329 lines
37 KiB
XML
329 lines
37 KiB
XML
<document id="11CE14C1990722E2722997137710A909" ID-CLB-Dataset="112027" ID-DOI="10.11646/zootaxa.5169.3.3" ID-GBIF-Dataset="1f9fb092-7402-431a-beb4-89fee8ba6b9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952381" ID-ZooBank="4AD63AE5-0E34-4BF6-931B-1C013C19001E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659438748806" checkinUser="plazi" docAuthor="Martínez-Luque, Erick Omar, Jones, Robert Wallace & Niño-Maldonado, Santiago" docDate="2022" docId="52682200FFF8FFCA9882C60AFA7B4347" docLanguage="en" docName="zootaxa.5169.3.3.pdf" docOrigin="Zootaxa 5169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scaptolenus" docType="treatment" docVersion="5" lastPageNumber="277" masterDocId="AE515A78FFF1FFC09815C234FFE74725" masterDocTitle="New species, notes on the biology of Scaptolenus LeConte (Coleoptera: Elateridae) and a key to the “ vagans-group ”" masterLastPageNumber="278" masterPageNumber="267" pageNumber="276" updateTime="1699362632238" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="0E03CC8C06343FCB3C0FC9DCB29C143A">New species, notes on the biology of Scaptolenus LeConte (Coleoptera: Elateridae) and a key to the “ vagans-group ”</mods:title>
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<mods:name id="A3329C2404174935361F6B6B92B5420D" type="personal">
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<mods:namePart id="1D2A68E649276E4075025FD8B52B475B">Martínez-Luque, Erick Omar</mods:namePart>
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<mods:namePart id="9DC6FC3A59B11C76D5140E5F30D625BB">Jones, Robert Wallace</mods:namePart>
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<mods:affiliation id="DAC0706915288457DF7746A609020FD0">rjones @ uaq. mx; https: // orcid. org / 0000 - 0001 - 9125 - 8848</mods:affiliation>
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<mods:nameIdentifier id="929313FD283106D53315941C548CFAFB" type="email">rjones@uaq.mx</mods:nameIdentifier>
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<mods:namePart id="84AB8DFCAA1FE6F8E57DD20EC33ADBF3">Niño-Maldonado, Santiago</mods:namePart>
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<mods:affiliation id="D1907D83B898563C2AF5EE4B830D59EF">Facultad de Ingeniería y Ciencias, Universidad Autónoma de Tamaulipas, Centro Universitario Victoria, Ciudad Victoria, Tamaulipas, México. C. P. 87149</mods:affiliation>
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<mods:title id="57A25A9E442F01EA0CA7BF4C9888CD68">Zootaxa</mods:title>
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<mods:date id="05EA71AB0CC1C5607AA548089E93DC26">2022</mods:date>
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<mods:number id="3276B72F99824FD4EE7F9CCBB3DCFB98">2022-07-29</mods:number>
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<treatment id="52682200FFF8FFCA9882C60AFA7B4347" ID-DOI="http://doi.org/10.5281/zenodo.6958093" ID-GBIF-Taxon="198574535" ID-Zenodo-Dep="6958093" LSID="urn:lsid:plazi:treatment:52682200FFF8FFCA9882C60AFA7B4347" httpUri="http://treatment.plazi.org/id/52682200FFF8FFCA9882C60AFA7B4347" lastPageId="10" lastPageNumber="277" pageId="9" pageNumber="276">
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<subSubSection id="92DBC09DFFF8FFC99882C60AFDCA437C" box="[151,557,1086,1113]" pageId="9" pageNumber="276" type="nomenclature">
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<paragraph id="DA7E9316FFF8FFC99882C60AFDCA437C" blockId="9.[151,557,1086,1113]" box="[151,557,1086,1113]" pageId="9" pageNumber="276">
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<heading id="8136247AFFF8FFC99882C60AFDCA437C" bold="true" box="[151,557,1086,1113]" fontSize="11" level="1" pageId="9" pageNumber="276" reason="1">
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<emphasis id="E8B54F04FFF8FFC99882C60AFDCA437C" bold="true" box="[151,557,1086,1113]" pageId="9" pageNumber="276">
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Biology of the genus
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<taxonomicName id="1DC1E895FFF8FFC9998FC60AFDCA437C" box="[410,557,1086,1113]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC9998FC60AFDCA437C" bold="true" box="[410,557,1086,1113]" italics="true" pageId="9" pageNumber="276">Scaptolenus</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="92DBC09DFFF8FFCA9882C6B3FA994641" lastPageId="10" lastPageNumber="277" pageId="9" pageNumber="276" type="discussion">
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<paragraph id="DA7E9316FFF8FFC99882C6B3FC7C4174" blockId="9.[151,1437,1159,2013]" pageId="9" pageNumber="276">
