223 lines
19 KiB
XML
223 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.690.13751" ID-GBIF-Dataset="356a4c29-470b-4ca5-8503-76191b4fef6f" ID-PMC="PMC5672587" ID-Pensoft-Pub="1313-2970-690-1" ID-PubMed="29118595" ID-ZBK="C1B8D7C059E54C3A944F69F4FDE96B20" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-690-1" ModsDocOrigin="ZooKeys 690" ModsDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" checkinTime="1502920743207" checkinUser="pensoft" docAuthor="Shpeley, Danny, Hunting, Wesley & Ball, George E." docDate="2017" docId="FA83870A19F5DE70263243279F3CF7AC" docLanguage="en" docName="ZooKeys 690: 1-195" docOrigin="ZooKeys 690" docSource="http://dx.doi.org/10.3897/zookeys.690.13751" docTitle="Selenophorus alternans Dejean" docType="treatment" docVersion="5" lastPageNumber="73" masterDocId="FE026465FF86843AE44A7B6EFF8DC422" masterDocTitle="A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography" masterLastPageNumber="195" masterPageNumber="1" pageNumber="73" updateTime="1668164690231" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Shpeley, Danny</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Hunting, Wesley</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Ball, George E.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2017</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>690</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>195</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.690.13751</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.690.13751</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-690-1</mods:identifier>
|
||
<mods:identifier type="ZBK">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
|
||
<mods:identifier type="ZooBank">C1B8D7C059E54C3A944F69F4FDE96B20</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="132785158" LSID="urn:lsid:plazi:treatment:FA83870A19F5DE70263243279F3CF7AC" httpUri="http://treatment.plazi.org/id/FA83870A19F5DE70263243279F3CF7AC" lastPageNumber="73" pageId="72" pageNumber="73">
|
||
<subSubSection pageId="72" pageNumber="73" type="nomenclature">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
<taxonomicName authority="Dejean" class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus alternans" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alternans">Selenophorus alternans Dejean</taxonomicName>
|
||
Figs 49A, 50
|
||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||
, 53
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="reference_group">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus alternans" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alternans">Selenophorus alternans</taxonomicName>
|
||
Dejean, 1829: 86. In
|
||
<normalizedToken originalValue="Chaudoir-Oberthür">Chaudoir-Oberthuer</normalizedToken>
|
||
Collection, 33 specimens in front of following box label:
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori alternans" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alternans">alternans</taxonomicName>
|
||
/ Dej./ Bresil/ C. Dejean// LECTOTYPE (here selected), labelled: [male]//
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori alternans" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alternans">alternans</taxonomicName>
|
||
m/ in Brasilia [green paper, handwritten]// [MNHP].-
|
||
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
|
||
: 265.-
|
||
<bibRefCitation author="Gundlach, DJ" journalOrPublisher="Anales de la Sociedad Espanola de Historia Natural" pageId="104" pageNumber="105" pagination="259 - 344" title="Apuntes para la fauna Puerto-Riquena. Octava Parte. Neuropteros, Hemipteros, Coleopteros." volume="22" year="1894">Gundlach 1894</bibRefCitation>
|
||
: 294.-
|
||
<bibRefCitation pageId="72" pageNumber="73">Csiki 1932</bibRefCitation>
|
||
: 1196.-
|
||
<bibRefCitation author="Darlington, PJ Jr" journalOrPublisher="Psyche" pageId="103" pageNumber="104" pagination="66 - 131" title="New West Indian Carabidae with a list of the Cuban species." url="https://doi.org/10.1155/1934/10161" volume="41" year="1934">Darlington 1934</bibRefCitation>
|
||
: 104.-
|
||
<bibRefCitation pageId="72" pageNumber="73">Blackwelder 1944</bibRefCitation>
|
||
: 49.-
|
||
<bibRefCitation author="Erwin, TL" journalOrPublisher="Quaestiones Entomologicae" pageId="104" pageNumber="105" pagination="350 - 466" title="Carabid beetles of the West Indies (Insects: Coleoptera): a synopsis of the genera and checklists of tribes of Caraboidea, and of the West Indian species." volume="20" year="1984">Erwin and Sims 1984</bibRefCitation>
|
||
: 440.-
|
||
<bibRefCitation author="Bennett, FD" journalOrPublisher="Caribbean Agricultural Research and Development Institute, Wildey" pageId="102" pageNumber="103" title="An annotated check-list of the insects and allied terrestrial arthropods of Barbados." year="1985">Bennett and Alam 1985</bibRefCitation>
|
||
: 20.-
|
||
<bibRefCitation pageId="72" pageNumber="73">Ball 1992</bibRefCitation>
|
||
: 85.-
|
||
<bibRefCitation author="Lorenz, W" journalOrPublisher="Lorenz, Tutzing" pageId="105" pageNumber="106" title="Systematic list of ground beetles of the World (InsectaColeoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). W." year="1998">Lorenz 1998</bibRefCitation>
|
||
: 355;
|
||
<bibRefCitation pageId="72" pageNumber="73">Lorenz 2005</bibRefCitation>
|
||
: 376.-
|
||
<bibRefCitation author="Peck, SB" journalOrPublisher="Florida Department of Agriculture and Consumer Services, Gainesville" pageId="106" pageNumber="107" title="A checklist of the beetles of Cuba, with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and neighboring land areas, Volume 18." year="2005">Peck 2005</bibRefCitation>
|
||
: 32.-
|
||
<bibRefCitation pageId="72" pageNumber="73">Ivie et al. 2008</bibRefCitation>
|
||
: 238.-
|
||
<bibRefCitation author="Perez-Gelabert, DE" journalOrPublisher="Zootaxa" pageId="106" pageNumber="107" pagination="1 - 530" title="Arthropods of Hispaniola (Dominican Republic and Haiti): A checklist and bibliography." volume="1831" year="2008">Perez-Gelabert 2008</bibRefCitation>
|
||
: 79.-
|
||
<bibRefCitation author="Turnbow, RH" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 66" title="An annotated checklist of the Coleoptera (Insecta) of the Bahamas." volume="0034" year="2008">Turnbow and Thomas 2008</bibRefCitation>
|
||
: 14.-
|
||
<bibRefCitation author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="106" pageNumber="107" pagination="1 - 51" title="The Beetles of Barbados, West Indies (Insecta: Coleoptera): diversity, distribution and faunal structure." volume="0073" year="2009">Peck 2009</bibRefCitation>
|
||
: 13.
|
||
</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus lineatopunctatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="lineatopunctatus">Selenophorus lineatopunctatus</taxonomicName>
|
||
Dejean, 1829: 86. TYPE MATERIAL: male, in front of the alternans box label (see above); LECTOTYPE (here selected), labelled: [male]//
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophori" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Selenophori lineatopunctatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="lineatopunctatus">lineatopunctatus</taxonomicName>
|
||
m./ Cayenne [green paper, handwritten]// [MNHP].-
|
||
<bibRefCitation author="Gemminger, M" journalOrPublisher="Gummi, Monachii" pageId="104" pageNumber="105" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tom. I. Cicindelidae- Carabidae. E. H." year="1868">Gemminger and Harold 1868</bibRefCitation>
|
||
: 266.-
|
||
<bibRefCitation author="Putzeys, JADH" journalOrPublisher="Stettiner Entomologische Zeitung" pageId="106" pageNumber="107" pagination="3 - 73" title="Description des Selenophorus de l'Amerique." volume="39" year="1878 a">Putzeys 1878a</bibRefCitation>
|
||
: 13.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="type locality">
|
||
<paragraph pageId="72" pageNumber="73">Type locality.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">Vicinity of Rio de Janeiro, State of Rio de Janeiro, Brazil.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="diagnosis">
|
||
<paragraph pageId="72" pageNumber="73">Diagnosis.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">
|
||
This species is readily separated from the similarly colored member of the palliatus species group,
|
||
<taxonomicName lsidName="S. woodruffi" pageId="72" pageNumber="73" rank="species" species="woodruffi">S. woodruffi</taxonomicName>
|
||
, by the impunctate intervals next to the basal ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="descriptive notes">
|
||
<paragraph pageId="72" pageNumber="73">Descriptive notes.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">Data for SBL in Table 1. Habitus as in Fig. 49A. Clypeus and labrum with anterior margin of each shallowly concave. Head, pronotum and elytra with mesh pattern isodiametric. Mouthparts and legs testaceous to slightly darker; antennae darker than legs. Dorsal and ventral surface rufo-brunneous to dark brunneous; dorsal surface with faint aeneous metallic luster. Elytron bicolored, with apical fascia testaceous to slightly darker, length of pale marking nearly the same in intervals 2-9, forming a diagonal pale fascia; pale marking of interval 1longer than that of interval 2; intervals 3-5 may be darker just in front of pale apical fascia. Elytral epipleuron pale, same color as the legs. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Punctures of striae 2, 5 and 7 markedly foveate. Elytron with intervals impunctate basally near basal ridge. Both males and females with four terminal setae near the posterior margin on sternum VII.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Male genitalia. Fig. 50
|
||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||
. Very similar to that of
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
. For details, see this topic for
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
, below.
|
||
</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Ovipositor and female reproductive tract. Very similar to that of
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
, Fig. 52A. For details, see this topic for
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
, below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="distribution">
|
||
<paragraph pageId="72" pageNumber="73">Geographical distribution.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">Fig. 53. The known range of this species extends through the Lesser Antilles north-westward to the Virgin Islands, Puerto Rico, Hispaniola and to the islands of Andros, Mayaguana and New Providence in the Bahamas.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="chorological affinities and relationships">
|
||
<paragraph pageId="72" pageNumber="73">Chorological affinities and relationships.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">
|
||
The West Indian range of this species is overlapped by the ranges of
|
||
<taxonomicName lsidName="S. palliatus" pageId="72" pageNumber="73" rank="species" species="palliatus">S. palliatus</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="S. woodruffi" pageId="72" pageNumber="73" rank="species" species="woodruffi">S. woodruffi</taxonomicName>
|
||
, all members of the palliatus species group. Relationships of
|
||
<taxonomicName lsidName="S. alternans" pageId="72" pageNumber="73" rank="species" species="alternans">S. alternans</taxonomicName>
|
||
are not postulated beyond species group membership.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="73" type="materials_examined">
|
||
<paragraph pageId="72" pageNumber="73">Material examined.</paragraph>
|
||
<paragraph pageId="72" pageNumber="73">In addition to type material, we have seen a total of 678 specimens (310 males, 365 females, 3 unknown). See Appendix for details.</paragraph>
|
||
<caption pageId="72" pageNumber="73">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Figure 49. Habitus digital images of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus palliatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="palliatus">Selenophorus palliatus</taxonomicName>
|
||
species group, dorsal aspect. A
|
||
<taxonomicName lsidName="S. alternans" pageId="72" pageNumber="73" rank="species" species="alternans">S. alternans</taxonomicName>
|
||
Dejean B
|
||
<taxonomicName lsidName="S. palliatus" pageId="72" pageNumber="73" rank="species" species="palliatus">S. palliatus</taxonomicName>
|
||
(Fabricius) C
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
Dejean D
|
||
<taxonomicName lsidName="S. woodruffi" pageId="72" pageNumber="73" rank="species" species="woodruffi">S. woodruffi</taxonomicName>
|
||
, Ball & Shpeley. Scale bars: A, B, D 5 mm; C 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="72" pageNumber="73">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Figure 50. Digital images of male genitalia of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus palliatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="palliatus">Selenophorus palliatus</taxonomicName>
|
||
species group, in part. A, D, G right lateral aspect B, E, H dorsal aspect C, F, I left lateral aspect.
|
||
<normalizedToken originalValue="A–C">A-C</normalizedToken>
|
||
<taxonomicName lsidName="S. alternans" pageId="72" pageNumber="73" rank="species" species="alternans">S. alternans</taxonomicName>
|
||
Dejean
|
||
<normalizedToken originalValue="D–F">D-F</normalizedToken>
|
||
<taxonomicName lsidName="S. palliatus" pageId="72" pageNumber="73" rank="species" species="palliatus">S. palliatus</taxonomicName>
|
||
(Fabricius)
|
||
<normalizedToken originalValue="G–I">G-I</normalizedToken>
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
Dejean. Scale bars 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="72" pageNumber="73">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Figure 51. Digital images of male genitalia of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus palliatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="palliatus">Selenophorus palliatus</taxonomicName>
|
||
species group, in part,
|
||
<taxonomicName lsidName="S. woodruffi" pageId="72" pageNumber="73" rank="species" species="woodruffi">S. woodruffi</taxonomicName>
|
||
Ball & Shpeley. A right lateral aspect B dorsal aspect C left lateral aspect. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="72" pageNumber="73">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Figure 52. Line drawing of female reproductive tract of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus palliatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="palliatus">Selenophorus palliatus</taxonomicName>
|
||
species group, in part, ventral aspect. A
|
||
<taxonomicName lsidName="S. pyritosus" pageId="72" pageNumber="73" rank="species" species="pyritosus">S. pyritosus</taxonomicName>
|
||
Dejean B
|
||
<taxonomicName lsidName="S. woodruffi" pageId="72" pageNumber="73" rank="species" species="woodruffi">S. woodruffi</taxonomicName>
|
||
Ball & Shpeley. Legend: bc bursa copulatrix; co common oviduct gc1 gonocoxite 1 gc2 gonocoxite 2 sp spermatheca spg spermathecal gland spgd spermathecal gland duct v valvifer. Scale bars 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="72" pageNumber="73">
|
||
<paragraph pageId="72" pageNumber="73">
|
||
Figure 53. Map of West Indies showing known localities for species of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Selenophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Selenophorus palliatus" order="Coleoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="palliatus">Selenophorus palliatus</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |