253 lines
33 KiB
XML
253 lines
33 KiB
XML
<document id="8035A187262221D028F48D12341EE81E" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA031AFF9AFF2FF88DFEFB14DB" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus astrophilus Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="55" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="53" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5DAC79BC7F5C0073D93101EDC9C9E168">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:roleTerm id="1F605335A69FBE8DAE86D12FFB47D4F7">Author</mods:roleTerm>
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<mods:namePart id="DF26B51565F109E61F103F02CAFF0B52">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="5814B97CE33E215F1C0A965B8DC94931">text</mods:typeOfResource>
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<mods:title id="6BFEC262C79B85AF947E220260F1AE29">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F8F5D29E51A59DD2E042073DEE53C48B">
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<mods:date id="8CC7CC210F20927EC7C84A0456EAB2F6">2009</mods:date>
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<mods:detail id="521F019B20DD17DFBCB8543DD51415EF" type="pubDate">
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<mods:number id="240905C4DDCE0F00B0C75EEE2ABB1FD6">2009-08-06</mods:number>
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</mods:detail>
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<mods:detail id="C49ED55200C061A33ED8E02E76F439DB" type="volume">
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<mods:number id="22250EAB3F411C8D24E0D66D4EFD0322">2178</mods:number>
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<mods:start id="8D8140D1CE2A6EBA0D11703D9E132094">1</mods:start>
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<mods:end id="E915D2B99DC836E3DCFB377032D990FF">72</mods:end>
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<mods:location id="B935C12972AE28BC1EB7C827F618AA02">
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<mods:url id="3CEF362480C73C8A5D13DBEF0A7E5629">https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1</mods:url>
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</mods:location>
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<mods:classification id="7BE06DD2508AB01960AEBA58DB641566">journal article</mods:classification>
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<mods:identifier id="449F3D2746E3EC89AA94F732613F9621" type="DOI">10.11646/zootaxa.2178.1.1</mods:identifier>
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<mods:identifier id="E73FED1B369DFF76A3886E3889EADC9E" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="69F550C65164AC91D4B7448F228CFD02" type="Zenodo-Dep">5312276</mods:identifier>
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<treatment id="733A87FA031AFF9AFF2FF88DFEFB14DB" ID-DOI="http://doi.org/10.5281/zenodo.5320104" ID-Zenodo-Dep="5320104" LSID="urn:lsid:plazi:treatment:733A87FA031AFF9AFF2FF88DFEFB14DB" httpUri="http://treatment.plazi.org/id/733A87FA031AFF9AFF2FF88DFEFB14DB" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
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<subSubSection id="B3896567031AFF98FF2FF88DFE1D126B" box="[151,465,1853,1879]" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="FB2C36EC031AFF98FF2FF88DFE1D126B" blockId="52.[151,465,1853,1914]" box="[151,465,1853,1879]" pageId="52" pageNumber="53">
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<heading id="A0648180031AFF98FF2FF88DFE1D126B" bold="true" box="[151,465,1853,1879]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
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<emphasis id="C9E7EAFE031AFF98FF2FF88DFE1D126B" bold="true" box="[151,465,1853,1879]" pageId="52" pageNumber="53">
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<taxonomicName id="3C934D6F031AFF98FF2FF88DFE4B126B" ID-CoL="57ZDL" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,391,1853,1879]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
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<emphasis id="C9E7EAFE031AFF98FF2FF88DFE4B126B" bold="true" box="[151,391,1853,1879]" italics="true" pageId="52" pageNumber="53">Trechus astrophilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D2D45785031AFF98FE37F88DFE1D126B" box="[399,465,1853,1879]" pageId="52" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B3896567031AFF98FF2FF8D0FEAC1246" box="[151,352,1888,1914]" pageId="52" pageNumber="53" type="description">