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Very little is known about the biology of the
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<taxonomicName id="1DC1E895FFF8FFC99A80C6B3FCF64384" authorityName="Latreille" authorityYear="1802" box="[661,785,1159,1185]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="tribe" tribe="Cebrionini">Cebrionini</taxonomicName>
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. Much of the information is based on casual observations obtained in the field, mostly of the genus
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<taxonomicName id="1DC1E895FFF8FFC99A76C69FFCAD43E0" authority="Olivier, 1790" authorityName="Olivier" authorityYear="1790" box="[611,842,1195,1221]" class="Insecta" family="Elateridae" genus="Cebrio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99A76C69FFD5643E0" box="[611,689,1195,1221]" italics="true" pageId="9" pageNumber="276">Cebrio</emphasis>
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Olivier, 1790
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</taxonomicName>
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(mainly European), and on some observations generated during the collection of North American species of the genus
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<taxonomicName id="1DC1E895FFF8FFC99B6CC6FBFBE743CC" box="[889,1024,1231,1257]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99B6CC6FBFBE743CC" box="[889,1024,1231,1257]" italics="true" pageId="9" pageNumber="276">Scaptolenus</emphasis>
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</taxonomicName>
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. However, a large part of the species of the genus is described from localities of
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<collectingCountry id="A2D6D386FFF8FFC99A6DC6C7FD294228" box="[632,718,1267,1293]" name="Mexico" pageId="9" pageNumber="276">México</collectingCountry>
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and Central America. One of the first works in which biological data of this group of insects is presented is in
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<bibRefCitation id="BE50EEE7FFF8FFC99AB5C723FC5B4214" author="Guerin-Meneville, M. F. E." box="[672,956,1303,1329]" pageId="9" pageNumber="276" pagination="403 - 404" refId="ref7371" refString="Guerin-Meneville, M. F. E. (1844) Note sur la decouverte faite en Algerie, par M. de docteur Guyon, d´une nouvelle espece du genre Cebrio. Revue Zoologique, par la Societe Cuvierienne, 7, 403 - 404." type="journal article" year="1844">Guérin-Méneville (1844)</bibRefCitation>
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and
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<bibRefCitation id="BE50EEE7FFF8FFC99BE6C723FB7C4214" author="Werner, F. G." box="[1011,1179,1303,1329]" pageId="9" pageNumber="276" pagination="26 - 27" refId="ref8204" refString="Werner, F. G. (1969) Two flights of Scaptolenus (Coleoptera: Cebrionidae). The Coleopterists Bulletin, 23 (1), 26 - 27." type="journal article" year="1969">Werner (1969)</bibRefCitation>
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. The first observations of the behavior of female individuals is with
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<taxonomicName id="1DC1E895FFF8FFC99A89C70FFBDB4270" authority="(Fall, 1932)" baseAuthorityName="Fall" baseAuthorityYear="1932" box="[668,1084,1339,1365]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="species" species="fuscipennis">
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<emphasis id="E8B54F04FFF8FFC99A89C70FFC4D4270" box="[668,938,1339,1365]" italics="true" pageId="9" pageNumber="276">Scaptolenus fuscipennis</emphasis>
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(
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<bibRefCitation id="BE50EEE7FFF8FFC99BAEC70FFBD34270" author="Fall, H. C." box="[955,1076,1339,1365]" pageId="9" pageNumber="276" pagination="56 - 62" refId="ref7261" refString="Fall, H. C. (1932) New Coleoptera XV. The Canadian Entomologist, 64 (3), 56 - 62. https: // doi. org / 10.4039 / Ent 6456 - 3" type="journal article" year="1932">Fall, 1932</bibRefCitation>
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)
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</taxonomicName>
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. Collected females of this species were found partially buried in the ground, from where they do not fully emerge which is very similar to that observed in
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<collectingCountry id="A2D6D386FFF8FFC99931C7B7FE8042B8" box="[292,359,1411,1437]" name="Spain" pageId="9" pageNumber="276">Spain</collectingCountry>
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,
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<collectingCountry id="A2D6D386FFF8FFC99966C7B7FE4142B8" box="[371,422,1411,1437]" name="Italy" pageId="9" pageNumber="276">Italy</collectingCountry>
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and
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<collectingCountry id="A2D6D386FFF8FFC999F4C7B7FDCA42B8" box="[481,557,1411,1437]" name="France" pageId="9" pageNumber="276">France</collectingCountry>
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with females of the genus
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<taxonomicName id="1DC1E895FFF8FFC99B73C7B7FC5342B8" authorityName="Olivier" authorityYear="1790" box="[870,948,1411,1437]" class="Insecta" family="Elateridae" genus="Cebrio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99B73C7B7FC5342B8" box="[870,948,1411,1437]" italics="true" pageId="9" pageNumber="276">Cebrio</emphasis>
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</taxonomicName>
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. In this genus, females are exposed on the ground and males emerge and fly in search of the females for copulation during and after rainstorms (
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<bibRefCitation id="BE50EEE7FFF8FFC99D12C793FA7042E4" author="Mittre, M. H." box="[1287,1431,1447,1473]" pageId="9" pageNumber="276" pagination="53 - 61" refId="ref7936" refString="Mittre, M. H. (1839) Notice sur l´accouplement du Cebrio gigas. Revue Zoologie par la Societe Cuvierienne, 2, 53 - 61." type="journal article" year="1839">Mittré, 1839</bibRefCitation>
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;
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<bibRefCitation id="BE50EEE7FFF8FFC99882C7FFFEA542C0" author="Moroder, S. E." box="[151,322,1483,1509]" pageId="9" pageNumber="276" pagination="5 - 8" refId="ref7965" refString="Moroder, S. E. (1920) Sobre los Cebrio carbonarius y superbus en la region valenciana. Anales del Instituto General y Tecnico de Valencia, 4, 5 - 8." type="journal article" year="1920">Moroder, 1920</bibRefCitation>
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;
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<bibRefCitation id="BE50EEE7FFF8FFC9995AC7FFFE3142C0" author="Rattu, R." box="[335,470,1483,1509]" pageId="9" pageNumber="276" pagination="45 - 50" refId="ref8000" refString="Rattu, R. (2012) Osserbazioni sulla biologia di Cebrio sardous Perris, 1869 (Insecta, Coleoptera, Elateridae, Cebrioninae). Bollettino del Museo di Storia Naturale di Naturale di Venezia, 63, 45 - 50." type="journal article" year="2012">Rattu, 2012</bibRefCitation>
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;
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<bibRefCitation id="BE50EEE7FFF8FFC999F7C7FFFDF942C0" author="Rattu, R." box="[482,542,1483,1509]" pageId="9" pageNumber="276" pagination="235 - 248" refId="ref8043" refString="Rattu, R. (2016) Contributo alla conosceza delle femmine delle specie sarde di Cebrio (Coleoptera, Elateridae, Elaterina, Cebrionini). Annali del Museo Civico de Storia Naturale " Giacomo Doria ", 108, 235 - 248." type="journal article" year="2016">2016</bibRefCitation>
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;
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<bibRefCitation id="BE50EEE7FFF8FFC99A3FC7FFFC6442C0" author="Zapata, J. L. & Sanchez-Ruiz, A." box="[554,899,1483,1509]" pageId="9" pageNumber="276" pagination="125 - 136" refId="ref8236" refString="Zapata, J. L. & Sanchez-Ruiz, A. (2016) Datos ara el estudio del genero Cebrio Olivier, 1790: las especies del grupo Cebrio amorii (Coleoptera: Elateridae: Elaterinae: Cebrionini). Arquivos Entomoloxicos Galegos, 15, 125 - 136." type="journal article" year="2016">Zapata & Sánchez-Ruiz, 2016</bibRefCitation>
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). Another genus of
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<taxonomicName id="1DC1E895FFF8FFC99C74C7FFFB3E42C0" authorityName="Latreille" authorityYear="1802" box="[1121,1241,1483,1509]" class="Insecta" family="Elateridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="tribe" tribe="Cebrionini">Cebrionini</taxonomicName>
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that has been reported with very similar activities in females and males, are some species of the genus
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<taxonomicName id="1DC1E895FFF8FFC99C74C7DBFA7F412C" authority="(Latreille, 1834)" baseAuthorityName="Latreille" baseAuthorityYear="1834" box="[1121,1432,1519,1545]" class="Insecta" family="Elateridae" genus="Selonodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99C74C7DBFB30412C" box="[1121,1239,1519,1545]" italics="true" pageId="9" pageNumber="276">Selonodon</emphasis>
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(
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<bibRefCitation id="BE50EEE7FFF8FFC99CF3C7DBFA77412C" author="Latreille, P. A." box="[1254,1424,1519,1545]" pageId="9" pageNumber="276" pagination="113 - 170" refId="ref7501" refString="Latreille, P. A. (1834) Distribution methodique et naturelle des genres de diverses tribus d'insectes Colepteres de la famille des serricornes. Annual de la Societe Entomologique de France, 3, 113 - 170." type="journal article" year="1834">Latreille, 1834</bibRefCitation>
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)
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</taxonomicName>
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, where the females of some species of the genus remain partially exposed in the soil, and the males present a very marked flight activity during the heavy summer rains (
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<bibRefCitation id="BE50EEE7FFF8FFC99AEFC403FC684174" author="Galley, K. E. M." box="[762,911,1591,1617]" pageId="9" pageNumber="276" pagination="1 - 48" refId="ref7300" refString="Galley, K. E. M. (1999) Revision of the genus Selonodon Latreille (Coleoptera: Cebrionidae). Occasional Papers of the Florida State Collection of Arthropods, 10, 1 - 48." type="journal article" year="1999">Galley, 1999</bibRefCitation>
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).
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</paragraph>
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<paragraph id="DA7E9316FFF8FFC998D2C46FFB044054" blockId="9.[151,1437,1159,2013]" pageId="9" pageNumber="276">
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We observed this behavior in
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<collectingRegion id="18055DF4FFF8FFC99A18C46FFD674150" box="[525,640,1627,1653]" country="Mexico" name="Queretaro de Arteaga" pageId="9" pageNumber="276">Querétaro</collectingRegion>
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,
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<collectingCountry id="A2D6D386FFF8FFC99A9FC46FFD054150" box="[650,738,1627,1653]" name="Mexico" pageId="9" pageNumber="276">México</collectingCountry>
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, for an undescribed species of
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<taxonomicName id="1DC1E895FFF8FFC99C26C46FFB5D4150" box="[1075,1210,1627,1653]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99C26C46FFB5D4150" box="[1075,1210,1627,1653]" italics="true" pageId="9" pageNumber="276">Scaptolenus</emphasis>
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</taxonomicName>
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, where we collected
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<specimenCount id="CCC7589FFFF8FFC99882C44AFF1641BC" box="[151,241,1662,1689]" pageId="9" pageNumber="276" type="male">45 male</specimenCount>
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individuals on the university campus in Juriquilla after one of the first heavy seasonal monsoon rains. Interestingly, all males died in less than 24 hours, suggesting that mating flights are very short and male mortality is high, resulting in a short opportunity for collection, regardless of population size. Subsequently, a search was started at various points in the soil of the scrub vegetation surrounding the campus and some larvae of the genus were collected. In addition, the probable apterous condition of the females and their behavior of remaining buried limit the chances of detection and collection. This also suggests that the genus
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<taxonomicName id="1DC1E895FFF8FFC99B8DC507FBF84068" box="[920,1055,1843,1869]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="276" phylum="Arthropoda" rank="genus">
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<emphasis id="E8B54F04FFF8FFC99B8DC507FBF84068" box="[920,1055,1843,1869]" italics="true" pageId="9" pageNumber="276">Scaptolenus</emphasis>
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</taxonomicName>
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may be more common than previously reported, and that collecting larval forms may give a better estimation of population density.
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</paragraph>
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<paragraph id="DA7E9316FFF8FFCA98D2C54FFA9E47DC" blockId="9.[151,1437,1159,2013]" lastBlockId="10.[151,1436,151,249]" lastPageId="10" lastPageNumber="277" pageId="9" pageNumber="276">
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In the case of the new species presented in this work, there are no data associated with their biology. However, the periods in which the specimens of the
|
||
<typeStatus id="057A2DB4FFF8FFC99A79C5ABFD7B409C" box="[620,668,1951,1977]" pageId="9" pageNumber="276">type</typeStatus>
|
||
series of both species were collected coincide with the rainy seasons in
|
||
<collectingCountry id="A2D6D386FFF8FFC998A6C5F7FEEC40F8" box="[179,267,1987,2013]" name="Mexico" pageId="9" pageNumber="276">Mexico</collectingCountry>
|
||
, which is linked to high levels of environmental humidity and this in turn is related to a flush of vegetative growth and root growth. These rains also produce water saturation and soften soils, which would presumably favor the movement of the larvae and emergence of adults of these edaphic organisms. This seasonality is probably closely linked to the life history events of
|
||
<taxonomicName id="1DC1E895FFFBFFCA9A20C2EBFD5B47DC" box="[565,700,223,249]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E8B54F04FFFBFFCA9A20C2EBFD5B47DC" box="[565,700,223,249]" italics="true" pageId="10" pageNumber="277">Scaptolenus</emphasis>
|
||
</taxonomicName>
|
||
in these highly seasonal habitats of tropical deciduous forests.
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C37EFA994641" blockId="10.[151,1406,330,357]" box="[151,1406,330,357]" pageId="10" pageNumber="277">
|
||
<heading id="8136247AFFFBFFCA9882C37EFA994641" bold="true" box="[151,1406,330,357]" fontSize="11" level="1" pageId="10" pageNumber="277" reason="1">
|
||
<emphasis id="E8B54F04FFFBFFCA9882C37EFA994641" bold="true" box="[151,1406,330,357]" pageId="10" pageNumber="277">
|
||
A key to the groups of
|
||
<taxonomicName id="1DC1E895FFFBFFCA99A4C37EFDA34640" box="[433,580,330,357]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E8B54F04FFFBFFCA99A4C37EFDA34640" bold="true" box="[433,580,330,357]" italics="true" pageId="10" pageNumber="277">Scaptolenus</emphasis>
|
||
</taxonomicName>
|
||
(males) (based on
|
||
<bibRefCitation id="BE50EEE7FFFBFFCA9B39C37EFC1A4641" author="Champion, G. C." box="[812,1021,330,356]" pageId="10" pageNumber="277" pagination="557 - 573" refId="ref7152" refString="Champion, G. C. (1896) Fam. Cebrionidae. In: Goodman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 3. Part 1. Serraticornia. R H. Porter, London. pp. 557 - 573." type="book chapter" year="1896">Champion, 1896</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="BE50EEE7FFFBFFCA9C1EC37FFB624641" author="Fall, H. C." box="[1035,1157,330,357]" pageId="10" pageNumber="277" pagination="56 - 62" refId="ref7261" refString="Fall, H. C. (1932) New Coleoptera XV. The Canadian Entomologist, 64 (3), 56 - 62. https: // doi. org / 10.4039 / Ent 6456 - 3" type="journal article" year="1932">Fall, 1932</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="BE50EEE7FFFBFFCA9CD6C37FFA914641" author="Johnson, P. J." box="[1219,1398,330,357]" pageId="10" pageNumber="277" pagination="105 - 110" refId="ref7450" refString="Johnson, P. J. (2013) A new species of Scaptolenus LeConte (Coleoptera: Elateridae: Cebrioninae) from Sonora, Mexico, with a checklist of species of northern Mexico and The United States. Dugesiana, 20 (2), 105 - 110." type="journal article" year="2013">Johnson, 2013</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92DBC09DFFFBFFCA9882C3A4FA7A46D8" pageId="10" pageNumber="277" type="key">
|
||
<key id="AE50C427FFFBFFCA9882C3A4FA7A46D8" pageId="10" pageNumber="277">
|
||
<keyStep id="61358BB3FFFBFFCA9882C3A4FA7B46E7" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C3A4FA7A4680" blockId="10.[151,1437,400,510]" box="[151,1437,400,422]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C3A4FA7A4680" box="[151,1437,400,422]" pageId="10" pageNumber="277">
|
||
1. Antennomeres with subparallel shape.........................................................
|
||
<emphasis id="E8B54F04FFFBFFCA9CE9C3A4FABB4680" box="[1276,1372,400,421]" italics="true" pageId="10" pageNumber="277">chevrolati</emphasis>
|
||
-group
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C399FA7B46E7" blockId="10.[151,1437,400,510]" box="[151,1436,429,451]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C399FA7B46E7" box="[151,1436,429,451]" pageId="10" pageNumber="277">- Antennomeres without subparallel shape................................................................... 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C3FEFA7A46D8" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C3FEFA7A46C5" blockId="10.[151,1437,400,510]" box="[151,1437,458,480]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C3FEFA7A46C5" box="[151,1437,458,480]" pageId="10" pageNumber="277">
|
||
2. Antennomeres angularly extended, “L” shaped....................................................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D0DC3FFFABB46C5" box="[1304,1372,459,480]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="vagans">
|
||
<emphasis id="E8B54F04FFFBFFCA9D0DC3FFFABB46C5" box="[1304,1372,459,480]" italics="true" pageId="10" pageNumber="277">vagans</emphasis>
|
||
</taxonomicName>
|
||
-group
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C3DCFA7A46D8" blockId="10.[151,1437,400,510]" box="[151,1437,488,510]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C3DCFA7A46D8" box="[151,1437,488,510]" pageId="10" pageNumber="277">
|
||
- Antennomeres serrated (in all its variants).........................................................
|
||
<emphasis id="E8B54F04FFFBFFCA9D36C3DCFABB46D8" box="[1315,1372,488,509]" italics="true" pageId="10" pageNumber="277">fulvus</emphasis>
|
||
-group
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="92DBC09DFFFBFFCA9882C064FAA2454F" box="[151,1349,591,618]" pageId="10" pageNumber="277" type="discussion">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C064FAA2454F" blockId="10.[151,1349,591,618]" box="[151,1349,591,618]" pageId="10" pageNumber="277">
|
||
<heading id="8136247AFFFBFFCA9882C064FAA2454F" bold="true" box="[151,1349,591,618]" fontSize="11" level="1" pageId="10" pageNumber="277" reason="1">
|
||
<emphasis id="E8B54F04FFFBFFCA9882C064FAA2454F" bold="true" box="[151,1349,591,618]" pageId="10" pageNumber="277">
|
||
A Key to the
|
||
<taxonomicName id="1DC1E895FFFBFFCA992FC064FE68454F" box="[314,399,592,618]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="vagans">
|
||
<emphasis id="E8B54F04FFFBFFCA992FC064FE68454F" bold="true" box="[314,399,592,618]" italics="true" pageId="10" pageNumber="277">vagans</emphasis>
|
||
</taxonomicName>
|
||
-group of
|
||
<taxonomicName id="1DC1E895FFFBFFCA9A1FC07BFD7A454F" box="[522,669,591,618]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E8B54F04FFFBFFCA9A1FC07BFD7A454F" bold="true" box="[522,669,591,618]" italics="true" pageId="10" pageNumber="277">Scaptolenus</emphasis>
|
||
</taxonomicName>
|
||
(males) (based on
|
||
<bibRefCitation id="BE50EEE7FFFBFFCA9B93C064FBDF454F" author="Johnson, P. J." box="[902,1080,592,618]" pageId="10" pageNumber="277" pagination="105 - 110" refId="ref7450" refString="Johnson, P. J. (2013) A new species of Scaptolenus LeConte (Coleoptera: Elateridae: Cebrioninae) from Sonora, Mexico, with a checklist of species of northern Mexico and The United States. Dugesiana, 20 (2), 105 - 110." type="journal article" year="2013">Johnson, 2013</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="BE50EEE7FFFBFFCA9C63C064FADA454F" author="Champion, G. C." box="[1142,1341,592,618]" pageId="10" pageNumber="277" pagination="557 - 573" refId="ref7152" refString="Champion, G. C. (1896) Fam. Cebrionidae. In: Goodman, F. D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 3. Part 1. Serraticornia. R H. Porter, London. pp. 557 - 573." type="book chapter" year="1896">Champion 1896</bibRefCitation>
|
||
)
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="92DBC09DFFFBFFCA9882C0A1FA7B4347" pageId="10" pageNumber="277" type="key">
|
||
<key id="AE50C427FFFBFFCA9882C0A1FA7B4347" pageId="10" pageNumber="277">
|
||
<keyStep id="61358BB3FFFBFFCA9882C0A1FA7B45E2" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C0A1FA7B458F" blockId="10.[151,1437,661,1123]" box="[151,1436,661,683]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C0A1FA7B458F" box="[151,1436,661,683]" pageId="10" pageNumber="277">1. Antennomeres 4-10 slightly apically dilated................................................................ 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C087FA7B45E2" blockId="10.[151,1437,661,1123]" box="[151,1436,690,712]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C087FA7B45E2" box="[151,1436,690,712]" pageId="10" pageNumber="277">
|
||
- Antennomers 4-10 clearly dilated to subangulated ventrally at the apices..............................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D1AC087FA7B45E2" box="[1295,1436,690,712]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="amplipennis">
|
||
<emphasis id="E8B54F04FFFBFFCA9D1AC087FA7B45E2" box="[1295,1436,690,712]" italics="true" pageId="10" pageNumber="277">S. amplipennis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C0E4FA7B4427" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C0E4FA7B45C0" blockId="10.[151,1437,661,1123]" box="[151,1436,720,742]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C0E4FA7B45C0" box="[151,1436,720,742]" pageId="10" pageNumber="277">2. Elytra with inter-grooves flat or slightly elevated; antennae do not reach the middle length of the elytra................. 3</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C0D9FA7B4427" blockId="10.[151,1437,661,1123]" box="[151,1436,749,771]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C0D9FA7B4427" box="[151,1436,749,771]" pageId="10" pageNumber="277">- Elytra with clearly raised striae, giving a sulcate appearance of the striae; antennae longer extending to elytral mid-length.. 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C13EFA7B44B0" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C13EFA7B4418" blockId="10.[151,1437,661,1123]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C13EFA7B4418" pageId="10" pageNumber="277">
|
||
3. Pronotum wider than the elytral base; head, thorax and scutellum densely covered with long and decumbent fulvous setae; parameres without small spurs at the genital base.....................................................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D55C11CFA7B4418" box="[1344,1436,808,829]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="vagans">
|
||
<emphasis id="E8B54F04FFFBFFCA9D55C11CFA7B4418" box="[1344,1436,808,829]" italics="true" pageId="10" pageNumber="277">S. vagans</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C171FA7B44B0" blockId="10.[151,1437,661,1123]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C171FA7B44B0" pageId="10" pageNumber="277">
|
||
- Pronotum similar in length to the elytral base; head, thorax and scutellum region, moderately lined with separate setation, setae long, fulvous and decumbent; parameres with small sclerotized spines in the first third very close to the genital base, these spines project towards the base and pass behind the middle lobe (
|
||
<figureCitation id="42FA8F93FFFBFFCA9B41C1B4FC7D44B0" box="[852,922,896,918]" captionStart="FIGURE 6–10" captionStartId="5.[151,250,1729,1753]" captionTargetBox="[161,1427,289,1689]" captionTargetId="figure-64@5.[151,1435,270,1702]" captionTargetPageId="5" captionText="FIGURE 6–10. Scaptolenus zuritai sp. nov.; Figs. 6, 7, 8 dorsal, ventral and lateral view; Figs. 9, 10 ventral and dorsal view of the aedeagus. PHA: phallobase; PAR: paramerous; ML: median lobe; SG: setation group." figureDoi="http://doi.org/10.5281/zenodo.6952385" httpUri="https://zenodo.org/record/6952385/files/figure.png" pageId="10" pageNumber="277">Fig. 10</figureCitation>
|
||
)..........................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9CE6C1B4FAAD44B0" authority="Martínez-Luque & Jones & Niño-Maldonado, 2022" authorityName="Martínez-Luque & Jones & Niño-Maldonado" authorityYear="2022" box="[1267,1354,896,917]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="zuritai" status="sp. nov.">
|
||
<emphasis id="E8B54F04FFFBFFCA9CE6C1B4FAAD44B0" box="[1267,1354,896,917]" italics="true" pageId="10" pageNumber="277">S. zuritai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8B54F04FFFBFFCA9D45C1B4FA7B44B0" bold="true" box="[1360,1436,896,917]" pageId="10" pageNumber="277">
|
||
<taxonomicNameLabel id="F386F27FFFFBFFCA9D45C1B4FA7B44B0" box="[1360,1436,896,917]" pageId="10" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C1A9FA7B44F5" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C1A9FA7B4496" blockId="10.[151,1437,661,1123]" box="[151,1436,925,947]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C1A9FA7B4496" box="[151,1436,925,947]" pageId="10" pageNumber="277">4. Head, pronotum and scutellum black, which contrast with the chestnut elytra...................................... 6</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C18FFA7B44F5" blockId="10.[151,1437,661,1123]" box="[151,1436,954,976]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C18FFA7B44F5" box="[151,1436,954,976]" pageId="10" pageNumber="277">- Head black, pronotum and scutellum with different coloration from the head...................................... 5</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C1E3FA7B4302" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C1E3FA7A44C8" blockId="10.[151,1437,661,1123]" box="[151,1437,983,1006]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C1E3FA7A44C8" box="[151,1437,983,1006]" pageId="10" pageNumber="277">
|
||
5. Pronotum and scutellum light brown in color similar to the coloration of the elytra; fulvus colored legs.........
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D2FC1ECFA7A44C8" authorityName="Johnson" authorityYear="2013" box="[1338,1437,984,1005]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="paltingi">
|
||
<emphasis id="E8B54F04FFFBFFCA9D2FC1ECFA7A44C8" box="[1338,1437,984,1005]" italics="true" pageId="10" pageNumber="277">S. paltingi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C1C1FA7B4302" blockId="10.[151,1437,661,1123]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C1C1FA7B4302" pageId="10" pageNumber="277">
|
||
- Pronotum and scutellum of testaceous color (yellow-brownish) and elytra of brown color; fuscous colored legs (
|
||
<figureCitation id="42FA8F93FFFBFFCA9D02C1C1FA8B432F" box="[1303,1388,1013,1035]" captionStart="FIGURE 1–5" captionStartId="3.[151,250,1933,1957]" captionTargetBox="[163,1433,316,1889]" captionTargetId="figure-86@3.[154,1433,306,1906]" captionTargetPageId="3" captionText="FIGURE 1–5. Scaptolenus pacofierroi sp. nov.; Figs. 1, 2, 3 dorsal, ventral and lateral view; Figs. 4, 5 ventral and dorsal view of the aedeagus. PHA: phallobase; PAR: paramere; ML: median lobe; SG: setation group." figureDoi="http://doi.org/10.5281/zenodo.6952383" httpUri="https://zenodo.org/record/6952383/files/figure.png" pageId="10" pageNumber="277">Figs. 1-3</figureCitation>
|
||
).....................................................................................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9CD3C627FAAE4302" authority="Martínez-Luque & Jones & Niño-Maldonado, 2022" authorityName="Martínez-Luque & Jones & Niño-Maldonado" authorityYear="2022" box="[1222,1353,1042,1064]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="pacofierroi" status="sp. nov.">
|
||
<emphasis id="E8B54F04FFFBFFCA9CD3C627FAAE4302" box="[1222,1353,1042,1064]" italics="true" pageId="10" pageNumber="277">S. pacofierroi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E8B54F04FFFBFFCA9D45C626FA7B4302" bold="true" box="[1360,1436,1042,1063]" pageId="10" pageNumber="277">
|
||
<taxonomicNameLabel id="F386F27FFFFBFFCA9D45C626FA7B4302" box="[1360,1436,1042,1063]" pageId="10" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="61358BB3FFFBFFCA9882C604FA7B4347" pageId="10" pageNumber="277">
|
||
<paragraph id="DA7E9316FFFBFFCA9882C604FA7B4360" blockId="10.[151,1437,661,1123]" box="[151,1436,1072,1094]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C604FA7B4360" box="[151,1436,1072,1094]" pageId="10" pageNumber="277">
|
||
6. Posterolateral angles of pronotum subparallel; fuscous-colored antennae.................................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D2DC604FA7B4360" box="[1336,1436,1072,1093]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="lecontei">
|
||
<emphasis id="E8B54F04FFFBFFCA9D2DC604FA7B4360" box="[1336,1436,1072,1093]" italics="true" pageId="10" pageNumber="277">S. lecontei</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="DA7E9316FFFBFFCA9882C679FA7B4347" blockId="10.[151,1437,661,1123]" box="[151,1436,1101,1123]" pageId="10" pageNumber="277">
|
||
<keyLead id="61303023FFFBFFCA9882C679FA7B4347" box="[151,1436,1101,1123]" pageId="10" pageNumber="277">
|
||
- Posterolateral angles of pronotum divergent; fusco-testaceous colored antennae........................
|
||
<taxonomicName id="1DC1E895FFFBFFCA9D1AC679FA7B4347" authorityName="Champion" authorityYear="1896" box="[1295,1436,1101,1122]" class="Insecta" family="Elateridae" genus="Scaptolenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="E8B54F04FFFBFFCA9D1AC679FA7B4347" box="[1295,1436,1101,1122]" italics="true" pageId="10" pageNumber="277">S. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |