refString="Heyden, L. (1885) Weitere Beitrage zur Coleoperen-Fauna des Amur-Gebietes. Deutsche Entomologische Zeitschrift, Berlin, 29, 299 - 307, pl. 4." type="journal article" year="1885">Heyden, 1885</bibRefCitation>
: 306
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</taxonomicName>
.
<treatmentCitation id="7C3E10DFFFADFFBF1FEB7C99FC33FB54" author="Semenow" box="[701,936,1186,1209]" page="499" pageId="19" pageNumber="20" year="1893">
<bibRefCitation id="990E4B3FFFADFFBF1FEB7C99FCF5FB54" author="Semenow" box="[701,878,1186,1209]" pageId="19" pageNumber="28" refString="Semenow, A. (also cited as Semenov) (1893) Coleoptera asiatica nova. Horae Societatis Entomologicae Rossicae, variis sermonibus in Rossia usitatis editae, 27, 494 - 507." type="journal article" year="1893">Semenow, 1893</bibRefCitation>
: 499
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.
<treatmentCitation id="7C3E10DFFFADFFBF1EE07C99FB33FB54" author="Champion" box="[950,1192,1186,1209]" page="114" pageId="19" pageNumber="20" year="1894">
<bibRefCitation id="990E4B3FFFADFFBF1EE07C99FBF4FB54" author="Champion" box="[950,1135,1186,1209]" pageId="19" pageNumber="26" refString="Champion, G. C. (1894) [Note] The genus Pseudonosoderma, v. Heyd. Entomologist's Monthly Magazine, 30, 114." type="journal article" year="1894">Champion, 1894</bibRefCitation>
: 114
</treatmentCitation>
. Fairmaire, 1894: C1.
<treatmentCitation id="7C3E10DFFFADFFBF1D917CF9FE1FFB34" author="Lewis" box="[199,388,1218,1241]" page="447" pageId="19" pageNumber="20" year="1895">
<bibRefCitation id="990E4B3FFFADFFBF1D917CF9FED5FB34" author="Lewis" box="[199,334,1218,1241]" pageId="19" pageNumber="27" refString="Lewis, G. (1895) On the Cistelidae and other heteromerous species of Japan. Annals and Magazine of Natural History (London) 6, 422 - 448, pl. viii." type="journal article" year="1895">Lewis, 1895</bibRefCitation>
: 447
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.
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</paragraph>
<paragraph id="FD2036CEFFADFFBF1DC17CD9FA9CFAF4" blockId="19.[151,1436,1186,1370]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="DD8F11E0FFADFFBF1DC17CD9FA9CFAF4" pageId="19" pageNumber="20">
<taxonomicName id="3A9F4D4DFFADFFBF1DC17CD9FDC8FB14" authority="Reitter, 1916: 130" authorityName="Reitter" authorityPageNumber="130" authorityYear="1916" box="[151,595,1250,1273]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="CFEBEADCFFADFFBF1DC17CD9FEE0FB14" box="[151,379,1250,1273]" italics="true" pageId="19" pageNumber="20">Phellopsis amurensis</emphasis>
:
<treatmentCitation id="7C3E10DFFFADFFBF1CDA7CD9FDC8FB14" author="Reitter" box="[396,595,1250,1273]" page="130" pageId="19" pageNumber="20" year="1916">
<bibRefCitation id="990E4B3FFFADFFBF1CDA7CD9FD80FB14" author="Reitter" box="[396,539,1250,1273]" pageId="19" pageNumber="28" refString="Reitter, V. E. (1916) Bestimmungstabelle der Tenebrioniden, enthaltend die Zopherini, Elenophorini, Leptodini, Stenosini und Lachnogyini. Wiener Entomologische Zeitung, 35, 129 - 171." type="journal article" year="1916">Reitter, 1916</bibRefCitation>
: 130
</treatmentCitation>
</taxonomicName>
.
<treatmentCitation id="7C3E10DFFFADFFBF1F097CD9FC8DFB14" author="Leng" box="[607,790,1250,1273]" page="223" pageId="19" pageNumber="20" year="1920">
<bibRefCitation id="990E4B3FFFADFFBF1F097CD9FD45FB14" author="Leng" box="[607,734,1250,1273]" pageId="19" pageNumber="27" refString="Leng, C. W. (1920) Catalogue of the Coleoptera of America, North of Mexico. John D. Sherman, Jr., Mount Vernon, New York. x + 470 pp." type="book" year="1920">Leng, 1920</bibRefCitation>
: 223
</treatmentCitation>
.
<treatmentCitation id="7C3E10DFFFADFFBF1E747CD9FC74FB14" author="Gebien" box="[802,1007,1250,1273]" page="668" pageId="19" pageNumber="20" year="1936">
<bibRefCitation id="990E4B3FFFADFFBF1E747CD9FC23FB14" author="Gebien" box="[802,952,1250,1273]" pageId="19" pageNumber="26" refString="Gebien, H. (1936) Katalog der Tenebrioniden (Col.: Heteromera). Teil I. (abgeschlossen August 1936) Pubblicasioni del Museo Entomologico &quot; Pietro Rossi &quot; 2, 505 - 883." type="journal article" year="1936">Gebien, 1936</bibRefCitation>
: 668
</treatmentCitation>
.
<treatmentCitation id="7C3E10DFFFADFFBF1EAA7CD9FA15FB14" author="Keleinikova" box="[1020,1422,1250,1273]" page="124" pageId="19" pageNumber="20" year="1971">
<bibRefCitation id="990E4B3FFFADFFBF1EAA7CD9FAC1FB14" author="Keleinikova" box="[1020,1370,1250,1273]" pageId="19" pageNumber="27" refString="Keleinikova, S. I. &amp; Mamaev, B. M. (1971) Phellopsis amurensis Heyd. (Coleoptera, Zopheridae) a specific wood destroyer in the South Primorski Krai. Builletin, Moskovskoe Obshchestvo Ispytatelei Prirody, Novaya Seriya. Otdel Biologicheskii, 76 (4), 124 - 128. [in Russian]" type="journal article" year="1971">Keleinikova and Mamaev, 1971</bibRefCitation>
: 124
</treatmentCitation>
128.
<treatmentCitation id="7C3E10DFFFADFFBF1DAA7D39FE4AFAF4" author="Gaedike" box="[252,465,1282,1305]" page="360" pageId="19" pageNumber="20" year="1986">
<bibRefCitation id="990E4B3FFFADFFBF1DAA7D39FE00FAF4" author="Gaedike" box="[252,411,1282,1305]" pageId="19" pageNumber="26" refString="Gaedike, H. (1986) Katalog der in den Sammlungen der Abteilung Taxonomie der Insekten des Institutes fur Pflanzenschutzforschung, Bereich Eberswalde (ehemals Deutsches Entomologisches Institut) aufbewahrten Typen-XXIV. Beitrage zur Entomologie, 36, 321 - 429." type="journal article" year="1986">Gaedike, 1986</bibRefCitation>
: 360
</treatmentCitation>
.
<treatmentCitation id="7C3E10DFFFADFFBF1C8D7D39FD27FAF4" author="Masumoto" box="[475,700,1282,1305]" page="91" pageId="19" pageNumber="20" year="1990">
<bibRefCitation id="990E4B3FFFADFFBF1C8D7D39FD08FAF4" author="Masumoto" box="[475,659,1282,1305]" pageId="19" pageNumber="27" refString="Masumoto, K. (1990) A new Phellopsis species from South Korea (Zopheridae, Coleoptera). Entomological Review of Japan, 45, 87 - 91." type="journal article" year="1990">Masumoto, 1990</bibRefCitation>
: 91
</treatmentCitation>
.
<treatmentCitation id="7C3E10DFFFADFFBF1F907D39FC15FAF4" author="Egorov" box="[710,910,1282,1305]" page="505" pageId="19" pageNumber="20" year="1992">
<bibRefCitation id="990E4B3FFFADFFBF1F907D39FCC2FAF4" author="Egorov" box="[710,857,1282,1305]" pageId="19" pageNumber="26" refString="Egorov, A. B. (1992) 94. Cem. Zopheridae, pp. 504 - 505. In: P. A. Ler, (ed.) Key to the Insects of the Russian Far East. Vol. III. Part 2. Dal'nauka, Vladivostok [in Russian]." type="book chapter" year="1992">Egorov, 1992</bibRefCitation>
: 505
</treatmentCitation>
. Ś lipiński and Lawrence, 1999: 23.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FD2036CEFFADFFBF1DC17D19FCB3FAD7" blockId="19.[151,1436,1186,1370]" box="[151,808,1314,1338]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="DD8F11E0FFADFFBF1DC17D19FCB3FAD7" box="[151,808,1314,1338]" pageId="19" pageNumber="20">
<taxonomicName id="3A9F4D4DFFADFFBF1DC17D19FDA0FAD4" authority="Masumoto, 1990: 87" authorityName="Masumoto" authorityPageNumber="87" authorityYear="1990" box="[151,571,1314,1337]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="CFEBEADCFFADFFBF1DC17D19FECFFAD4" box="[151,340,1314,1337]" italics="true" pageId="19" pageNumber="20">Phellopsis imurai</emphasis>
<treatmentCitation id="7C3E10DFFFADFFBF1C0C7D19FDA0FAD4" author="Masumoto" box="[346,571,1314,1337]" page="87" pageId="19" pageNumber="20" year="1990">
<bibRefCitation id="990E4B3FFFADFFBF1C0C7D19FD88FAD4" author="Masumoto" box="[346,531,1314,1337]" pageId="19" pageNumber="27" refString="Masumoto, K. (1990) A new Phellopsis species from South Korea (Zopheridae, Coleoptera). Entomological Review of Japan, 45, 87 - 91." type="journal article" year="1990">Masumoto, 1990</bibRefCitation>
: 87
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</taxonomicName>
.