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||
<paragraph id="FB2C36EC031AFF98FF2FF8D0FEAC1246" blockId="52.[151,465,1853,1914]" box="[151,352,1888,1914]" pageId="52" pageNumber="53">
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(
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<figureCitation id="63A82A69031AFF98FF27F8D0FEE21246" box="[159,302,1888,1914]" captionStart="FIGURES 65–80" captionStartId="31.[151,270,1797,1821]" captionTargetBox="[292,1292,195,1761]" captionTargetId="figure-161@31.[291,1304,194,1761]" captionTargetPageId="31" captionText="FIGURES 65–80. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 65–71, 77–80; Figs. 65, 68–71, 79, 80 in addition with parameres) and in dorsal view (Figs. 72–76). Figs. 65, 74, Trechus astrophilus sp. n., paratype. Fig. 66, T. lama sp. n., paratype. Fig. 67, T. singularis sp. n., holotype. Fig. 68, T. tsampa sp. n., paratype. Figs. 69, 76, T. yak yak ssp. n., paratype. Figs. 70, 73, T. religiosus sp. n., paratype. Figs. 71, 72, T. chaklaensis sp. n., paratype. Fig. 75, T. yak shogulaensis ssp. n., paratype. Fig. 77, T. antonini sp. n., non-type specimen from Lhachen La (= locus typicus). Fig. 78, T. folwarcznyi Deuve, 1997, non-type specimen from Shogu La (= locus typicus). Fig. 79, T. rarus sp. n., holotype. Fig. 80, T. tseringi sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312298" httpUri="https://zenodo.org/record/5312298/files/figure.png" pageId="52" pageNumber="53">Figs. 65, 74</figureCitation>
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,
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<figureCitation id="63A82A69031AFF98FE81F8D0FE9B1246" box="[313,343,1888,1914]" captionStart="FIGURES 87–92" captionStartId="33.[151,270,1891,1915]" captionTargetBox="[161,1404,187,1850]" captionTargetId="figure-70@33.[151,1436,168,1867]" captionTargetPageId="33" captionText="FIGURES 87–92. Habitus. Fig. 87, Trechus martinae sp. n., paratype, female. Fig. 88, T. mieheorum sp. n., holotype. Fig. 89, T. solhoeyi sp. n., paratype, male. Fig. 90, T. yeti sp. n., holotype. Fig. 91, T. astrophilus sp. n., paratype, male. Fig. 92, T. tsampa sp. n., holotype." figureDoi="http://doi.org/10.5281/zenodo.5312302" httpUri="https://zenodo.org/record/5312302/files/figure.png" pageId="52" pageNumber="53">91</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="B3896567031AFF99FF2FF81DFD0015C3" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="FB2C36EC031AFF98FF2FF81DFC6A12D2" blockId="52.[151,1436,1965,2030]" pageId="52" pageNumber="53">
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<emphasis id="C9E7EAFE031AFF98FF2FF81DFE9312FB" bold="true" box="[151,351,1965,1991]" pageId="52" pageNumber="53">Type material:</emphasis>
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<materialsCitation id="4BFB3CB1031AFF98FED1F81DFC6A12D2" collectionCode="BMNH" elevation="5450" latitude="30.185276" location="South Central" longLatPrecision="20" longitude="90.47833" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
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<typeStatus id="2428884E031AFF98FED1F81DFE1212FB" box="[361,478,1965,1991]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
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male, with label data “
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<collectingRegion id="3957F80E031AFF98FCB1F81DFCA812FB" box="[777,868,1965,1991]" country="China" name="Xizang" pageId="52" pageNumber="53">TIBET</collectingRegion>
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(
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<location id="FE4C6037031AFF98FCCFF81DFBE012FB" LSID="urn:lsid:plazi:treatment:733A87FA031AFF9AFF2FF88DFEFB14DB:FE4C6037031AFF98FCCFF81DFBE012FB" box="[887,1068,1965,1991]" latitude="30.185276" longLatPrecision="20" longitude="90.47833" name="South Central" pageId="52" pageNumber="53" stateProvince="Xizang">South Central</location>
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||
) 19.VI.07, Budha Vall.
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<locationDeviation id="EA9E2FCE031AFF98FADBF81DFEFE12D2" location="Yangpachem" pageId="52" pageNumber="53">N of Yangpachem</locationDeviation>
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||
, ca.
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||
<geoCoordinate id="9EA7502B031AFF98FEDEF864FE2B12D2" box="[358,487,2004,2030]" degrees="30" direction="north" minutes="11" orientation="latitude" pageId="52" pageNumber="53" precision="15" seconds="07" value="30.185276">30°11’07N</geoCoordinate>
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||
<geoCoordinate id="9EA7502B031AFF98FE56F864FDBC12D2" box="[494,624,2004,2030]" degrees="90" direction="east" minutes="28" orientation="longitude" pageId="52" pageNumber="53" precision="15" seconds="42" value="90.47833">90°28’42E</geoCoordinate>
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,
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||
<quantity id="3C6B9B09031AFF98FDC4F864FCEC12D2" box="[636,800,2004,2030]" metricMagnitude="3" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="52" pageNumber="53" unit="m" value="5450.0" valueMax="5600.0" valueMin="5300.0">
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||
<elevation id="70BED1DF031AFF98FDC4F864FCEC12D2" box="[636,800,2004,2030]" metricMagnitude="3" metricUnit="m" metricValue="5.45" metricValueMax="5.6" metricValueMin="5.3" pageId="52" pageNumber="53" unit="m" value="5450.0" valueMax="5600.0" valueMin="5300.0">5300–5600 m</elevation>
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</quantity>
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” (
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<collectionCode id="9D82AE29031AFF98FC83F864FC5512D2" box="[827,921,2004,2030]" pageId="52" pageNumber="53">BMNH</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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<paragraph id="FB2C36EC031BFF99FF7DFF28FD0015C3" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">
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||
<materialsCitation id="4BFB3CB1031BFF99FF7DFF28FEAD15E4" collectionCode="BMNH, CKAB, CSCHM, CWR, MNHN, SMNS" elevation="5450" latitude="30.185276" location="South Central" longLatPrecision="20" longitude="90.47833" pageId="53" pageNumber="54" specimenCount="64" specimenCount-female="12" specimenCount-male="52" stateProvince="Xizang" typeStatus="paratype">
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<typeStatus id="2428884E031BFF99FF7DFF28FE8D158E" box="[197,321,152,178]" pageId="53" pageNumber="54" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="ED95FD65031BFF99FEF5FF28FE72158E" box="[333,446,152,178]" pageId="53" pageNumber="54" type="male">52 males</specimenCount>
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,
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<specimenCount id="ED95FD65031BFF99FE74FF28FD99158E" box="[460,597,152,178]" pageId="53" pageNumber="54" type="female">12 females</specimenCount>
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||
, with same label data as holotype (
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<collectionCode id="9D82AE29031BFF99FBBEFF28FBA9158E" box="[1030,1125,152,178]" pageId="53" pageNumber="54">BMNH</collectionCode>
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||
,
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<collectionCode id="9D82AE29031BFF99FBCAFF28FB05158E" box="[1138,1225,152,178]" pageId="53" pageNumber="54">CKAB</collectionCode>
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||
,
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<collectionCode id="9D82AE29031BFF99FB6EFF28FA8E158E" box="[1238,1346,152,178]" pageId="53" pageNumber="54">CSCHM</collectionCode>
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||
,
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<collectionCode id="9D82AE29031BFF99FAE8FF28FA5B158E" box="[1360,1431,152,178]" pageId="53" pageNumber="54">CWR</collectionCode>
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,
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<collectionCode id="9D82AE29031BFF99FF2FFF0EFF3915E4" box="[151,245,190,216]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="53" pageNumber="54">MNHN</collectionCode>
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,
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<collectionCode id="9D82AE29031BFF99FEBAFF0EFE9415E4" box="[258,344,190,216]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="53" pageNumber="54">SMNS</collectionCode>
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||
)
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</materialsCitation>
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;
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<materialsCitation id="4BFB3CB1031BFF99FED5FF0EFD0415C3" collectionCode="CSCHM" elevation="5100" latitude="30.182222" location="South Central" longLatPrecision="20" longitude="90.48917" pageId="53" pageNumber="54" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
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<specimenCount id="ED95FD65031BFF99FED5FF0EFE0E15E4" box="[365,450,190,216]" pageId="53" pageNumber="54" type="male">1 male</specimenCount>
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||
,
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<specimenCount id="ED95FD65031BFF99FE77FF0EFDF615E4" box="[463,570,190,216]" pageId="53" pageNumber="54" type="female">1 female</specimenCount>
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||
,
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||
<location id="FE4C6037031BFF99FDFFFF0EFD2215E4" LSID="urn:lsid:plazi:treatment:733A87FA031AFF9AFF2FF88DFEFB14DB:FE4C6037031BFF99FDFFFF0EFD2215E4" box="[583,750,190,216]" latitude="30.182222" longLatPrecision="20" longitude="90.48917" name="South Central" pageId="53" pageNumber="54" stateProvince="Xizang">South Central</location>
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||
<collectingRegion id="3957F80E031BFF99FD4EFF0EFCFB15E4" box="[758,823,190,216]" country="China" name="Xizang" pageId="53" pageNumber="54">Tibet</collectingRegion>
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||
, Budha Vall.
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<locationDeviation id="EA9E2FCE031BFF99FC63FF0EFB7B15E4" box="[987,1207,190,216]" location="Yangpachem" pageId="53" pageNumber="54">N of Yangpachem</locationDeviation>
|
||
,
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||
<quantity id="3C6B9B09031BFF99FB7CFF0EFAA115E4" box="[1220,1389,190,216]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="53" pageNumber="54" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">
|
||
<elevation id="70BED1DF031BFF99FB7CFF0EFAA115E4" box="[1220,1389,190,216]" metricMagnitude="3" metricUnit="m" metricValue="5.1" metricValueMax="5.2" metricValueMin="5.0" pageId="53" pageNumber="54" unit="m" value="5100.0" valueMax="5200.0" valueMin="5000.0">5000–5200 m</elevation>
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||
</quantity>
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||
, ca.
|
||
<geoCoordinate id="9EA7502B031BFF99FF2FFF55FED615C3" box="[151,282,229,255]" degrees="30" direction="north" minutes="10" orientation="latitude" pageId="53" pageNumber="54" precision="15" seconds="56" value="30.182222">30°10’56N</geoCoordinate>
|
||
<geoCoordinate id="9EA7502B031BFF99FE99FF55FE6E15C3" box="[289,418,229,255]" degrees="90" direction="east" minutes="29" orientation="longitude" pageId="53" pageNumber="54" precision="15" seconds="21" value="90.48917">90°29’21E</geoCoordinate>
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||
, 17–20.VI.07 (
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||
<collectionCode id="9D82AE29031BFF99FDEAFF55FD0C15C3" box="[594,704,229,255]" pageId="53" pageNumber="54">CSCHM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="B3896567031BFF99FF7DFEBBFAA11092" pageId="53" pageNumber="54" type="description">
|
||
<paragraph id="FB2C36EC031BFF99FF7DFEBBFD57141A" blockId="53.[151,1436,152,2034]" box="[197,667,267,294]" pageId="53" pageNumber="54">
|
||
<emphasis id="C9E7EAFE031BFF99FF7DFEBBFE931419" bold="true" box="[197,351,267,293]" pageId="53" pageNumber="54">Description:</emphasis>
|
||
Body length:
|
||
<quantity id="3C6B9B09031BFF99FDBEFEBCFD5B141A" box="[518,663,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.9" metricValueMin="3.2" pageId="53" pageNumber="54" unit="mm" value="3.55" valueMax="3.9" valueMin="3.2">3.2–3.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031BFF99FF7DFE82FB3014A6" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Colour: Dorsal surface dark brown, shiny, with pronotum dark reddish brown. Scapus, basal segment of maxillary palpus and legs yellowish brown; distal half of antennal segment II, antennal segments IV–XI on the whole, second segment and basal portion of the third segment of maxillary palpus darkened.</paragraph>
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<paragraph id="FB2C36EC031BFF99FF7DFE16FE8B1708" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Microsculpture: Supraorbital area and disc of pronotum almost polished, with very faintly engraved meshes, visible under high magnification only (x100). More deeply engraved almost isodiametric meshes in frontal furrows of head and in pronotal basal depression. Slightly engraved slightly transverse meshes on disc of elytra (x60).</paragraph>
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<paragraph id="FB2C36EC031BFF99FF7DFDF1FD5D17F3" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Head: Average, with eyes small and slightly protruding; temples approximately 4/5 of length of eyes and strongly wrinkled to the neck. Frontal furrows slightly flattened at level of hind suborbital seta. Antennae average, 3 antennomeres extend beyond the pronotal base. Antennomere III slightly longer than antennomere II and IV, both the latter are alike in length.</paragraph>
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<paragraph id="FB2C36EC031BFF99FF7DFD6CFE49168B" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Pronotum: Relatively small, cordate, moderately transverse and strongly contracted towards base; proportions WP/LP = 1.25–1.31, WP/WPB = 1.35–1.39, WP/WH = 1.15–1.20, WE/WP = 1.72–1.81. Surface strongly convex. Sides concave anterad of hind angles; the latter moderately large, sometimes slightly bent outwards, rectangular or slightly obtuse. Marginal gutter narrow, slightly widened anterad of laterobasal depressions. Base rectilinear or weakly convex in middle, more strongly curved anteriorly at outer fifth; basal depressions smooth.</paragraph>
|
||
<paragraph id="FB2C36EC031BFF99FF7DFC74FDC411D1" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Elytra: Oval, broadest almost at mid-length, with proportion WE/LE = 1.48–1.54. Surface strongly convex, not flattened on disc. Shoulders rounded, indistinct. Striae faintly or indistinctly punctate, first stria fully deep impressed, striae II–IV deeply impressed on disc but reduced at base and extreme apex, outer striae shallower, stria VII very faintly impressed but present. Stria VIII moderately impressed from level of the fifth umbilicate pore backwards and more deeply impressed at levels of seventh and eighth pores. Recurrent elytral preapical sulcus connected with the end of the fifth or seventh stria. Intervals I–IV (-V) strongly convex. Preapical seta is located in the third interval distinctly before the end of the second stria and at the beginning of the last seventh of the elytra.</paragraph>
|
||
<paragraph id="FB2C36EC031BFF99FF7DFB49FE24102F" blockId="53.[151,1436,152,2034]" box="[197,488,1273,1299]" pageId="53" pageNumber="54">Legs: Relatively slender.</paragraph>
|
||
<paragraph id="FB2C36EC031BFF99FF7DFA90FAA11092" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">Male genitalia: Aedeagal median lobe average in length (LE/LA = 2.55–2.78), more strongly curved in basal half, slightly elongate towards apex, but with terminal lamella short; the latter slightly curved upwards. Basal bulb average. The structure of the sclerotized internal sac portion is distinctly bipartite, in dorsal view with a long needle-like copulatory piece which is surrounded by a more sac like sheet. Parameres slender.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B3896567031BFF9AFF7DFA0AFA4A158E" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="etymology">
|
||
<paragraph id="FB2C36EC031BFF99FF7DFA0AFC97131E" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">
|
||
<emphasis id="C9E7EAFE031BFF99FF7DFA0AFE9610E8" bold="true" box="[197,346,1466,1492]" pageId="53" pageNumber="54">Etymology:</emphasis>
|
||
The specific name is derived from the Latinized Greek words “astrum” (starry sky) and “philum” (friend) and refers to the habitat of the new species which extends to exceptionally high altitudes, and which offer ideal condition to watch the stars; adjective.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031BFF99FF7DF99EFCD81381" blockId="53.[151,1436,152,2034]" pageId="53" pageNumber="54">
|
||
<emphasis id="C9E7EAFE031BFF99FF7DF99EFE4E1374" bold="true" box="[197,386,1582,1608]" pageId="53" pageNumber="54">Identification:</emphasis>
|
||
In external characters similar to
|
||
<taxonomicName id="3C934D6F031BFF99FC9CF99FFBB31374" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" box="[804,1151,1582,1608]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="folwarcznyi">
|
||
<emphasis id="C9E7EAFE031BFF99FC9CF99FFC1F1374" box="[804,979,1583,1608]" italics="true" pageId="53" pageNumber="54">T. folwarcznyi</emphasis>
|
||
<bibRefCitation id="9F024B1D031BFF99FC65F99EFBB31374" author="Deuve, T." box="[989,1151,1582,1608]" pageId="53" pageNumber="54" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
, but with body size on average larger, with head, antennae and legs somewhat slender, with temporae longer and with antennae darker. Moreover,
|
||
<taxonomicName id="3C934D6F031BFF99FED6F9CCFDC213A9" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[366,526,1660,1685]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031BFF99FED6F9CCFDC213A9" box="[366,526,1660,1685]" italics="true" pageId="53" pageNumber="54">T. astrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031BFF99FDACF9CBFD9913A9" bold="true" box="[532,597,1659,1685]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="D2D45785031BFF99FDACF9CBFD9913A9" box="[532,597,1659,1685]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is easily distinguished from all other species of the
|
||
<taxonomicName id="3C934D6F031BFF99FB17F9CCFAE113A9" authorityName="Deuve" authorityYear="1997" box="[1199,1325,1660,1685]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="antonini">
|
||
<emphasis id="C9E7EAFE031BFF99FB17F9CCFAE113A9" box="[1199,1325,1660,1685]" italics="true" pageId="53" pageNumber="54">T. antonini</emphasis>
|
||
</taxonomicName>
|
||
