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<document id="347670543200411619636064F5FB03F6" ID-DOI="10.11646/zootaxa.2178.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5312276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629627601103" checkinUser="felipe" docAuthor="Schmidt, Joachim" docDate="2009" docId="733A87FA0318FF9BFF2FFDF1FB5D10E9" docLanguage="en" docName="zt02178p072.pdf" docOrigin="Zootaxa 2178 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2178.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trechus lama Schmidt 2009, sp. n." docType="treatment" docVersion="4" lastPageNumber="56" masterDocId="8F03FF82032EFFACFFB8FFB0FFCC153C" masterDocTitle="Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178" masterLastPageNumber="72" masterPageNumber="1" pageNumber="55" updateTime="1699086330132" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="38B106660E466F72F1A9BEB0C60CCB49">Taxonomic and biogeographical review of the genus Trechus Clairville, 1806, from the Tibetan Himalaya and the southern central Tibetan Plateau (Coleoptera: Carabidae: Trechini) 2178</mods:title>
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<mods:namePart id="4C910BF788040CACCDB1AC5C09D113D6">Schmidt, Joachim</mods:namePart>
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<mods:date id="049C376FA70A8123040D2A51AE96522B">2009</mods:date>
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<mods:number id="6B990FBCEE3C06295EA7BD754989F62B">2009-08-06</mods:number>
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<treatment id="733A87FA0318FF9BFF2FFDF1FB5D10E9" ID-DOI="http://doi.org/10.5281/zenodo.5320106" ID-Zenodo-Dep="5320106" LSID="urn:lsid:plazi:treatment:733A87FA0318FF9BFF2FFDF1FB5D10E9" httpUri="http://treatment.plazi.org/id/733A87FA0318FF9BFF2FFDF1FB5D10E9" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<subSubSection id="B38965670318FF9AFF2FFDF1FE4A1766" box="[151,390,576,603]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="FB2C36EC0318FF9AFF2FFDF1FE4A1766" blockId="54.[151,390,576,638]" box="[151,390,576,603]" pageId="54" pageNumber="55">
<heading id="A06481800318FF9AFF2FFDF1FE4A1766" bold="true" box="[151,390,576,603]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="C9E7EAFE0318FF9AFF2FFDF1FE4A1766" bold="true" box="[151,390,576,603]" pageId="54" pageNumber="55">
<taxonomicName id="3C934D6F0318FF9AFF2FFDF1FEF01767" ID-CoL="57ZR7" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[151,316,577,603]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
<emphasis id="C9E7EAFE0318FF9AFF2FFDF1FEF01767" bold="true" box="[151,316,577,603]" italics="true" pageId="54" pageNumber="55">Trechus lama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D457850318FF9AFEFCFDF0FE4A1766" box="[324,390,576,602]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B38965670318FF9AFF2FFDD4FEF91742" box="[151,309,612,638]" pageId="54" pageNumber="55" type="description">
<paragraph id="FB2C36EC0318FF9AFF2FFDD4FEF91742" blockId="54.[151,390,576,638]" box="[151,309,612,638]" pageId="54" pageNumber="55">
(
<figureCitation id="63A82A690318FF9AFF27FDD4FECE1742" box="[159,258,612,638]" captionStart="FIGURES 6580" captionStartId="31.[151,270,1797,1821]" captionTargetBox="[292,1292,195,1761]" captionTargetId="figure-161@31.[291,1304,194,1761]" captionTargetPageId="31" captionText="FIGURES 6580. Aedeagal median lobe with sclerotized portions of internal sac in lateral view (Figs. 6571, 7780; Figs. 65, 6871, 79, 80 in addition with parameres) and in dorsal view (Figs. 7276). Figs. 65, 74, Trechus astrophilus sp. n., paratype. Fig. 66, T. lama sp. n., paratype. Fig. 67, T. singularis sp. n., holotype. Fig. 68, T. tsampa sp. n., paratype. Figs. 69, 76, T. yak yak ssp. n., paratype. Figs. 70, 73, T. religiosus sp. n., paratype. Figs. 71, 72, T. chaklaensis sp. n., paratype. Fig. 75, T. yak shogulaensis ssp. n., paratype. Fig. 77, T. antonini sp. n., non-type specimen from Lhachen La (= locus typicus). Fig. 78, T. folwarcznyi Deuve, 1997, non-type specimen from Shogu La (= locus typicus). Fig. 79, T. rarus sp. n., holotype. Fig. 80, T. tseringi sp. n., paratype." figureDoi="http://doi.org/10.5281/zenodo.5312298" httpUri="https://zenodo.org/record/5312298/files/figure.png" pageId="54" pageNumber="55">Figs. 66</figureCitation>
,
<figureCitation id="63A82A690318FF9AFEB6FDD4FEE01742" box="[270,300,612,638]" captionStart="FIGURES 9395" captionStartId="34.[151,273,1047,1071]" captionTargetBox="[175,1400,218,1010]" captionTargetId="figure-300@34.[151,1436,194,1023]" captionTargetPageId="34" captionText="FIGURES 9395. Habitus. Trechus lama sp. n., paratype, male (93). T. religiosus sp. n., paratype, male (94). T. chaklaensis sp. n., paratype, female (95)." figureDoi="http://doi.org/10.5281/zenodo.5312306" httpUri="https://zenodo.org/record/5312306/files/figure.png" pageId="54" pageNumber="55">93</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B38965670318FF9AFF2FFD00FEC31603" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="FB2C36EC0318FF9AFF2FFD00FEC31603" blockId="54.[151,1436,688,870]" pageId="54" pageNumber="55">
<emphasis id="C9E7EAFE0318FF9AFF2FFD00FE9C17F6" bold="true" box="[151,336,688,714]" pageId="54" pageNumber="55">Type material:</emphasis>
<materialsCitation id="4BFB3CB10318FF9AFEE0FD01FE531624" collectionCode="BMNH" elevation="5050" latitude="29.705" location="South Central" longLatPrecision="20" longitude="90.58778" pageId="54" pageNumber="55" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="2428884E0318FF9AFEE0FD01FE0917F7" box="[344,453,689,715]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
male, with label data “
<collectingRegion id="3957F80E0318FF9AFD6FFD01FCE117F7" box="[727,813,689,715]" country="China" name="Xizang" pageId="54" pageNumber="55">TIBET</collectingRegion>
(
<location id="FE4C60370318FF9AFC85FD01FC2817F7" LSID="urn:lsid:plazi:treatment:733A87FA0318FF9BFF2FFDF1FB5D10E9:FE4C60370318FF9AFC85FD01FC2817F7" box="[829,996,689,715]" latitude="29.705" longLatPrecision="20" longitude="90.58778" name="South Central" pageId="54" pageNumber="55" stateProvince="Xizang">South Central</location>
) 29.VI.07, Dulong, Kurum vall. NW Lhasa,
<quantity id="3C6B9B090318FF9AFF4BFD68FE6F17CE" box="[243,419,728,754]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="54" pageNumber="55" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">
<elevation id="70BED1DF0318FF9AFF4BFD68FE6F17CE" box="[243,419,728,754]" metricMagnitude="3" metricUnit="m" metricValue="5.05" metricValueMax="5.2" metricValueMin="4.9" pageId="54" pageNumber="55" unit="m" value="5050.0" valueMax="5200.0" valueMin="4900.0">49005200 m</elevation>
</quantity>
, ca.
<geoCoordinate id="9EA7502B0318FF9AFE58FD68FDA617CE" box="[480,618,728,754]" degrees="29" direction="north" minutes="42" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="18" value="29.705">29°4218N</geoCoordinate>
<geoCoordinate id="9EA7502B0318FF9AFDCDFD68FCCD17CE" box="[629,769,728,754]" degrees="90" direction="east" minutes="35" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="16" value="90.58778">90°3516E</geoCoordinate>
”, “south ascent of Tsubu side valley, above Tsurphu Monastery” (
<collectionCode id="9D82AE290318FF9AFE8EFD4EFE5B1624" box="[310,407,766,792]" pageId="54" pageNumber="55">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="4BFB3CB10318FF9AFE14FD4EFEC71603" collectionCode="BMNH, CKAB, CSCHM" elevation="5050" latitude="29.705" location="South Central" longLatPrecision="20" longitude="90.58778" pageId="54" pageNumber="55" specimenCount="28" specimenCount-female="7" specimenCount-male="21" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="2428884E0318FF9AFE14FD4EFDEA1624" box="[428,550,766,792]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="ED95FD650318FF9AFD8BFD4EFD6E1624" box="[563,674,766,792]" pageId="54" pageNumber="55" type="male">21 males</specimenCount>
,
<specimenCount id="ED95FD650318FF9AFD08FD4EFCE51624" box="[688,809,766,792]" pageId="54" pageNumber="55" type="female">7 females</specimenCount>
, with same label data as holotype (
<collectionCode id="9D82AE290318FF9AFB6DFD4EFAFF1624" box="[1237,1331,766,792]" pageId="54" pageNumber="55">BMNH</collectionCode>
,
<collectionCode id="9D82AE290318FF9AFAF9FD4EFA5B1624" box="[1345,1431,766,792]" pageId="54" pageNumber="55">CKAB</collectionCode>
,
<collectionCode id="9D82AE290318FF9AFF2FFC95FECF1603" box="[151,259,805,831]" pageId="54" pageNumber="55">CSCHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B38965670318FF9BFF7DFCFBFA591656" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph id="FB2C36EC0318FF9AFF7DFCFBFD57165A" blockId="54.[151,1436,688,870]" box="[197,667,843,870]" pageId="54" pageNumber="55">
<emphasis id="C9E7EAFE0318FF9AFF7DFCFBFE931659" bold="true" box="[197,351,843,869]" pageId="54" pageNumber="55">Description:</emphasis>
Body length: 3.54.0 mm.
</paragraph>
<caption id="AFEC66640318FF9AFF7DF8F0FDE812DE" ID-DOI="http://doi.org/10.5281/zenodo.5312308" ID-Zenodo-Dep="5312308" httpUri="https://zenodo.org/record/5312308/files/figure.png" pageId="54" pageNumber="55" startId="54.[197,301,1856,1880]" targetBox="[172,1412,916,1846]" targetPageId="54">
<paragraph id="FB2C36EC0318FF9AFF7DF8F0FDE812DE" blockId="54.[151,1436,1856,2018]" pageId="54" pageNumber="55">
<emphasis id="C9E7EAFE0318FF9AFF7DF8F0FE9B1264" bold="true" box="[197,343,1856,1880]" pageId="54" pageNumber="55">FIGURE 96.</emphasis>
Glacier formed landscape on central Nyainqentanglha Shan Massif with the highest occurrences of carabid beetles worldwide: Under big stones (foreground) in an altitude of 5600 m on the anticline of an older moraine three carabid beetle species were found frequently:
<taxonomicName id="3C934D6F0318FF9AFD76F836FB5012A0" authority="Hieke, 1995" authorityName="Hieke" authorityYear="1995" box="[718,1180,1925,1949]" class="Insecta" family="Carabidae" genus="Amara" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="altiphila" subGenus="Bradytulus">
<emphasis id="C9E7EAFE0318FF9AFD76F836FCD412A1" box="[718,792,1926,1949]" italics="true" pageId="54" pageNumber="55">Amara</emphasis>
(
<emphasis id="C9E7EAFE0318FF9AFC90F836FC5312A1" box="[808,927,1926,1949]" italics="true" pageId="54" pageNumber="55">Bradytulus</emphasis>
)
<emphasis id="C9E7EAFE0318FF9AFC08F836FBC012A1" box="[944,1036,1926,1949]" italics="true" pageId="54" pageNumber="55">altiphila</emphasis>
Hieke, 1995
</taxonomicName>
, a hitherto undescribed species of the
<taxonomicName id="3C934D6F0318FF9AFE8CF819FE3412FC" authorityName="Toledano" authorityYear="2000" box="[308,504,1961,1984]" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="baehri">
<emphasis id="C9E7EAFE0318FF9AFE8CF819FE3412FC" box="[308,504,1961,1984]" italics="true" pageId="54" pageNumber="55">Bembidion baehri</emphasis>
</taxonomicName>
group, and
<taxonomicName id="3C934D6F0318FF9AFDC4F819FC9E12FC" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[636,850,1961,1984]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0318FF9AFDC4F819FC9E12FC" box="[636,850,1961,1984]" italics="true" pageId="54" pageNumber="55">Trechus astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0318FF9AFCE2F818FC5412FC" bold="true" box="[858,920,1960,1984]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="D2D457850318FF9AFCE2F818FC5412FC" box="[858,920,1960,1984]" pageId="54" pageNumber="55" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(Photo taken by the author, 19.VI.2007, above Budha Valley north of Yangpachem).
</paragraph>
</caption>
<paragraph id="FB2C36EC0319FF9BFF7DFF28FCC115C3" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">Colour: Dorsal surface dark brown, shiny, with pronotum in most specimens somewhat lightened reddish brown. Antennae, palpi and legs yellowish brown; in some specimens distal half of antennal segment III or IV and antennal segments IVXI on the whole darkened.</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFEBCFCE21419" blockId="55.[151,1436,152,1493]" box="[197,814,267,294]" pageId="55" pageNumber="56">
Microsculpture: As described in
<taxonomicName id="3C934D6F0319FF9BFDFDFEBCFD291419" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[581,741,268,293]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFDFDFEBCFD291419" box="[581,741,268,293]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFD54FEBBFCE21419" bold="true" box="[748,814,267,293]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFD54FEBBFCE21419" box="[748,814,267,293]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFE82FD69144E" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
Head: Temples 4/5 5/6 of length of eyes, in some specimens almost as long as eyes. In all other characters agreeing with
<taxonomicName id="3C934D6F0319FF9BFE04FEE9FD90144E" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[444,604,345,370]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFE04FEE9FD90144E" box="[444,604,345,370]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFDDBFEE8FD69144E" bold="true" box="[611,677,344,370]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFDDBFEE8FD69144E" box="[611,677,344,370]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFE30FB6F1708" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
Pronotum: Proportions: WP/LP = 1.191.29, WP/WPB = 1.291.36, WP/WH = 1.191.23, WE/WP = 1.691.79. Sides more strongly concave anterad of hind angles, the latter large, sometimes slightly bent outwards, slightly obtuse or rectangular. Base slightly curved anteriorly at outer fifth. Basal depression rugose both sides of middle of pronotum due to the presence of several small longitudinal wrinkles in addition to the convex surfaces of sculpticells. In all other characters agreeing with
<taxonomicName id="3C934D6F0319FF9BFC02FDAAFB96170F" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[954,1114,538,563]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFC02FDAAFB96170F" box="[954,1114,538,563]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFBD9FDAAFB6F1708" bold="true" box="[1121,1187,538,564]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFBD9FDAAFB6F1708" box="[1121,1187,538,564]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFDF1FB0B1794" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
Elytra: Broader oval, with proportion WE/LE = 1.501.55. Sides with shoulders almost evenly rounded. Striae more deeply punctate. Preapical seta is located close to the second stria and at the beginning of the posterior elytral seventh or eighth. In all other characters agreeing with
<taxonomicName id="3C934D6F0319FF9BFC66FD3FFBB21794" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[990,1150,655,680]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFC66FD3FFBB21794" box="[990,1150,655,680]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFB3DFD3EFB0B1794" bold="true" box="[1157,1223,654,680]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFB3DFD3EFB0B1794" box="[1157,1223,654,680]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFD05FE2417F3" blockId="55.[151,1436,152,1493]" box="[197,488,693,719]" pageId="55" pageNumber="56">Legs: Relatively slender.</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFD6CFA591656" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">Male genitalia: Aedeagal median lobe average in length (LE/LA = 2.692.88), more strongly curved in basal half, slightly elongate towards apex, but with terminal lamella short; the latter distinctly curved upwards, its base slightly stepped from level of ventral margin of median lobe. Basal bulb average. Internal sac with slightly sclerotized longitudinal sheets below median lobe ostium, without distinct copulatory piece.</paragraph>
</subSubSection>
<subSubSection id="B38965670319FF9BFF7DFCC6FAB1102F" pageId="55" pageNumber="56" type="etymology">
<paragraph id="FB2C36EC0319FF9BFF7DFCC6FABB168B" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
<emphasis id="C9E7EAFE0319FF9BFF7DFCC6FE9A16AC" bold="true" box="[197,342,886,912]" pageId="55" pageNumber="56">Etymology:</emphasis>
The specific name refers to the Buddhist monks of
<collectingRegion id="3957F80E0319FF9BFC03FCC6FC3616AC" box="[955,1018,886,912]" country="China" name="Xizang" pageId="55" pageNumber="56">Tibet</collectingRegion>
, and especially to the monks of the very old Tsurphu Monastery which is located near the
<typeStatus id="2428884E0319FF9BFCADFC2DFC8B168B" box="[789,839,925,951]" pageId="55" pageNumber="56">type</typeStatus>
locality of the new species; noun in apposition.
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFC73FB6611D1" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
<emphasis id="C9E7EAFE0319FF9BFF7DFC73FEB416E1" bold="true" box="[197,376,963,989]" pageId="55" pageNumber="56">Identification:</emphasis>
This new species is very similar to the above described
<taxonomicName id="3C934D6F0319FF9BFBB5FC74FB6116E1" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1037,1197,964,989]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFBB5FC74FB6116E1" box="[1037,1197,964,989]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFB0CFC73FB3A16E1" bold="true" box="[1204,1270,963,989]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFB0CFC73FB3A16E1" box="[1204,1270,963,989]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but differs in having antennae paler, pronotal hind angles larger, basal depressions more distinctly rugose, elytra broader oval, and especially by having aedeagal internal sac only weakly sclerotized. It is also similar to
<taxonomicName id="3C934D6F0319FF9BFB40FBA1FEF9116E" authority="Deuve, 1997" authorityName="Deuve" authorityYear="1997" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="folwarcznyi">
<emphasis id="C9E7EAFE0319FF9BFB40FBA1FA501116" box="[1272,1436,1041,1066]" italics="true" pageId="55" pageNumber="56">T. folwarcznyi</emphasis>
<bibRefCitation id="9F024B1D0319FF9BFF2FFB88FEF9116E" author="Deuve, T." box="[151,309,1080,1106]" pageId="55" pageNumber="56" pagination="139 - 154" refId="ref48219" refString="Deuve, T. (1997) Nouveaux Trechini du Tibet et des regions limitrophes (Coleoptera, Trechidae). Revue Francaise d'Entomologie (Nouvelle serie), 19 (3 - 4), 139 - 154." type="journal article" year="1997">Deuve, 1997</bibRefCitation>
</taxonomicName>
, but has larger body size, head, antennae and legs slender, temporae longer and elytra much broader. In male genitalia characters
<taxonomicName id="3C934D6F0319FF9BFDF2FBEFFD6D1144" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[586,673,1119,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="lama" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFDF2FBEFFD6D1144" box="[586,673,1119,1144]" italics="true" pageId="55" pageNumber="56">T. lama</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D2D457850319FF9BFD11FBEEFD241144" box="[681,744,1118,1144]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
can easily be distinguished from both
<taxonomicName id="3C934D6F0319FF9BFB09FBEFFA9E1144" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[1201,1362,1119,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="budhaensis" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFB09FBEFFA9E1144" box="[1201,1362,1119,1144]" italics="true" pageId="55" pageNumber="56">T. budhaensis</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFAE2FBEEFA501144" bold="true" box="[1370,1436,1118,1144]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFAE2FBEEFA501144" box="[1370,1436,1118,1144]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="3C934D6F0319FF9BFF74FB35FED511A2" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[204,281,1157,1182]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="yeti" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFF74FB35FED511A2" box="[204,281,1157,1182]" italics="true" pageId="55" pageNumber="56">T. yeti</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFE9AFB35FEA411A3" bold="true" box="[290,360,1157,1183]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFE9AFB35FEA411A3" box="[290,360,1157,1183]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, which also have more slender appendages, broader oval elytra and weak internal sac sclerotisation of aedeagal median lobe, by the much stouter aedeagal median lobe with a shorter terminal lamella which is slightly stepped from the level of the ventral margin of the median lobe.
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFB48FAB1102F" blockId="55.[151,1436,152,1493]" box="[197,1405,1272,1299]" pageId="55" pageNumber="56">
<emphasis id="C9E7EAFE0319FF9BFF7DFB48FEB4102E" bold="true" box="[197,376,1272,1298]" pageId="55" pageNumber="56">Relationships:</emphasis>
This species is the presumed sister species of
<taxonomicName id="3C934D6F0319FF9BFC2EFB49FBFA102E" authority="Schmidt, 2009" authorityName="Schmidt" authorityYear="2009" box="[918,1078,1273,1298]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="astrophilus" status="sp. nov.">
<emphasis id="C9E7EAFE0319FF9BFC2EFB49FBFA102E" box="[918,1078,1273,1298]" italics="true" pageId="55" pageNumber="56">T. astrophilus</emphasis>
</taxonomicName>
<emphasis id="C9E7EAFE0319FF9BFB85FB48FBB3102E" bold="true" box="[1085,1151,1272,1298]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="D2D457850319FF9BFB85FB48FBB3102E" box="[1085,1151,1272,1298]" pageId="55" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, see the latter, above.
</paragraph>
</subSubSection>
<subSubSection id="B38965670319FF9BFF7DFAAFFB5D10E9" pageId="55" pageNumber="56" type="distribution">
<paragraph id="FB2C36EC0319FF9BFF7DFAAFFDFF10BB" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
<emphasis id="C9E7EAFE0319FF9BFF7DFAAFFEA41005" bold="true" box="[197,360,1311,1337]" pageId="55" pageNumber="56">Distribution:</emphasis>
<figureCitation id="63A82A690319FF9BFED6FA90FE181006" box="[366,468,1312,1338]" captionStart="FIGURE 100" captionStartId="67.[151,256,1106,1130]" captionTargetBox="[151,1436,194,1082]" captionTargetId="figure-404@67.[151,1436,194,1082]" captionTargetPageId="67" captionText="FIGURE 100. Map of Southern Central Tibet showing distribution of species of the Trechus antonini group (white circles). 1, T. antonini Deuve, 1997. 2, T. religiosus sp. n. 3, T. astrophilus sp. n. 4, T. budhaensis sp. n. 5, T. rarus sp. n. 6, T. yeti sp. n. 7, T. folwarcznyi Deuve, 1997. 8, T. yak yak ssp. n. 9, T. yak shogulaensis ssp. n. 10, T. tseringi sp. n. 11, T. singularis sp. n. 12, T. tsampa sp. n. 13, T. lama sp. n." figureDoi="http://doi.org/10.5281/zenodo.5312320" httpUri="https://zenodo.org/record/5312320/files/figure.png" pageId="55" pageNumber="56">Fig. 100</figureCitation>
. Transhimalaya: Currently only known from the source area of a snow water brook on south western side of middle Tsubu Valley (or Tsurphu Valley) which is the north western side valley of the Kurum Valley north west of Lhasa.
</paragraph>
<paragraph id="FB2C36EC0319FF9BFF7DFA23FB5D10E9" blockId="55.[151,1436,152,1493]" pageId="55" pageNumber="56">
<emphasis id="C9E7EAFE0319FF9BFF7DFA23FEE21091" bold="true" box="[197,302,1427,1453]" pageId="55" pageNumber="56">Habitat:</emphasis>
Higher alpine zone. The specimens were found under stones on gently inclined slopes of eastern exposition and close to small melt water brooks at altitudes between 5100 and
<quantity id="3C6B9B090319FF9BFB88FA0BFB4010E9" box="[1072,1164,1467,1493]" metricMagnitude="3" metricUnit="m" metricValue="5.2" pageId="55" pageNumber="56" unit="m" value="5200.0">5200 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>