<emphasis id="CFEBEADCFFADFFBF1F137D19FCB3FAD7" bold="true" box="[581,808,1314,1338]" pageId="19" pageNumber="20">NEW SYNONYMY</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="FD2036CEFFADFFBF1DC17D79FABBFAB7" blockId="19.[151,1436,1186,1370]" box="[151,1312,1346,1370]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="DD8F11E0FFADFFBF1DC17D79FABBFAB7" box="[151,1312,1346,1370]" pageId="19" pageNumber="20">
<taxonomicName id="3A9F4D4DFFADFFBF1DC17D79FDF7FAB4" authority="Weon et al., 2000: 119" authorityName="Weon et al." authorityPageNumber="119" authorityYear="2000" box="[151,620,1346,1369]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="subaenea">
<emphasis id="CFEBEADCFFADFFBF1DC17D79FEE9FAB4" box="[151,370,1346,1369]" italics="true" pageId="19" pageNumber="20">Phellopsis subaenea</emphasis>
<treatmentCitation id="7C3E10DFFFADFFBF1C2E7D79FDF7FAB4" author="Weon" box="[376,620,1346,1369]" page="119" pageId="19" pageNumber="20" year="2000">
<bibRefCitation id="990E4B3FFFADFFBF1C2E7D79FDA3FAB4" author="Weon" box="[376,568,1346,1369]" pageId="19" pageNumber="28" refString="Weon, G. - J., Jang, W. - Y., Oh, K. - S., Shin, S. - C. &amp; Park, J. - D. (2000) List of Species in Insect Collection of the Korean Forest Research Institute, 7. Coleoptera (except Superfamily Chrysomeloidea). Korean Forest Research Institute, Journal of Forestry Science, 63, 108 - 123." type="journal article" year="2000">
Weon
<emphasis id="CFEBEADCFFADFFBF1CEA7D79FE6AFAB4" box="[444,497,1346,1369]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2000
</bibRefCitation>
: 119
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</taxonomicName>
. (
<emphasis id="CFEBEADCFFADFFBF1F297D79FC88FAB4" box="[639,787,1346,1369]" italics="true" pageId="19" pageNumber="20">lapsus calami</emphasis>
for
<taxonomicName id="3A9F4D4DFFADFFBF1E697D79FB93FAB4" box="[831,1032,1346,1369]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="suberea">
<emphasis id="CFEBEADCFFADFFBF1E697D79FB93FAB4" box="[831,1032,1346,1369]" italics="true" pageId="19" pageNumber="20">Phellopsis suberea</emphasis>
</taxonomicName>
, MISIDENTIFICATION
<emphasis id="CFEBEADCFFADFFBF18417D79FABBFAB7" bold="true" box="[1303,1312,1346,1370]" pageId="19" pageNumber="20">)</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="B5856545FFADFFBF1DC17DB1FED2F9F1" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="FD2036CEFFADFFBF1DC17DB1FED2F9F1" blockId="19.[151,1437,1418,2004]" pageId="19" pageNumber="20">
<emphasis id="CFEBEADCFFADFFBF1DC17DB1FE82FA49" bold="true" box="[151,281,1418,1444]" pageId="19" pageNumber="20">Diagnosis:</emphasis>
This species is similar in general elytral structure to
<taxonomicName id="3A9F4D4DFFADFFBF1E287DB0FB8AFA49" box="[894,1041,1419,1444]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="obcordata">
<emphasis id="CFEBEADCFFADFFBF1E287DB0FB8AFA49" box="[894,1041,1419,1444]" italics="true" pageId="19" pageNumber="20">P. obcordata</emphasis>
</taxonomicName>
but has the lateral elytral margins serrate and suprantennal frontal ridges strongly elevated like the other Asian species. It is distinguished from these species by the more stout form, rounded elytral punctures and presence of strongly notched humeri (
<figureCitation id="65A42A4BFFADFFBF1DC97E39FF74F9F1" box="[159,239,1538,1564]" captionStart="FIGURE 3" captionStartId="10.[151,255,1971,1995]" captionTargetBox="[308,1293,353,1940]" captionTargetId="figure@10.[151,1436,325,1988]" captionTargetPageId="10" captionText="FIGURE 3. Dorsal habitus photos of the Asian Phellopsis spp. A. Phellopsis amurensis; B. P. s u b e re a; C. P. chinensis; D. P. yulongensis." httpUri="https://zenodo.org/record/180608/files/figure.png" pageId="19" pageNumber="20">Figs. 3</figureCitation>
A, 4A).
</paragraph>
</subSubSection>
<subSubSection id="B5856545FFADFFB81D907E11FCF8FD76" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="FD2036CEFFADFFBF1D907E11FD47F8D9" blockId="19.[151,1437,1418,2004]" pageId="19" pageNumber="20">
<emphasis id="CFEBEADCFFADFFBF1D907E11FE23F9A9" bold="true" box="[198,440,1578,1604]" pageId="19" pageNumber="20">Description (male):</emphasis>
Length
<quantity id="3A679B2BFFADFFBF1F4E7E11FD2BF9A9" box="[536,688,1578,1604]" metricMagnitude="-2" metricUnit="m" metricValue="1.675" metricValueMax="1.9" metricValueMin="1.45" pageId="19" pageNumber="20" unit="mm" value="16.75" valueMax="19.0" valueMin="14.5">14.519 mm</quantity>
. Dark brown to black; dorsal vestiture consisting of golden hairlike setae. Head with dorsal surface (
<figureCitation id="65A42A4BFFADFFBF1F0C7E69FD39F981" box="[602,674,1618,1644]" captionStart="FIGURE 6" captionStartId="14.[151,255,1583,1607]" captionTargetBox="[173,1415,345,1535]" captionTargetId="figure@14.[152,1436,320,1560]" captionTargetPageId="14" captionText="FIGURE 6. Dorsal head surface of the Asian Phellopsis spp. A. Phellopsis amurensis; B. P. s u b e re a; C. P. chinensis; D. P. yulongensis." httpUri="https://zenodo.org/record/180611/files/figure.png" pageId="19" pageNumber="20">Fig. 6</figureCitation>
A) with tuberculation between the frons weak; outer margin of suprantennal frontal ridges smooth; post occipital suture narrow and deep; lateral margin of epistoma anterior to suprantennal ridge longer (
<quantity id="3A679B2BFFADFFBF1CB87E99FD04F951" box="[494,671,1698,1724]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="19" pageNumber="20" unit="mm" value="0.265" valueMax="0.29" valueMin="0.24">0.240.29 mm</quantity>
); ventral surface of head with distinct raised tubercles; gula wide, narrowest point at gular pits; subgenal ridge produced apically, rounded margin; subgenal ridge shorter, not extending below eye; eye at same level as gena. Ligula subquadrate with slightly emarginate apical edge; mandibular prostheca with brush of setae at apex.
</paragraph>
<paragraph id="FD2036CEFFADFFBF1D907F79FEBDF841" blockId="19.[151,1437,1418,2004]" pageId="19" pageNumber="20">Pronotum evenly tuberculate, apical margins acutely rounded; paired elevations on apical margin of pronotum distinctly separated along midline; lateral margin weakly bisinuate; hypomeron lacking intertuberculate setae.</paragraph>
<paragraph id="FD2036CEFFADFFB81D907F81FC30FE16" blockId="19.[151,1437,1418,2004]" lastBlockId="20.[151,1437,152,1747]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">Lateral profile of elytra flattened medially in dorsal plane; elytral humerus flattened with a distinct notch; lateral margins of elytra serrate; scutellum oval set well below elytral ridges; scutellary striole distinct and deep; 1012 rounded elytral punctures between striole and start of nodule at apical declivity; 3rd and 4th rows of punctures forming distinct rows at midpoint; ridge in 5th elytral interval starting at mid-point of meta-sternum and extending to the apical margin of ventrite 3, distinct gap between ridge and nodule of declivity; nodule in 3rd elytral interval connected to ridge; median subapical nodule larger than lateral one; nodule at apex of elytron smaller than median subapical nodule; paired subapical nodules of declivity overlapping for a significant width in the same lateral plane, almost entire width. Mesepisternum with regular small punctures; metasternum bituberculate. Ventrites 13 flat, with numerous uniformly distributed small tubercles. Tarsus with ventral setae modified as spurs, distinctly heavier than dorsal setae.</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907A32FF6EFDA6" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">Aedeagus with basal stop of parameres only moderately enlarged, shorter in length; apical margin of stop smooth.</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907A62FB95FD9E" blockId="20.[151,1437,152,1747]" box="[198,1038,601,627]" pageId="20" pageNumber="21">FEMALE: Similar to male except lacking setose pit on the submentum.</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907ABAFCF8FD76" blockId="20.[151,1437,152,1747]" box="[198,867,641,667]" pageId="20" pageNumber="21">
LARVA: Described by
<bibRefCitation id="990E4B3FFFAAFFB81C817ABAFCC4FD76" author="Keleinikova" box="[471,863,641,667]" pageId="20" pageNumber="27" refString="Keleinikova, S. I. &amp; Mamaev, B. M. (1971) Phellopsis amurensis Heyd. (Coleoptera, Zopheridae) a specific wood destroyer in the South Primorski Krai. Builletin, Moskovskoe Obshchestvo Ispytatelei Prirody, Novaya Seriya. Otdel Biologicheskii, 76 (4), 124 - 128. [in Russian]" type="journal article" year="1971">Keleinikova and Mamaev (1971)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B5856545FFAAFFB81D907A93FE56FC5E" pageId="20" pageNumber="21" type="distribution">
<paragraph id="FD2036CEFFAAFFB81D907A93FF6BFCFE" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
<emphasis id="CFEBEADCFFAAFFB81D907A93FE4FFD2F" bold="true" box="[198,468,680,706]" pageId="20" pageNumber="21">
Distribution (
<figureCitation id="65A42A4BFFAAFFB81C397A93FE2EFD2F" box="[367,437,680,706]" captionStart="FIGURE 1. A" captionStartId="5.[151,255,1929,1953]" captionTargetBox="[186,1365,204,1928]" captionTargetId="figure@5.[151,1436,152,1956]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of specimen records for North American Phellopsis spp. P. p o rc a t a, green circles; P. obcordata red diamonds. B. Map of specimen records for Asian Phellopsis spp. P. yulongensis red diamonds; P. chinensis green circles; P. a m u re n s i s red cross; P. suberea green triangles." httpUri="https://zenodo.org/record/180606/files/figure.png" pageId="20" pageNumber="21">Fig. 1</figureCitation>
B)
</emphasis>
: Probably relatively widely distributed geographically from the coastal mountains of the Amur River valley in
<collectingCountry id="8588765EFFAAFFB81F577AEAFDC9FD06" box="[513,594,721,747]" name="Russia" pageId="20" pageNumber="21">Russia</collectingCountry>
, through southern portion of the Korean Peninsula (including
<collectingCountry id="8588765EFFAAFFB8180E7AEAFF7FFCFE" name="North Korea" pageId="20" pageNumber="21">North Korea</collectingCountry>
).
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907B1BFE56FC5E" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
<emphasis id="CFEBEADCFFAAFFB81D907B1BFE7EFCD7" bold="true" box="[198,485,800,826]" pageId="20" pageNumber="21">Recorded distribution:</emphasis>
Complete label data may be found in
<bibRefCitation id="990E4B3FFFAAFFB81EF97B1AFBD7FCD6" author="Foley" box="[943,1100,801,827]" pageId="20" pageNumber="26" refString="Foley, I. A. (2006) A Review of the Ironclad Beetle's of the World (Coleoptera Zopheridae: Phellopsini and Zopherini). Master's Thesis, Montana State University. 218 pp." type="book" year="2006">Foley (2006)</bibRefCitation>
. A summary of the distribution from the
<specimenCount id="EB99FD47FFAAFFB81C167B72FE7AFC8E" box="[320,481,841,867]" pageId="20" pageNumber="21" type="generic">19 specimens</specimenCount>
examined (BMNH, HNHM, NHMB, ZIN) is presented here,
<collectingCountry id="8588765EFFAAFFB819867B72FAA6FC8E" box="[1232,1341,841,867]" name="Russia" pageId="20" pageNumber="21">RUSSIA</collectingCountry>
: coastal Mountains from Amur River valley to Khrebet Sikhote Alin Mountains north of Vladivostok.
<collectingCountry id="8588765EFFAAFFB8186C7B4AFE9AFC5E" name="South Korea" pageId="20" pageNumber="21">SOUTH KOREA</collectingCountry>
: Chonwangbong.
</paragraph>
</subSubSection>
<subSubSection id="B5856545FFAAFFB81D907BFBFABCF93E" pageId="20" pageNumber="21" type="description">
<paragraph id="FD2036CEFFAAFFB81D907BFBFA98FC36" blockId="20.[151,1437,152,1747]" box="[198,1283,960,987]" pageId="20" pageNumber="21">
<emphasis id="CFEBEADCFFAAFFB81D907BFBFED8FC37" bold="true" box="[198,323,960,986]" pageId="20" pageNumber="21">Variation:</emphasis>
Little morphological variation is seen in specimens throughout the species range.
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907BD3FBC8FBEE" blockId="20.[151,1437,152,1747]" box="[198,1107,1000,1027]" pageId="20" pageNumber="21">
<emphasis id="CFEBEADCFFAAFFB81D907BD3FE8CFBEF" bold="true" box="[198,279,1000,1026]" pageId="20" pageNumber="21">
<typeStatus id="2224886CFFAAFFB81D907BD3FE96FBEF" box="[198,269,1000,1026]" pageId="20" pageNumber="21">Types</typeStatus>
:
</emphasis>
<taxonomicName id="3A9F4D4DFFAAFFB81C767BD2FD40FBEE" authority="Heyden." authorityName="Heyden." box="[288,731,1001,1027]" class="Insecta" family="Zopheridae" genus="Pseudonosoderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="amurense">
<emphasis id="CFEBEADCFFAAFFB81C767BD2FDEAFBEF" box="[288,625,1001,1026]" italics="true" pageId="20" pageNumber="21">Pseudonosoderma amurense</emphasis>
Heyden.
</taxonomicName>
<typeStatus id="2224886CFFAAFFB81FB47BD2FCD4FBEE" box="[738,847,1001,1027]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
in DEI not examined.
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907C2AFB18FB4E" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB81D907C2AFDBAFBC6" authority="Masumoto." authorityName="Masumoto." box="[198,545,1041,1067]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="CFEBEADCFFAAFFB81D907C2AFE0EFBC7" box="[198,405,1041,1066]" italics="true" pageId="20" pageNumber="21">Phellopsis imurai</emphasis>
Masumoto.
</taxonomicName>
<typeStatus id="2224886CFFAAFFB81F717C2AFD22FBC6" box="[551,697,1041,1067]" pageId="20" pageNumber="21" type="paratype">PARATYPE</typeStatus>
ɗ in HNHM with median lobe on card, missing both mesotarsi and left meta-leg. Chonwangbong, 950-;
<quantity id="3A679B2BFFAAFFB81FD37C02FD40FBBE" box="[645,731,1081,1107]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">1500m</quantity>
, Mt. Chiri-san; Kyongsangnam-do;
<date id="8921100EFFAAFFB819CE7C02FA94FBBE" box="[1176,1295,1081,1107]" pageId="20" pageNumber="21" value="1989-06-07">7.VI.1989</date>
[
<collectingCountry id="8588765EFFAAFFB818747C02FA0AFBBE" box="[1314,1425,1081,1107]" name="South Korea" pageId="20" pageNumber="21">KOREA</collectingCountry>
]; Imura &amp; Mizunuma lg/ pink rectangle label “
<typeStatus id="2224886CFFAAFFB81FE87C5AFCB6FB96" box="[702,813,1121,1147]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
”; handwritten “
<emphasis id="CFEBEADCFFAAFFB81EBC7C5AFBEFFB97" box="[1002,1140,1121,1146]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB81EBC7C5AFBF1FB97" box="[1002,1130,1121,1146]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phellopsis</taxonomicName>
</emphasis>
;
<taxonomicName id="3A9F4D4DFFAAFFB819D07C5AFA07FB96" authority="Masumoto. The" authorityName="Masumoto. The" box="[1158,1436,1121,1147]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="CFEBEADCFFAAFFB819D07C5AFB4FFB97" box="[1158,1236,1121,1146]" italics="true" pageId="20" pageNumber="21">imurai</emphasis>
Masumoto. The
</taxonomicName>
<typeStatus id="2224886CFFAAFFB81DC17CB2FF65FB4E" box="[151,254,1161,1187]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
is deposited in the National Science Museum, Tokyo and was not examined.
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907C8BFD28FAAF" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
<emphasis id="CFEBEADCFFAAFFB81D907C8BFE8DFB27" bold="true" box="[198,278,1200,1226]" pageId="20" pageNumber="21">Notes:</emphasis>
While correctly identifying the generic synonymy of
<taxonomicName id="3A9F4D4DFFAAFFB81EC37C8AFAD6FB26" authority="Champion (1894)" authorityName="Champion" authorityYear="1894" box="[917,1357,1201,1227]" class="Insecta" family="Zopheridae" genus="Pseudonosoderma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="CFEBEADCFFAAFFB81EC37C8AFBF6FB27" box="[917,1133,1201,1226]" italics="true" pageId="20" pageNumber="21">Pseudonosoderma</emphasis>
,
<bibRefCitation id="990E4B3FFFAAFFB8192B7C8AFAD6FB26" author="Champion" box="[1149,1357,1201,1227]" pageId="20" pageNumber="26" refString="Champion, G. C. (1894) [Note] The genus Pseudonosoderma, v. Heyd. Entomologist's Monthly Magazine, 30, 114." type="journal article" year="1894">Champion (1894)</bibRefCitation>
</taxonomicName>
incorrectly synonymized
<emphasis id="CFEBEADCFFAAFFB81CDF7CE2FE64FB1F" box="[393,511,1241,1266]" italics="true" pageId="20" pageNumber="21">P. s u b e re a</emphasis>
with
<emphasis id="CFEBEADCFFAAFFB81F1C7CE2FD73FB1F" box="[586,744,1241,1266]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB81F1C7CE2FD7FFB1F" box="[586,740,1241,1266]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="amurensis">P. amurensis</taxonomicName>
.
</emphasis>
<bibRefCitation id="990E4B3FFFAAFFB81FA77CE2FC0BFB1E" author="Lewis" box="[753,912,1241,1267]" pageId="20" pageNumber="27" refString="Lewis, G. (1895) On the Cistelidae and other heteromerous species of Japan. Annals and Magazine of Natural History (London) 6, 422 - 448, pl. viii." type="journal article" year="1895">Lewis (1895)</bibRefCitation>
corrected a previous misidentification of
<taxonomicName id="3A9F4D4DFFAAFFB818D07CE2FE94FAF7" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="CFEBEADCFFAAFFB818D07CE2FE94FAF7" italics="true" pageId="20" pageNumber="21">P. amurensis</emphasis>
</taxonomicName>
as a
<typeStatus id="2224886CFFAAFFB81C057D3AFE35FAF6" box="[339,430,1281,1307]" pageId="20" pageNumber="21" type="syntype">syntype</typeStatus>
of
<taxonomicName id="3A9F4D4DFFAAFFB81C8A7D3AFDCCFAF7" box="[476,599,1281,1306]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="suberea">
<emphasis id="CFEBEADCFFAAFFB81C8A7D3AFDCCFAF7" box="[476,599,1281,1306]" italics="true" pageId="20" pageNumber="21">P. suberea</emphasis>
</taxonomicName>
and recognized both species as distinct returning all of the available Asian names to valid status within
<emphasis id="CFEBEADCFFAAFFB81F677D12FD28FAAF" box="[561,691,1321,1346]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB81F677D12FD34FAAF" box="[561,687,1321,1346]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Phellopsis</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907D6AFE26FA0E" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
<bibRefCitation id="990E4B3FFFAAFFB81D907D6AFE03FA86" author="Masumoto" box="[198,408,1361,1387]" pageId="20" pageNumber="27" refString="Masumoto, K. (1990) A new Phellopsis species from South Korea (Zopheridae, Coleoptera). Entomological Review of Japan, 45, 87 - 91." type="journal article" year="1990">Masumoto (1990)</bibRefCitation>
commented that
<taxonomicName id="3A9F4D4DFFAAFFB81F357D6AFD55FA87" box="[611,718,1361,1386]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="CFEBEADCFFAAFFB81F357D6AFD55FA87" box="[611,718,1361,1386]" italics="true" pageId="20" pageNumber="21">P. imurai</emphasis>
</taxonomicName>
was a relative of
<emphasis id="CFEBEADCFFAAFFB81ECB7D6AFBB6FA87" box="[925,1069,1361,1386]" italics="true" pageId="20" pageNumber="21">P. a m u re n s i s</emphasis>
, but compared it to
<emphasis id="CFEBEADCFFAAFFB8184A7D6AFA00FA87" box="[1308,1435,1361,1386]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB8184A7D6AFA0CFA87" box="[1308,1431,1361,1386]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="suberea">P. suberea</taxonomicName>
,
</emphasis>
from which it clearly differs. A
<typeStatus id="2224886CFFAAFFB81F587D42FDE8FA7E" box="[526,627,1401,1427]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
of
<taxonomicName id="3A9F4D4DFFAAFFB81FCC7D42FC9EFA7F" box="[666,773,1401,1426]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="imurai">
<emphasis id="CFEBEADCFFAAFFB81FCC7D42FC9EFA7F" box="[666,773,1401,1426]" italics="true" pageId="20" pageNumber="21">P. imurai</emphasis>
</taxonomicName>
is the only Korean specimen examined, and is lighter in color (possibly teneral) than many typical specimens of
<emphasis id="CFEBEADCFFAAFFB81E127D9AFC43FA57" box="[836,984,1441,1466]" italics="true" pageId="20" pageNumber="21">P. a m u re n s i s</emphasis>
, but morphologically does not differ from Russian specimens.
</paragraph>
<paragraph id="FD2036CEFFAAFFB81D907DCAFABCF93E" blockId="20.[151,1437,152,1747]" pageId="20" pageNumber="21">
While only reported from the Sikhote-Alin Mountain Range of the Primorski Krai, Amur River valley and mountainous regions of
<collectingCountry id="8588765EFFAAFFB81CE57E22FDD2F9DE" box="[435,585,1561,1587]" name="South Korea" pageId="20" pageNumber="21">South Korea</collectingCountry>
, it is expected that
<taxonomicName id="3A9F4D4DFFAAFFB81E7D7E22FC5BF9DF" box="[811,960,1561,1586]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="amurensis">
<emphasis id="CFEBEADCFFAAFFB81E7D7E22FC5BF9DF" box="[811,960,1561,1586]" italics="true" pageId="20" pageNumber="21">P. amurensis</emphasis>
</taxonomicName>
occurs in suitable habitat areas all along the western slope of the Sea of
<collectingCountry id="8588765EFFAAFFB81F447E7AFDC2F9B6" box="[530,601,1601,1627]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
. This includes
<collectingCountry id="8588765EFFAAFFB81E5B7E7AFC3EF9B6" box="[781,933,1601,1627]" name="North Korea" pageId="20" pageNumber="21">North Korea</collectingCountry>
, and possibly far-eastern areas of the Jilin and/or Heilongjiang provinces of
<collectingCountry id="8588765EFFAAFFB81F797E52FDEEF96E" box="[559,629,1641,1667]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
in the Changbai Shan Mountains where deforestation may be less than total. Specimens from
<collectingCountry id="8588765EFFAAFFB81CB77EAAFDB2F946" box="[481,553,1681,1707]" name="South Korea" pageId="20" pageNumber="21">Korea</collectingCountry>
(identified as
<emphasis id="CFEBEADCFFAAFFB81F807EAAFCCCF947" box="[726,855,1681,1706]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="3A9F4D4DFFAAFFB81F807EAAFCC8F947" box="[726,851,1681,1706]" class="Insecta" family="Zopheridae" genus="Phellopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="suberea">P. suberea</taxonomicName>
,
</emphasis>
almost certainly incorrectly) have been reported from Cheongrangli, Hongcheon, Kangwon, Kwangheung, Kyeongki, and Seoul (
<bibRefCitation id="990E4B3FFFAAFFB8191A7E82FA80F93E" author="Weon" box="[1100,1307,1721,1747]" pageId="20" pageNumber="28" refString="Weon, G. - J., Jang, W. - Y., Oh, K. - S., Shin, S. - C. &amp; Park, J. - D. (2000) List of Species in Insect Collection of the Korean Forest Research Institute, 7. Coleoptera (except Superfamily Chrysomeloidea). Korean Forest Research Institute, Journal of Forestry Science, 63, 108 - 123." type="journal article" year="2000">
Weon
<emphasis id="CFEBEADCFFAAFFB819CF7E82FB4FF93F" box="[1177,1236,1721,1746]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
2000
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>