group by the more extensively sclerotized aedeagal internal sac.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC031BFF9AFF7DF979FA4A158E" blockId="53.[151,1436,152,2034]" lastBlockId="54.[151,1436,152,487]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
|
||
<emphasis id="C9E7EAFE031BFF99FF7DF979FE4B13DF" bold="true" box="[197,391,1737,1763]" pageId="53" pageNumber="54">Relationships:</emphasis>
|
||
<taxonomicName id="3C934D6F031BFF99FE29F979FD8C13DE" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[401,576,1737,1762]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031BFF99FE29F979FD8C13DE" box="[401,576,1737,1762]" italics="true" pageId="53" pageNumber="54">T. astrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031BFF99FDF3F979FD5F13DF" bold="true" box="[587,659,1737,1763]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="D2D45785031BFF99FDF3F979FD5F13DF" box="[587,659,1737,1763]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the below described
|
||
<taxonomicName id="3C934D6F031BFF99FC67F979FBF213DE" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[991,1086,1737,1762]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031BFF99FC67F979FBF213DE" box="[991,1086,1737,1762]" italics="true" pageId="53" pageNumber="54">T. lama</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031BFF99FBF1F979FB5D13DF" bold="true" box="[1097,1169,1737,1763]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="D2D45785031BFF99FBF1F979FB5D13DF" box="[1097,1169,1737,1763]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
together share some conspicuous similarities in the shape of body and aedeagus which are considered to be synapomorphies and therefore, which indicate a sister species relationship: Antennae and legs more slender, elytra more evenly rounded, aedeagal median lobe more elongated towards apex. The above newly described species
|
||
<taxonomicName id="3C934D6F031BFF99FA3DF88DFED71241" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="budhaensis" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031BFF99FA3DF88DFED71241" italics="true" pageId="53" pageNumber="54">T. budhaensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031BFF99FE9AF8D3FEA91241" bold="true" box="[290,357,1891,1917]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="D2D45785031BFF99FE9AF8D3FEA91241" box="[290,357,1891,1917]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3C934D6F031BFF99FE26F8D4FE2A1241" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[414,486,1892,1917]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="yeti" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE031BFF99FE26F8D4FE2A1241" box="[414,486,1892,1917]" italics="true" pageId="53" pageNumber="54">T. yeti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE031BFF99FE55F8D3FDFC1241" bold="true" box="[493,560,1891,1917]" pageId="53" pageNumber="54">
|
||
<taxonomicNameLabel id="D2D45785031BFF99FE55F8D3FDFC1241" box="[493,560,1891,1917]" pageId="53" pageNumber="54" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
share a similar body shape, however, based on apomorphic charater states of male genitalia both the latter species seem to belong to another evolutionary line which includes
|
||
<taxonomicName id="3C934D6F031BFF99FAA6F83BFEFA12F7" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="antonini">
|
||
<emphasis id="C9E7EAFE031BFF99FAA6F83BFA501298" box="[1310,1436,1931,1956]" italics="true" pageId="53" pageNumber="54">T. antonini</emphasis>
|
||
<bibRefCitation id="9F024B1D031BFF99FF2FF801FEFA12F7" author="Deuve, T." box="[151,310,1969,1995]" pageId="53" pageNumber="54" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
, a species with a shorter head and appendages and with less rounded elytra (see discussion above). Therefore, parallel evolution of similar external body shape characters are very likely in both the sister species pairs
|
||
<emphasis id="C9E7EAFE0318FF9AFEC0FF28FDB4158D" box="[376,632,152,178]" italics="true" pageId="54" pageNumber="55">
|
||
<taxonomicName id="3C934D6F0318FF9AFEC0FF28FDD4158D" authorityName="Schmidt" authorityYear="2009" box="[376,536,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="budhaensis">T. budhaensis</taxonomicName>
|
||
-
|
||
<taxonomicName id="3C934D6F0318FF9AFD89FF28FDB4158D" authorityName="Schmidt" authorityYear="2009" box="[561,632,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="yeti">T. yeti</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="C9E7EAFE0318FF9AFD09FF28FC0C158D" box="[689,960,152,178]" italics="true" pageId="54" pageNumber="55">
|
||
<taxonomicName id="3C934D6F0318FF9AFD09FF28FC9D158D" authorityName="Schmidt" authorityYear="2009" box="[689,849,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="astrophilus">T. astrophilus</taxonomicName>
|
||
-
|
||
<taxonomicName id="3C934D6F0318FF9AFCD1FF28FC0C158D" authorityName="Schmidt" authorityYear="2009" box="[873,960,152,177]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="lama">T. lama</taxonomicName>
|
||
</emphasis>
|
||
without indicating close relationships.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B38965670318FF9AFF7DFF0EFEFB14DB" pageId="54" pageNumber="55" type="distribution">
|
||
<paragraph id="FB2C36EC0318FF9AFF7DFF0EFBB115C3" blockId="54.[151,1436,152,487]" pageId="54" pageNumber="55">
|
||
<emphasis id="C9E7EAFE0318FF9AFF7DFF0EFEA015E4" bold="true" box="[197,364,190,216]" pageId="54" pageNumber="55">Distribution:</emphasis>
|
||
<figureCitation id="63A82A690318FF9AFECEFF0EFE1215E4" box="[374,478,190,216]" captionStart="FIGURE 100" captionStartId="67.[151,256,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-404@67.[151,1436,194,1082]" captionTargetPageId="67" captionText="FIGURE 100. Map of Southern Central Tibet showing distribution of species of the Trechus antonini group (white circles). 1, T. antonini Deuve, 1997. 2, T. religiosus sp. n. 3, T. astrophilus sp. n. 4, T. budhaensis sp. n. 5, T. rarus sp. n. 6, T. yeti sp. n. 7, T. folwarcznyi Deuve, 1997. 8, T. yak yak ssp. n. 9, T. yak shogulaensis ssp. n. 10, T. tseringi sp. n. 11, T. singularis sp. n. 12, T. tsampa sp. n. 13, T. lama sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312320" httpUri="https://zenodo.org/record/5312320/files/figure.png" pageId="54" pageNumber="55">Fig. 100</figureCitation>
|
||
. Currently only known from the source area of the glacier brook of the Budha Valley on south slope of central Nyainqentanglha Shan Massif north of Yangpachem.
|
||
</paragraph>
|
||
<paragraph id="FB2C36EC0318FF9AFF7DFEBBFEFB14DB" blockId="54.[151,1436,152,487]" pageId="54" pageNumber="55">
|
||
<emphasis id="C9E7EAFE0318FF9AFF7DFEBBFEFE1419" bold="true" box="[197,306,267,293]" pageId="54" pageNumber="55">Habitat:</emphasis>
|
||
Higher alpine zone. The species seems to be strictly limited to altitudes above
|
||
<quantity id="3C6B9B090318FF9AFAB7FEBCFABC141A" box="[1295,1392,268,294]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="54" pageNumber="55" unit="m" value="5200.0">5200 m</quantity>
|
||
. Its vertical distribution extends to the upper limit of the alpine zone and adjoins the nival zone. Up to an altitude of
|
||
<quantity id="3C6B9B090318FF9AFF00FEE9FEDE144F" box="[184,274,345,371]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="54" pageNumber="55" unit="m" value="5600.0">5600 m</quantity>
|
||
the species was frequently found under stones on gently inclined slopes in southern and western expositions as well on the top of an older moraine. At its highest occurrences,
|
||
<taxonomicName id="3C934D6F0318FF9AFBDBFE30FAC614A5" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1123,1290,384,409]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
|
||
<emphasis id="C9E7EAFE0318FF9AFBDBFE30FAC614A5" box="[1123,1290,384,409]" italics="true" pageId="54" pageNumber="55">T. astrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="C9E7EAFE0318FF9AFAACFECFFA9514A5" bold="true" box="[1300,1369,383,409]" pageId="54" pageNumber="55">
|
||
<taxonomicNameLabel id="D2D457850318FF9AFAACFECFFA9514A5" box="[1300,1369,383,409]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lives sympatrically with
|
||
<taxonomicName id="3C934D6F0318FF9AFEC7FE17FD1014FC" authority="Hieke, 1995" authorityName="Hieke" authorityYear="1995" box="[383,732,422,448]" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="altiphila">
|
||
<emphasis id="C9E7EAFE0318FF9AFEC7FE17FDF214FC" box="[383,574,423,448]" italics="true" pageId="54" pageNumber="55">Amara altiphila</emphasis>
|
||
Hieke, 1995
|
||
</taxonomicName>
|
||
, and with a hitherto undescribed species of the
|
||
<taxonomicName id="3C934D6F0318FF9AFAA3FE17FF2814DA" authorityName="Toledano" authorityYear="2000" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="C9E7EAFE0318FF9AFAA3FE17FF2814DA" italics="true" pageId="54" pageNumber="55">Bembidion baehri</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |