297 lines
38 KiB
XML
297 lines
38 KiB
XML
<document id="58DB89246E603BC0F9042C8F5BD4F142" ID-DOI="10.11646/palaeoentomology.6.3.12" ID-ISSN="2624-2834" ID-Zenodo-Dep="8209281" ID-ZooBank="0B00D332-FB99-48C2-9ECE-9D62BEF3A851" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691005694799" checkinUser="plazi" docAuthor="BRAZIDEC, MANUEL" docDate="2023" docId="525187E7FFBDFF9DF4CFF970FCA4B98D" docLanguage="en" docName="palaeoentomology.6.3.12.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.12" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Argemiones stupeflip BRAZIDEC 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="297" masterDocId="AE68FF9FFFBFFF98F447FF84FFAEBB49" masterDocTitle="Argemiones stupeflip gen. et sp. nov., a new spathiopterygid wasp (Hymenoptera: Diaprioidea) from the mid-Cretaceous Kachin amber" masterLastPageNumber="299" masterPageNumber="292" pageNumber="294" updateTime="1691006100581" updateUser="ExternalLinkService">
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<mods:title id="76B8933DD4F5D82490D563A3D8DB643F">Argemiones stupeflip gen. et sp. nov., a new spathiopterygid wasp (Hymenoptera: Diaprioidea) from the mid-Cretaceous Kachin amber</mods:title>
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</mods:titleInfo>
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<mods:name id="DF808A3197D6A4583141AEEB2DC01C75" type="personal">
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<mods:roleTerm id="BAB7706D7B2520FFF7939428904F6BC6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BB430130B986E34D17BC8C5157F0BB6E">BRAZIDEC, MANUEL</mods:namePart>
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<mods:typeOfResource id="FC4C45FF90580905C06D69AA3638AC7E">text</mods:typeOfResource>
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<mods:title id="5C994108C27465BEDAE8EDE0D60104C9">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="669A6B0289DAF1AE85355BC1D5C8772A">
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<mods:date id="398882C2B42180387189F9D446FC7245">2023</mods:date>
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<mods:number id="0AEE1C4B43E0DC0E29D8DF05DE4AC151">2023-06-28</mods:number>
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<mods:number id="106282F6F6A29D8B136BE45E43A69D86">6</mods:number>
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<mods:detail id="1B2D63D8772FFCC9D338ECCEA64E0455" type="issue">
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<mods:number id="AE573280C6B407B274881213A90389AD">3</mods:number>
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<mods:start id="C7A9D0D3449BD043D0061391EC471FE6">292</mods:start>
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<mods:url id="A643C4A81F09F571D9D1AAF8B3FD2163">http://dx.doi.org/10.11646/palaeoentomology.6.3.12</mods:url>
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</mods:location>
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<mods:classification id="C076BBF9337B690095AEF03C062AA1B8">journal article</mods:classification>
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<mods:identifier id="E91534EA1B72D181AD466EC6C8CB2611" type="DOI">10.11646/palaeoentomology.6.3.12</mods:identifier>
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<mods:identifier id="865F2F66F6DAB983418471814FD6CDBB" type="ISSN">2624-2834</mods:identifier>
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<mods:identifier id="CC2DA5E24543F83E35D734FC04572112" type="Zenodo-Dep">8209281</mods:identifier>
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<mods:identifier id="66FF8F5069DCF88EA0FE5361A501260C" type="ZooBank">0B00D332-FB99-48C2-9ECE-9D62BEF3A851</mods:identifier>
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</mods:mods>
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<treatment id="525187E7FFBDFF9DF4CFF970FCA4B98D" LSID="urn:lsid:plazi:treatment:525187E7FFBDFF9DF4CFF970FCA4B98D" httpUri="http://treatment.plazi.org/id/525187E7FFBDFF9DF4CFF970FCA4B98D" lastPageId="5" lastPageNumber="297" pageId="2" pageNumber="294">
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<subSubSection id="92E2657AFFBDFF9AF4CFF970FE71BC47" box="[136,479,1780,1807]" pageId="2" pageNumber="294" type="nomenclature">
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<paragraph id="DA4736F1FFBDFF9AF4CFF970FE71BC47" blockId="2.[136,778,1780,1914]" box="[136,479,1780,1807]" pageId="2" pageNumber="294">
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<heading id="810F819DFFBDFF9AF4CFF970FE71BC47" bold="true" box="[136,479,1780,1807]" fontSize="10" level="2" pageId="2" pageNumber="294" reason="3">
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<emphasis id="E88CEAE3FFBDFF9AF4CFF970FE71BC47" bold="true" box="[136,479,1780,1807]" pageId="2" pageNumber="294">
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<taxonomicName id="1DF84D72FFBDFF9AF4CFF970FED0BC46" authority="BRAZIDEC, 2023" authorityName="BRAZIDEC" authorityYear="2023" box="[136,382,1780,1807]" class="Insecta" family="Spathiopterygidae" genus="Argemiones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="294" phylum="Arthropoda" rank="species" species="stupeflip" status="sp. nov.">
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<emphasis id="E88CEAE3FFBDFF9AF4CFF970FED0BC46" bold="true" box="[136,382,1780,1807]" italics="true" pageId="2" pageNumber="294">Argemiones stupeflip</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F3BF5798FFBDFF9AF5C3F970FE71BC47" box="[388,479,1780,1806]" pageId="2" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9AF4CFF89CFE94BC33" pageId="2" pageNumber="294" type="description">
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<paragraph id="DA4736F1FFBDFF9AF4CFF89CFE94BC33" blockId="2.[136,778,1780,1914]" pageId="2" pageNumber="294">
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(
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<figureCitation id="42C32A74FFBDFF9AF4D7F89CFF5ABC7B" box="[144,244,1816,1842]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="2.[136,235,1354,1378]" captionStartId-1="3.[136,235,1428,1452]" captionStartId-2="4.[136,235,1478,1502]" captionTargetBox-0="[219,1367,179,1328]" captionTargetBox-1="[312,1275,189,1404]" captionTargetBox-2="[151,1436,170,1455]" captionTargetId-0="figure-266@2.[219,1367,179,1328]" captionTargetId-1="figure-237@3.[312,1275,189,1405]" captionTargetId-2="figure-205@4.[151,1436,170,1455]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 2. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, habitus in dorsal view. Scale bar = 1 mm." captionText-1="FIGURE 3. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, line drawing of habitus in dorsal view (pst = parastigma). Scale bar = 1 mm." captionText-2="FIGURE 4. Argemiones stupeflip gen. et sp. nov., specimen IGR.BU-076, A, Habitus in ventral view. B, Right fore wing and metasoma in dorsal view (arrow = remnant of hind wing). C, Left antenna in ventral view. D, Left mesotarsomere 5 in ventral view. Scale bars: A = 1 mm, B = 0.5 mm, C = 0.25 mm, D = 0.125 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8209285" figureDoi-1="http://doi.org/10.5281/zenodo.8209287" figureDoi-2="http://doi.org/10.5281/zenodo.8209289" httpUri-0="https://zenodo.org/record/8209285/files/figure.png" httpUri-1="https://zenodo.org/record/8209287/files/figure.png" httpUri-2="https://zenodo.org/record/8209289/files/figure.png" pageId="2" pageNumber="294">Figs 2–4</figureCitation>
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)
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<uri id="AE693AF3FFBDFF9AF4CFF8B8FE94BC33" pageId="2" pageNumber="294">
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||
urn:lsid:zoobank.org:act:
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<uuid id="AE5E0C24FFBDFF9AF5E7F8B8FE94BC33" pageId="2" pageNumber="294">BA3F9D4C-8803-4BF8-AE49- 05C75129137C</uuid>
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</uri>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9AF4CFF823FCACB365" pageId="2" pageNumber="294" type="materials_examined">
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<paragraph id="DA4736F1FFBDFF9AF4CFF823FCACB365" blockId="2.[136,778,1959,2093]" pageId="2" pageNumber="294">
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<emphasis id="E88CEAE3FFBDFF9AF4CFF823FF59BC88" bold="true" box="[136,247,1959,1985]" pageId="2" pageNumber="294">Material.</emphasis>
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<materialsCitation id="6A903CACFFBDFF9AF4B8F823FCACB365" country="France" county="University of Rennes" location="Geology Department" municipality="Museum" pageId="2" pageNumber="294" specimenCount="1" typeStatus="holotype">
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<typeStatus id="05438853FFBDFF9AF4B8F823FEC9BC88" box="[255,359,1959,1985]" pageId="2" pageNumber="294">Holotype</typeStatus>
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male IGR.BU-076, a well-preserved male, only lacking left pedicel, flagellomeres 1 and 2; housed in the amber collection of the
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<location id="DF27602AFFBDFF9AF667F86AFCA4B340" LSID="urn:lsid:plazi:treatment:525187E7FFBDFF9DF4CFF970FCA4B98D:DF27602AFFBDFF9AF667F86AFCA4B340" box="[544,778,2030,2057]" country="France" county="University of Rennes" municipality="Museum" name="Geology Department" pageId="2" pageNumber="294">Geology Department</location>
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and
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<collectingMunicipality id="3A23AC8BFFBDFF9AF4F0F797FEB7B364" box="[183,281,2067,2093]" pageId="2" pageNumber="294">Museum</collectingMunicipality>
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of the
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<collectingCounty id="33264E7DFFBDFF9AF52FF796FDF7B364" box="[360,601,2066,2093]" pageId="2" pageNumber="294">University of Rennes</collectingCounty>
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,
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<collectingCountry id="A2EF7661FFBDFF9AF623F797FD1EB364" box="[612,688,2067,2093]" name="France" pageId="2" pageNumber="294">France</collectingCountry>
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(IGR).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9AF718FA29FA99BD78" pageId="2" pageNumber="294" type="etymology">
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<paragraph id="DA4736F1FFBDFF9AF718FA29FA99BD78" blockId="2.[810,1452,1453,2093]" pageId="2" pageNumber="294">
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<emphasis id="E88CEAE3FFBDFF9AF718FA29FC49BE8E" bold="true" box="[863,999,1453,1479]" pageId="2" pageNumber="294">Etymology.</emphasis>
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This species is dedicated to the French musical band Stupeflip, exploring various genres such as hip-hop, punk, electronic music and pop. The specific epithet is to be treated as a name in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9AF718F9BEFBD9BD1C" box="[863,1143,1594,1621]" pageId="2" pageNumber="294" type="diagnosis">
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<paragraph id="DA4736F1FFBDFF9AF718F9BEFBD9BD1C" blockId="2.[810,1452,1453,2093]" box="[863,1143,1594,1621]" pageId="2" pageNumber="294">
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<emphasis id="E88CEAE3FFBDFF9AF718F9BEFC77BD1D" bold="true" box="[863,985,1594,1620]" pageId="2" pageNumber="294">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9AF718F9DBFA92BD8B" pageId="2" pageNumber="294" type="materials_examined">
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||
<paragraph id="DA4736F1FFBDFF9AF718F9DBFA92BD8B" blockId="2.[810,1452,1453,2093]" pageId="2" pageNumber="294">
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||
<materialsCitation id="6A903CACFFBDFF9AF718F9DBFA96BD8B" country="Myanmar" county="Ma" location="Noije Bum" municipality="Albian-early Cenomanian" pageId="2" pageNumber="294" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
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<emphasis id="E88CEAE3FFBDFF9AF718F9DBFB0DBD30" bold="true" box="[863,1187,1631,1657]" pageId="2" pageNumber="294">
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<typeStatus id="05438853FFBDFF9AF718F9DBFC32BD30" box="[863,924,1631,1657]" pageId="2" pageNumber="294">Type</typeStatus>
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locality and horizon.
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</emphasis>
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||
<location id="DF27602AFFBDFF9AF0EAF9DBFA81BD33" LSID="urn:lsid:plazi:treatment:525187E7FFBDFF9DF4CFF970FCA4B98D:DF27602AFFBDFF9AF0EAF9DBFA81BD33" box="[1197,1327,1631,1658]" country="Myanmar" county="Ma" municipality="Albian-early Cenomanian" name="Noije Bum" pageId="2" pageNumber="294" stateProvince="Kachin">Noije Bum</location>
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,
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<location id="DF27602AFFBDFF9AF179F9DBFCD9BDD4" LSID="urn:lsid:plazi:treatment:525187E7FFBDFF9DF4CFF970FCA4B98D:DF27602AFFBDFF9AF179F9DBFCD9BDD4" country="Myanmar" county="Ma" municipality="Albian-early Cenomanian" name="Hukawng Valley" pageId="2" pageNumber="294" stateProvince="Kachin">Hukawng Valley</location>
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,
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<collectingRegion id="183CF813FFBDFF9AF7CFF907FB8FBDD4" box="[904,1057,1667,1693]" country="Myanmar" name="Kachin" pageId="2" pageNumber="294">Kachin State</collectingRegion>
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, northern
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<collectingCountry id="A2EF7661FFBDFF9AF0D9F900FAA1BDD7" box="[1182,1295,1668,1694]" name="Myanmar" pageId="2" pageNumber="294">Myanmar</collectingCountry>
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; late
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<collectingMunicipality id="3A23AC8BFFBDFF9AF111F907FC57BD8B" pageId="2" pageNumber="294">Albian-early Cenomanian</collectingMunicipality>
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,
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<emphasis id="E88CEAE3FFBDFF9AF043F92CFBB0BD88" box="[1028,1054,1704,1729]" italics="true" pageId="2" pageNumber="294">ca</emphasis>
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. 99
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<collectingCounty id="33264E7DFFBDFF9AF017F92CFBD6BD8B" box="[1104,1144,1704,1730]" pageId="2" pageNumber="294">Ma</collectingCounty>
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, mid-Cretaceous
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="92E2657AFFBDFF9DF718F948FCA4B98D" lastPageId="5" lastPageNumber="297" pageId="2" pageNumber="294" type="description">
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<paragraph id="DA4736F1FFBDFF9AF718F948FC05BC42" blockId="2.[810,1452,1453,2093]" pageId="2" pageNumber="294">
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<emphasis id="E88CEAE3FFBDFF9AF718F948FC5FBDAF" bold="true" box="[863,1009,1740,1766]" pageId="2" pageNumber="294">Description.</emphasis>
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Body length
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<quantity id="1D009B14FFBDFF9AF0C2F948FB45BDAF" box="[1157,1259,1740,1766]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="294" unit="mm" value="0.76">0.76 mm</quantity>
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; integument dark castaneous.
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</paragraph>
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<paragraph id="DA4736F1FFBDFF9BF718F890FF78BC7B" blockId="2.[810,1452,1453,2093]" lastBlockId="3.[136,778,1564,2059]" lastPageId="3" lastPageNumber="295" pageId="2" pageNumber="294">
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Head hypognathous, globular (length
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||
<quantity id="1D009B14FFBDFF9AF172F890FA09BC66" box="[1333,1447,1812,1839]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="294" unit="mm" value="0.16">0.16 mm</quantity>
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, width above eyes
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||
<quantity id="1D009B14FFBDFF9AF048F8BDFB2EBC1A" box="[1039,1152,1849,1875]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="294" unit="mm" value="0.23">0.23 mm</quantity>
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||
); back of head coarsely punctured; compound eye bulging, occupying about half of head lateral surface, composed of numerous ommatidia (number over 60 for each eye); torulus inserted on frontal shelf, higher than eye; antenna filiform, longer than body; scape elongate, longer than F1 (
|
||
<quantity id="1D009B14FFBDFF9AF09CF86AFAE8B341" box="[1243,1350,2030,2056]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="294" unit="mm" value="0.1">0.10 mm</quantity>
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||
); pedicel globular, about twice as long as wide (length
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<quantity id="1D009B14FFBDFF9AF172F796FA31B365" box="[1333,1439,2066,2093]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="294" unit="mm" value="0.05">0.05 mm</quantity>
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); 12 flagellomeres cylindrical, elongate, F1 and F2 not modified, flagellomeres 1–11 progressively shortening (
|
||
<quantity id="1D009B14FFBCFF9BF4D7F9E0FF58BD36" box="[144,246,1636,1663]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="295" unit="mm" value="0.1">0.10 mm</quantity>
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||
> F1>
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||
<quantity id="1D009B14FFBCFF9BF516F9E0FE18BD36" box="[337,438,1636,1663]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="295" unit="mm" value="0.09">0.09 mm</quantity>
|
||
> F2–F4>
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||
<quantity id="1D009B14FFBCFF9BF67AF9E0FD0CBD36" box="[573,674,1636,1663]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="295" unit="mm" value="0.08">0.08 mm</quantity>
|
||
> F5–F6>
|
||
<quantity id="1D009B14FFBCFF9BF4E6F90CFEA9BDEA" box="[161,263,1672,1699]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="3" pageNumber="295" unit="mm" value="0.07">0.07 mm</quantity>
|
||
> F7–F10>
|
||
<quantity id="1D009B14FFBCFF9BF5E4F90CFDA7BDEA" box="[419,521,1672,1699]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="295" unit="mm" value="0.06">0.06 mm</quantity>
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||
> F11) and widening, flagellomere 12 acuminate, about as long as flagellomere 4; pubescence on flagellomeres denser toward apex of antenna, setae fine, not longer than a third of flagellomere length.
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</paragraph>
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<caption id="8E876679FFBCFF9BF4CFFA10FEECBE99" ID-DOI="http://doi.org/10.5281/zenodo.8209287" ID-Zenodo-Dep="8209287" httpUri="https://zenodo.org/record/8209287/files/figure.png" pageId="3" pageNumber="295" startId="3.[136,235,1428,1452]" targetBox="[312,1275,189,1404]" targetPageId="3" targetType="figure">
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<paragraph id="DA4736F1FFBCFF9BF4CFFA10FEECBE99" blockId="3.[136,1452,1428,1489]" pageId="3" pageNumber="295">
|
||
<emphasis id="E88CEAE3FFBCFF9BF4CFFA10FEACBEE5" bold="true" box="[136,258,1428,1452]" pageId="3" pageNumber="295">FIGURE 3.</emphasis>
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||
<taxonomicName id="1DF84D72FFBCFF9BF540FA11FE74BEE4" authority="BRAZIDEC, 2023" authorityName="BRAZIDEC" authorityYear="2023" box="[263,474,1429,1453]" class="Insecta" family="Spathiopterygidae" genus="Argemiones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="295" phylum="Arthropoda" rank="species" species="stupeflip" status="gen. et sp. nov.">
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||
<emphasis id="E88CEAE3FFBCFF9BF540FA11FE74BEE4" box="[263,474,1429,1453]" italics="true" pageId="3" pageNumber="295">Argemiones stupeflip</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E88CEAE3FFBCFF9BF598FA11FDD9BEE4" bold="true" box="[479,631,1429,1453]" pageId="3" pageNumber="295">
|
||
<taxonomicNameLabel id="F3BF5798FFBCFF9BF598FA11FDD9BEE4" box="[479,631,1429,1453]" pageId="3" pageNumber="295" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, specimen IGR.BU-076, line drawing of habitus in dorsal view (pst = parastigma). Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA4736F1FFBCFF9BF4FAF8B9FE80BCAF" blockId="3.[136,778,1564,2059]" pageId="3" pageNumber="295">
|
||
Mesosoma stout (length
|
||
<quantity id="1D009B14FFBCFF9BF5A8F8B8FDCEBC1F" box="[495,608,1852,1878]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="295" unit="mm" value="0.29">0.29 mm</quantity>
|
||
); mesoscutum large, convex, about half length of mesosoma, apparently smooth; notaulus present, convergent posteriorly; pronotum apparently without lateral striations, not visible in dorsal view.
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBCFF9BF4FAF875FCCFBC30" blockId="3.[136,778,1564,2059]" lastBlockId="3.[810,1452,1563,2057]" pageId="3" pageNumber="295">
|
||
Fore wing paddle-like, longer than body (length
|
||
<quantity id="1D009B14FFBCFF9BF76DF99FFC3CBD7C" box="[810,914,1563,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="3" pageNumber="295" unit="mm" value="1.13">1.13 mm</quantity>
|
||
); membrane slightly wrinkled medially, smooth apically, uniformly micropubescent; fringe of long setae (longest
|
||
<quantity id="1D009B14FFBCFF9BF790F9E7FBEDBD34" box="[983,1091,1635,1661]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="3" pageNumber="295" unit="mm" value="0.07">0.07 mm</quantity>
|
||
) present along apical margin; parastigma sclerotized and slightly thickened, extending approximately to half of wing; R, Rs, M, Cu and A veins spectral but visible apically, outlined by microtrichiae, not sclerotized basally; fork of Rs and M basad apex of parastigma; A longer than parastigma; B not sclerotized. Hind wing strongly reduced, only present as remnant of vein.
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBCFF9CF718F807FD5DBD80" blockId="3.[810,1452,1563,2057]" lastBlockId="4.[136,778,1636,1926]" lastPageId="4" lastPageNumber="296" pageId="3" pageNumber="295">Legs slender, long with respect to body, with sparse setae; tibia with inner row of setae; tibial spur formula 1-1-1, metatibial spur longest and slightly curved; probasitarsomere shorter than remainder of protarsus; meso- and metabasitarsomeres, respectively, distinctly longer than remainder of meso- and metatarsus; tarsal claws simple, angled apically; arolium well-developed.</paragraph>
|
||
<caption id="8E876679FFBBFF9CF4CFFA42FC88BD6F" ID-DOI="http://doi.org/10.5281/zenodo.8209289" ID-Zenodo-Dep="8209289" httpUri="https://zenodo.org/record/8209289/files/figure.png" pageId="4" pageNumber="296" startId="4.[136,235,1478,1502]" targetBox="[151,1436,170,1455]" targetPageId="4" targetType="figure">
|
||
<paragraph id="DA4736F1FFBBFF9CF4CFFA42FC88BD6F" blockId="4.[136,1452,1478,1575]" pageId="4" pageNumber="296">
|
||
<emphasis id="E88CEAE3FFBBFF9CF4CFFA42FEA8BE96" bold="true" box="[136,262,1478,1503]" pageId="4" pageNumber="296">FIGURE 4.</emphasis>
|
||
<taxonomicName id="1DF84D72FFBBFF9CF549FA43FE4BBE96" authority="BRAZIDEC, 2023" authorityName="BRAZIDEC" authorityYear="2023" box="[270,485,1479,1503]" class="Insecta" family="Spathiopterygidae" genus="Argemiones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="stupeflip" status="gen. et sp. nov.">
|
||
<emphasis id="E88CEAE3FFBBFF9CF549FA43FE4BBE96" box="[270,485,1479,1503]" italics="true" pageId="4" pageNumber="296">Argemiones stupeflip</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E88CEAE3FFBBFF9CF5A9FA43FD3FBE96" bold="true" box="[494,657,1479,1503]" pageId="4" pageNumber="296">
|
||
<taxonomicNameLabel id="F3BF5798FFBBFF9CF5A9FA43FD3FBE96" box="[494,657,1479,1503]" pageId="4" pageNumber="296" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, specimen IGR.BU-076,
|
||
<emphasis id="E88CEAE3FFBBFF9CF7DDFA42FC02BE97" bold="true" box="[922,940,1478,1502]" pageId="4" pageNumber="296">A</emphasis>
|
||
, Habitus in ventral view.
|
||
<emphasis id="E88CEAE3FFBBFF9CF0FCFA43FB62BE96" bold="true" box="[1211,1228,1479,1503]" pageId="4" pageNumber="296">B</emphasis>
|
||
, Right fore wing and metasoma in dorsal view (arrow = remnant of hind wing).
|
||
<emphasis id="E88CEAE3FFBBFF9CF697FA6EFD4CBD4B" bold="true" box="[720,738,1514,1538]" pageId="4" pageNumber="296">C</emphasis>
|
||
, Left antenna in ventral view.
|
||
<emphasis id="E88CEAE3FFBBFF9CF055FA6FFB8ABD4A" bold="true" box="[1042,1060,1515,1539]" pageId="4" pageNumber="296">D</emphasis>
|
||
, Left mesotarsomere 5 in ventral view. Scale bars:
|
||
<emphasis id="E88CEAE3FFBBFF9CF4BAF98AFEA1BD6F" bold="true" box="[253,271,1550,1574]" pageId="4" pageNumber="296">A</emphasis>
|
||
= 1 mm,
|
||
<emphasis id="E88CEAE3FFBBFF9CF536F98BFE2CBD6E" bold="true" box="[369,386,1551,1575]" pageId="4" pageNumber="296">B</emphasis>
|
||
= 0.5 mm,
|
||
<emphasis id="E88CEAE3FFBBFF9CF5B0F98AFDA7BD6F" bold="true" box="[503,521,1550,1574]" pageId="4" pageNumber="296">C</emphasis>
|
||
= 0.25 mm,
|
||
<emphasis id="E88CEAE3FFBBFF9CF6CCF98BFD33BD6E" bold="true" box="[651,669,1551,1575]" pageId="4" pageNumber="296">D</emphasis>
|
||
= 0.125 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DA4736F1FFBBFF9CF4FAF951FD4EBC5C" blockId="4.[136,778,1636,1926]" pageId="4" pageNumber="296">
|
||
Petiole visible, not obscured by gaster, slightly longer than wide (length
|
||
<quantity id="1D009B14FFBBFF9CF514F97FFE17BC5C" box="[339,441,1787,1813]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="296" unit="mm" value="0.06">0.06 mm</quantity>
|
||
, width
|
||
<quantity id="1D009B14FFBBFF9CF64CF97EFDDDBC5C" box="[523,627,1786,1813]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="296" unit="mm" value="0.05">0.05 mm</quantity>
|
||
), smooth.
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF4FAF8A4FF43BCCF" blockId="4.[136,778,1636,1926]" pageId="4" pageNumber="296">
|
||
Gaster globose (length
|
||
<quantity id="1D009B14FFBBFF9CF586F8A4FD87BC73" box="[449,553,1824,1851]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="296" unit="mm" value="0.25">0.25 mm</quantity>
|
||
); second metasomal tergite and sternite longest, hiding other tergites and sternites.
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF4CFF833FD69BC98" blockId="4.[136,711,1975,2001]" box="[136,711,1975,2001]" pageId="4" pageNumber="296">
|
||
Key to the genera and species of
|
||
<taxonomicName id="1DF84D72FFBBFF9CF5BCF833FD69BC98" authorityName="Engel & Ortega-Blanco" authorityYear="2013" box="[507,711,1975,2001]" class="Insecta" family="Spathiopterygidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="family">Spathiopterygidae</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF76DF9E1FA02BDFD" blockId="4.[810,1452,1637,2092]" pageId="4" pageNumber="296">
|
||
2. Compound eye bulging with few ommatidia (‘berry-like’), parastigma thin (thinner than other visible veins)..................................3 [
|
||
<taxonomicName id="1DF84D72FFBBFF9CF7BDF91BFA0BBDFD" authority="Engel & Ortega-Blanco, 2013" authorityName="Engel & Ortega-Blanco" authorityYear="2013" box="[1018,1445,1695,1717]" class="Insecta" family="Spathiopterygidae" genus="Spathiopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E88CEAE3FFBBFF9CF7BDF91BFBD1BDFD" box="[1018,1151,1695,1716]" italics="true" pageId="4" pageNumber="296">Spathiopteryx</emphasis>
|
||
<bibRefCitation id="BE694B00FFBBFF9CF0C2F91BFA0BBDFD" author="Engel, M. S. & Ortega-Blanco, J. & Soriano, C. & Grimaldi, D. A. & Delclos, X." box="[1157,1445,1695,1717]" pageId="4" pageNumber="296" pagination="1 - 23" refId="ref4226" refString="Engel, M. S., Ortega-Blanco, J., Soriano, C., Grimaldi, D. A. & Delclos, X. (2013) A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates, 3771, 1 - 23. https: // doi. org / 10.1206 / 3771.2" type="journal article" year="2013">Engel & Ortega-Blanco, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF76DF938FA05BC42" blockId="4.[810,1452,1637,2092]" pageId="4" pageNumber="296">- Compound eye with numerous ommatidia (more than 50 ommatidia), parastigma thick (at least as thick as other visible veins).......................................................................4</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF76DF896FA05BC28" blockId="4.[810,1452,1637,2092]" pageId="4" pageNumber="296">
|
||
3. Compound eye with about 40 ommatidia, pronotum with transverse striation, vertex punctured...........
|
||
<taxonomicName id="1DF84D72FFBBFF9CF158F8ABFA05BC28" authority="Engel & Ortega-Blanco, 2013" authorityName="Engel & Ortega-Blanco" authorityYear="2013" class="Insecta" family="Spathiopterygidae" genus="Spathiopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="alavarommopsis">
|
||
<emphasis id="E88CEAE3FFBBFF9CF158F8ABFBBDBC28" italics="true" pageId="4" pageNumber="296">Spathiopteryx alavarommopsis</emphasis>
|
||
<bibRefCitation id="BE694B00FFBBFF9CF071F8C8FA05BC28" author="Engel, M. S. & Ortega-Blanco, J. & Soriano, C. & Grimaldi, D. A. & Delclos, X." box="[1078,1451,1868,1890]" pageId="4" pageNumber="296" pagination="1 - 23" refId="ref4226" refString="Engel, M. S., Ortega-Blanco, J., Soriano, C., Grimaldi, D. A. & Delclos, X. (2013) A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates, 3771, 1 - 23. https: // doi. org / 10.1206 / 3771.2" type="journal article" year="2013">Engel & Ortega-Blanco, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF76DF8EDFA05BCF1" blockId="4.[810,1452,1637,2092]" pageId="4" pageNumber="296">
|
||
- Compound eye with about 20 ommatidia, pronotum smooth, vertex punctate to areolate..............................
|
||
<taxonomicName id="1DF84D72FFBBFF9CF733F826FA05BCF1" authority="Szabo, Brazidec & Perrichot, 2022" authorityName="Szabo, Brazidec & Perrichot" authorityYear="2022" box="[884,1451,1954,1976]" class="Insecta" family="Spathiopterygidae" genus="Spathiopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="soosi">
|
||
<emphasis id="E88CEAE3FFBBFF9CF733F826FB99BCF1" box="[884,1079,1954,1976]" italics="true" pageId="4" pageNumber="296">Spathiopteryx soosi</emphasis>
|
||
<bibRefCitation id="BE694B00FFBBFF9CF003F826FA05BCF1" author="Szabo, M. & Brazidec, M. & Perrichot, V. & Szenti, I. & Kukovecz, A. & Osi, A." box="[1092,1451,1954,1976]" pageId="4" pageNumber="296" pagination="105314" refId="ref5334" refString="Szabo, M., Brazidec, M., Perrichot, V., Szenti, I., Kukovecz, A. & Osi, A. (2022) A unique record of the Late Cretaceous of East-Central Europe: The first fossil wasps (Hymenoptera: Bethylidae, Spathiopterygidae) from the ajkaite amber (Bakony Mts., western Hungary). Cretaceous Research, 139, 105314. https: // doi. org / 10.1016 / j. cretres. 2022.105314" type="journal article" year="2022">Szabó, Brazidec & Perrichot, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBBFF9CF76DF83BFA05B347" blockId="4.[810,1452,1637,2092]" pageId="4" pageNumber="296">
|
||
4. Fore wing membrane entirely wrinkled, with a positive gradient of microtrichiae density towards apex.................. ..........
|
||
<taxonomicName id="1DF84D72FFBBFF9CF7F7F87EFA05B347" authority="Santer & Alvarez-Parra, 2022" authorityName="Santer & Alvarez-Parra" authorityYear="2022" box="[944,1451,2041,2065]" class="Insecta" family="Spathiopterygidae" genus="Diameneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="marveni">
|
||
<emphasis id="E88CEAE3FFBBFF9CF7F7F87EFBD1B346" box="[944,1151,2042,2063]" italics="true" pageId="4" pageNumber="296">Diameneura marveni</emphasis>
|
||
<bibRefCitation id="BE694B00FFBBFF9CF0C0F87DFA05B347" author="Santer, M. & Alvarez-Parra, S. & Nel, A. & Penalver, E. & Delclos, X." box="[1159,1451,2041,2065]" pageId="4" pageNumber="296" pagination="105128" refId="ref4907" refString="Santer, M., Alvarez-Parra, S., Nel, A., Penalver, E. & Delclos, X. (2022) New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea). Cretaceous Research, 133, 105128. https: // doi. org / 10.1016 / j. cretres. 2021.105128" type="journal article" year="2022">Santer & Álvarez-Parra, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<footnote id="B9E32AFFFFBBFF9CF4CFF87CFCA4B362" pageId="4" pageNumber="296">
|
||
<paragraph id="DA4736F1FFBBFF9CF4CFF87CFCA4B362" blockId="4.[136,778,2039,2092]" pageId="4" pageNumber="296">1. Fore wing with a well-defined marginal fringe of setae........2 - Fore wing without a marginal fringe of setae.....................7</paragraph>
|
||
</footnote>
|
||
<paragraph id="DA4736F1FFBBFF9DF76DF792FCA4BBFE" blockId="4.[810,1452,1637,2092]" lastBlockId="5.[136,778,161,708]" lastPageId="5" lastPageNumber="297" pageId="4" pageNumber="296">- Fore wing membrane partly smooth, microtrichiae distributed evenly on membrane....................................5</paragraph>
|
||
<paragraph id="DA4736F1FFBAFF9DF4CFFF3AFCA7BBB9" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
|
||
5. Fore wing with M+Cu desclerotized, hind wing reduced as a venal stub .............
|
||
<taxonomicName id="1DF84D72FFBAFF9DF5DCFF58FDC6BBB9" authority="BRAZIDEC, 2023" authorityName="BRAZIDEC" authorityYear="2023" box="[411,616,219,241]" class="Insecta" family="Spathiopterygidae" genus="Argemiones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="stupeflip" status="gen. et sp. nov.">
|
||
<emphasis id="E88CEAE3FFBAFF9DF5DCFF58FDC6BBB9" box="[411,616,219,241]" italics="true" pageId="5" pageNumber="297">Argemiones stupeflip</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E88CEAE3FFBAFF9DF628FF5FFCA7BBB9" bold="true" box="[623,777,219,240]" pageId="5" pageNumber="297">
|
||
<taxonomicNameLabel id="F3BF5798FFBAFF9DF628FF5FFCA7BBB9" box="[623,777,219,240]" pageId="5" pageNumber="297" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBAFF9DF4CFFF7DFCA4BA62" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
|
||
- Fore wing with M+Cu sclerotized, hind wing with membrane and hamuli 6 [
|
||
<taxonomicName id="1DF84D72FFBAFF9DF527FE92FCAABA62" authority="Engel & Ortega-Blanco, 2013" authorityName="Engel & Ortega-Blanco" authorityYear="2013" box="[352,772,278,300]" class="Insecta" family="Spathiopterygidae" genus="Mymaropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E88CEAE3FFBAFF9DF527FE92FE79BA62" box="[352,471,278,299]" italics="true" pageId="5" pageNumber="297">Mymaropsis</emphasis>
|
||
<bibRefCitation id="BE694B00FFBAFF9DF599FE92FCAABA62" author="Engel, M. S. & Ortega-Blanco, J. & Soriano, C. & Grimaldi, D. A. & Delclos, X." box="[478,772,278,300]" pageId="5" pageNumber="297" pagination="1 - 23" refId="ref4226" refString="Engel, M. S., Ortega-Blanco, J., Soriano, C., Grimaldi, D. A. & Delclos, X. (2013) A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates, 3771, 1 - 23. https: // doi. org / 10.1206 / 3771.2" type="journal article" year="2013">Engel & Ortega-Blanco, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
]
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBAFF9DF4CFFEB7FCA4BAE9" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
|
||
6. Scape slightly expanded ventrally, pedicel globular, flagellomeres 5–12 twice as wide as flagellomeres 1– 4 .......................................................................
|
||
<taxonomicName id="1DF84D72FFBAFF9DF6CBFEEAFCA4BAE9" authority="Krogmann, Azar, Rajaei & Nel, 2016" authorityName="Krogmann, Azar, Rajaei & Nel" authorityYear="2016" class="Insecta" family="Spathiopterygidae" genus="Mymaropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="baabdensis">
|
||
<emphasis id="E88CEAE3FFBAFF9DF6CBFEEAFE91BAD6" italics="true" pageId="5" pageNumber="297">Mymaropsis baabdensis</emphasis>
|
||
<bibRefCitation id="BE694B00FFBAFF9DF513FE0FFCA4BAE9" author="Krogmann, L. & Azar, D. & Rajaei, H. & Nel, A." box="[340,778,394,416]" pageId="5" pageNumber="297" pagination="483 - 487" refId="ref4422" refString="Krogmann, L., Azar, D., Rajaei, H. & Nel, A. (2016) Mymaropsis baabdaensis sp. n. from Lower Cretaceous Lebanese amber - the earliest spathiopterygid wasp and the first female known for the family. Comptes Rendus Palevol, 15, 483 - 487. https: // doi. org / 10.1016 / j. crpv. 2015.11.002" type="journal article" year="2016">Krogmann, Azar, Rajaei & Nel, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBAFF9DF4CFFE2CFCA4BABE" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
|
||
- Scape not ventrally expanded, pedicel almost globular, flagellomeres evenly wide.................................................. .......
|
||
<taxonomicName id="1DF84D72FFBAFF9DF546FE67FCA4BABE" authority="Engel & Ortega-Blanco, 2013" authorityName="Engel & Ortega-Blanco" authorityYear="2013" box="[257,778,482,504]" class="Insecta" family="Spathiopterygidae" genus="Mymaropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="turolensis">
|
||
<emphasis id="E88CEAE3FFBAFF9DF546FE67FE71BABE" box="[257,479,482,504]" italics="true" pageId="5" pageNumber="297">Mymaropsis turolensis</emphasis>
|
||
<bibRefCitation id="BE694B00FFBAFF9DF5A1FE66FCA4BABE" author="Engel, M. S. & Ortega-Blanco, J. & Soriano, C. & Grimaldi, D. A. & Delclos, X." box="[486,778,482,504]" pageId="5" pageNumber="297" pagination="1 - 23" refId="ref4226" refString="Engel, M. S., Ortega-Blanco, J., Soriano, C., Grimaldi, D. A. & Delclos, X. (2013) A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates, 3771, 1 - 23. https: // doi. org / 10.1206 / 3771.2" type="journal article" year="2013">Engel & Ortega-Blanco, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="DA4736F1FFBAFF9DF4CFFD84FCA4B925" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
|
||
7. Head coarsely punctured, antennae short, not longer than body length, gaster globose, nearly as long as mesosoma ........................................................
|
||
<emphasis id="E88CEAE3FFBAFF9DF6D6FDBEFEF8B925" italics="true" pageId="5" pageNumber="297">Spathiopria sayrevillensis</emphasis>
|
||
<bibRefCitation id="BE694B00FFBAFF9DF523FDD3FCA4B925" author="Engel, M. S. & Ortega-Blanco, J. & Soriano, C. & Grimaldi, D. A. & Delclos, X." box="[356,778,599,621]" pageId="5" pageNumber="297" pagination="1 - 23" refId="ref4226" refString="Engel, M. S., Ortega-Blanco, J., Soriano, C., Grimaldi, D. A. & Delclos, X. (2013) A new lineage of enigmatic diaprioid wasps in Cretaceous amber (Hymenoptera: Diaprioidea). American Museum Novitates, 3771, 1 - 23. https: // doi. org / 10.1206 / 3771.2" type="journal article" year="2013">Engel, Ortega-Blanco & Grimaldi, 2013</bibRefCitation>
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</paragraph>
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<paragraph id="DA4736F1FFBAFF9DF4CFFDF1FCA4B98D" blockId="5.[136,778,161,708]" pageId="5" pageNumber="297">
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- Head imbricate, antennae elongate, longer than body length, gaster ovoid, longer than mesosoma........................... .........................
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<taxonomicName id="1DF84D72FFBAFF9DF52EFD2AFCA4B98D" authority="Engel & Huang, 2015" authorityName="Engel & Huang" authorityYear="2015" box="[361,778,686,708]" class="Insecta" family="Spathiopterygidae" genus="Diaspathion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="ortegai">
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<emphasis id="E88CEAE3FFBAFF9DF52EFD2AFD82B98D" box="[361,556,686,708]" italics="true" pageId="5" pageNumber="297">Diaspathion ortegai</emphasis>
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<bibRefCitation id="BE694B00FFBAFF9DF674FD2AFCA4B98D" author="Engel, M. S. & Huang, D. Y. & Azar, D. & Nel, A. & Davis, S. R. & Alvarado, M. & Breitkreuz, L. C. V." box="[563,778,686,708]" pageId="5" pageNumber="297" pagination="95 - 100" refId="ref4138" refString="Engel, M. S., Huang, D. Y., Azar, D., Nel, A., Davis, S. R., Alvarado, M. & Breitkreuz, L. C. V. (2015) The wasp family Spathiopterygidae in mid-Cretaceous amber from Myanmar (Hymenoptera: Diaprioidea). Comptes Rendus Palevol, 14, 95 - 100. http: // dx. doi. org / 10.1016 / j. crpv. 2014.11.002" type="journal article" year="2015">Engel & Huang, 2015</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |