treatments-xml/data/AA/97/EF/AA97EF8662D6583C935854789638136D.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.205.82300" ID-Pensoft-Pub="1314-2003-205-59" ID-Pensoft-UUID="80FCD4712A5A55208D85A8328CD55CDD" ModsDocID="1314-2003-205-59" checkinTime="1661230231685" checkinUser="pensoft" docAuthor="Clark, Ruth P., Jiang, Kai-Wen &amp; Gagnon, Edeline" docDate="2022" docId="AA97EF8662D6583C935854789638136D" docLanguage="en" docName="PhytoKeys 205: 59-98" docOrigin="PhytoKeys 205" docPubDate="2022-08-22" docSource="http://dx.doi.org/10.3897/phytokeys.205.82300" docTitle="Ticanto crista (L.) R. Clark &amp; Gagnon 2022, comb. nov." docType="treatment" docVersion="1" id="80FCD4712A5A55208D85A8328CD55CDD" lastPageNumber="59" masterDocId="80FCD4712A5A55208D85A8328CD55CDD" masterDocTitle="Reinstatement of Ticanto (Leguminosae - Caesalpinioideae) - the final piece in the Caesalpinia group puzzle" masterLastPageNumber="98" masterPageNumber="59" pageNumber="59" updateTime="1661230533549" updateUser="ExternalLinkService">
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<mods:title>Reinstatement of Ticanto (Leguminosae - Caesalpinioideae) - the final piece in the Caesalpinia group puzzle</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Clark, Ruth P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9974-2933</mods:nameIdentifier>
<mods:affiliation>Accelerated Taxonomy Department, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
<mods:nameIdentifier type="email">r.clark@kew.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Kai-Wen</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5917-1846</mods:nameIdentifier>
<mods:affiliation>Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China &amp; Ningbo Botanical Garden, Ningbo 315201, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gagnon, Edeline</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3212-9688</mods:nameIdentifier>
<mods:affiliation>Royal Botanic Garden Edinburgh, 20 a Inverleith Row, Edinburgh EH 3 5 LR, UK &amp; Technical University of Munich, Chair of Phytopathology, TUM School of Life Sciences, Emil-Ramman-St. 2, D- 85354, Freising, Germany</mods:affiliation>
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<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>205</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>59</mods:start>
<mods:end>98</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.205.82300</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.205.82300</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-205-59</mods:identifier>
<mods:identifier type="Pensoft-UUID">80FCD4712A5A55208D85A8328CD55CDD</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:AA97EF8662D6583C935854789638136D" httpUri="http://treatment.plazi.org/id/AA97EF8662D6583C935854789638136D" lastPageNumber="59" pageId="0" pageNumber="59">
<subSubSection pageId="0" pageNumber="59" type="nomenclature">
<paragraph pageId="0" pageNumber="59">
2. 
<taxonomicName LSID="AA97EF86-62D6-583C-9358-54789638136D" authority="(L.) R. Clark &amp; Gagnon" authorityName="(L.) R. Clark &amp; Gagnon" authorityYear="2022" baseAuthorityName="L." class="Magnoliopsida" family="Fabaceae" genus="Ticanto" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ticanto crista" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="crista" status="comb. nov.">Ticanto crista (L.) R. Clark &amp; Gagnon</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="59">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="reference_group">
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina nuga" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="nuga" status="comb. nov.">= Guilandina nuga</taxonomicName>
L., Sp. Pl., 2. 1: 545. 1762. Type: [Indonesia]. East Indies, Ambon, 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" family="Fabaceae" genus="Nugae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Nugae silvarum" order="Fabales" pageId="0" pageNumber="59" rank="species" species="silvarum" status="comb. nov.">Nugae silvarum</taxonomicName>
Rumph. Herb. Am. 5. p.95, t. 50. 1750.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina axillaris" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="axillaris" status="comb. nov.">= Guilandina axillaris</taxonomicName>
Lam., Encycl. 1(2): 435. 1785. Type: [India]. Rheede. Hort. Mal. 6: t. 20. 1686.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authority="Adans." authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Ticanto" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Ticanto nuga" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="nuga" status="comb. nov.">= Ticanto nuga</taxonomicName>
(L.) Medik., Theodora 52. 1786. Type: based on 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina nuga" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="nuga" status="comb. nov.">Guilandina nuga</taxonomicName>
L.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina paniculata" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="paniculata" status="comb. nov.">= Guilandina paniculata</taxonomicName>
Lam., Encycl. 1(2): 435. 1785. Type: [India]. Malabar, Kaka Mullu vel Kaka Moullou (in caption Kaka Mullu) Rheede, Hort. Mal. 6: t. 19. 1686.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Genista" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Genista scandens" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="scandens" status="comb. nov.">= Genista scandens</taxonomicName>
Lour., Fl. Cochinch. 2: 428. 1790. Type: Cochinchina (
<emphasis italics="true" pageId="0" pageNumber="59">n.v.</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina parvifolia" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="parvifolia" status="comb. nov.">= Guilandina parvifolia</taxonomicName>
Stokes, Bot. Mat. Med. 2: 466. 1812. Type: [Indonesia]. East Indies, Ambon, 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" family="Fabaceae" genus="Nugae" higherTaxonomySource="treatment-meta" kingdom="Plantae" lsidName="Nugae silvarum" order="Fabales" pageId="0" pageNumber="59" rank="species" species="silvarum" status="comb. nov.">Nugae silvarum</taxonomicName>
Rumph. Herb. Am. 5. p.95, t.50. 1750.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia nuga" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="nuga" status="comb. nov.">= Caesalpinia nuga</taxonomicName>
(L.) W.T. Aiton, Hort. Kew, ed 2, 3: 32. 1811. Type: based on 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina nuga" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="nuga" status="comb. nov.">Guilandina nuga</taxonomicName>
L.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia paniculata" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="paniculata" status="comb. nov.">= Caesalpinia paniculata</taxonomicName>
(Lam.) Roxb., Hort. Beng. 32. 1814. Type: based on 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina paniculata" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="paniculata" status="comb. nov.">Guilandina paniculata</taxonomicName>
Lam.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia scandens" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="scandens" status="comb. nov.">= Caesalpinia scandens</taxonomicName>
Heyne ex Roth, Nov. Pl. Sp. 209. 1821. Neotype: (designated by 
<bibRefCitation DOI="https://doi.org/10.1002/tax.12449" author="Turner, IM" journalOrPublisher="Taxon" pageId="0" pageNumber="59" pagination="365 - 428" refId="B63" refString="Turner, IM, 2021. Heyne, Roth, Roemer and Schultes, and the plant names published in Novae plantarum species praesertim Indiae orientalis. Taxon 70 (2): 365 - 428, DOI: https://doi.org/10.1002/tax.12449" title="Heyne, Roth, Roemer and Schultes, and the plant names published in Novae plantarum species praesertim Indiae orientalis." url="https://doi.org/10.1002/tax.12449" volume="70" year="2021">Turner 2021</bibRefCitation>
): India, Samalcott, Mar 1811, 
<emphasis italics="true" pageId="0" pageNumber="59">B. Heyne s.n.</emphasis>
[K: K000789355!].
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia axillaris" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="axillaris" status="comb. nov.">= Caesalpinia axillaris</taxonomicName>
(Lam.) DC., Prodr. 2: 481. 1825. Type: based on 
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" baseAuthorityName="Clark &amp; Jiang &amp; Gagnon" baseAuthorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina axillaris" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="axillaris" status="comb. nov.">Guilandina axillaris</taxonomicName>
Lam.
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia laevigata" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="laevigata" status="comb. nov.">= Caesalpinia laevigata</taxonomicName>
Perr., 
<normalizedToken originalValue="Mém">Mem</normalizedToken>
. Soc. Linn. Paris 3: 104. 1825. Type: Philippines. 
<emphasis italics="true" pageId="0" pageNumber="59">Perrottet s.n.</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="59">n.v.</emphasis>
).
</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName authorityName="Clark &amp; Jiang &amp; Gagnon" authorityYear="2022" class="Magnoliopsida" family="Fabaceae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia crista subsp. var. var. parvistipula" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="variety" species="crista" status="comb. nov." subSpecies="var." variety="parvistipula">= Caesalpinia crista var. parvistipula</taxonomicName>
Urb., Symb. Antill. 2(2): 271. 1900. Type: Trinidad. Cult. Hort. Trinidad, 
<emphasis italics="true" pageId="0" pageNumber="59">Broadway 5589</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="59">n.v.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="basionym">
<paragraph pageId="0" pageNumber="59">Basionym.</paragraph>
<paragraph pageId="0" pageNumber="59">
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia crista" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="crista">
<emphasis italics="true" pageId="0" pageNumber="59">Caesalpinia crista</emphasis>
</taxonomicName>
L. Sp. Pl. 1: 380. 1753, emend Dandy &amp; Exell in J. Bot. 76: 179. 1938. ≡ 
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Guilandina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Guilandina crista" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="crista">
<emphasis italics="true" pageId="0" pageNumber="59">Guilandina crista</emphasis>
</taxonomicName>
(L.) Small, Fl. S.E. U.S. 591, 1331 (1903).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="materials_examined">
<paragraph pageId="0" pageNumber="59">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="59">
[
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Sri Lanka" location="Skeels in Science" specimenCount="1" typeStatus="lectotype">
<collectingCountry name="Sri Lanka">Sri Lanka</collectingCountry>
] Ceylon (
<normalizedToken originalValue="“Ceylan”">
&quot; 
<collectingCountry name="Sri Lanka">Ceylan</collectingCountry>
&quot;
</normalizedToken>
), 
<emphasis italics="true" pageId="0" pageNumber="59">Herb. Hermann</emphasis>
vol. 1, fol. 68, no. 157 (
<typeStatus>lectotype</typeStatus>
(designated by 
<location LSID="urn:lsid:plazi:treatment:AA97EF8662D6583C935854789638136D:55AB4E6A0B38F06242799D90FDB03E58" country="Sri Lanka" name="Skeels in Science">Skeels in Science</location>
, n.s., 37: 922. 1913): BM [BM000621459!]) (note: the sheet bearing this specimen was previously identified by a single barcode, BM000594500, which was subsequently replaced with four barcodes representing the four separate specimens on the sheet)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="description">
<paragraph pageId="0" pageNumber="59">Description.</paragraph>
<paragraph pageId="0" pageNumber="59">
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Habit</emphasis>
a liana or scrambling shrub to 15 m. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Stems</emphasis>
with few, scattered recurved prickles to 5 mm, sometimes with spine-tipped corky tubercles on older stems, or unarmed, glabrous or occasionally sparsely tomentose. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Stipules</emphasis>
persistent, triangular, ca. 1 
<normalizedToken originalValue="×">x</normalizedToken>
1 mm. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Leaves</emphasis>
with 3-6(-8) pairs pinnae, these opposite to occasionally slightly subopposite; petiole 1.8-5 cm; rachis 4-31 cm, usually armed with recurved prickles, these sometimes also on pinnae rachises, petiole and rachis usually glabrous, occasionally sparsely to moderately ferruginous tomentose; leaflets 2-4(-7) pairs per pinna, coriaceous, opposite, elliptic, base cuneate to obtuse, apex rounded to obtuse, less commonly acute, obtuse, or acuminate, terminal leaflets 2.1-7.2 
<normalizedToken originalValue="×">x</normalizedToken>
1-3.3 cm, lateral leaflets 2.1-5.8 
<normalizedToken originalValue="×">x</normalizedToken>
1-3.1 cm, upper surface glabrous, often glossy, lower surface glabrous or occasionally sparsely ferruginous tomentose, venation reticulate, anastomosing, visible on both surfaces. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Inflorescence</emphasis>
a raceme or panicle, axillary or terminal, 8-40 cm, axes glabrous or sparsely tomentose; pedicels 5-15 mm; bracts persistent, triangular or lunate, 1-1.5 
<normalizedToken originalValue="×">x</normalizedToken>
1 mm; bracteoles caducous, broadly elliptic, apex acute, margins sometimes with small teeth, 1-2.5 
<normalizedToken originalValue="×">x</normalizedToken>
0.5-1 mm. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Flowers</emphasis>
with a hypanthium 1-2 
<normalizedToken originalValue="×">x</normalizedToken>
3-6 mm, glabrous or sparsely ferruginous tomentose; lower calyx lobe 6-8(-10) 
<normalizedToken originalValue="×">x</normalizedToken>
3-4 mm, other lobes 5-6(- 8) 
<normalizedToken originalValue="×">x</normalizedToken>
2-3 mm, all lobes glabrous, or margins ciliate, or sparsely ferruginous tomentose; median petal 6-9 
<normalizedToken originalValue="×">x</normalizedToken>
3-7 mm, with dense circular patch of hairs on inner surface at base of blade; upper laterals 6-10 
<normalizedToken originalValue="×">x</normalizedToken>
3-5 mm, including claw ca. 1 mm, glabrous or inner surface of claw sparsely hairy; lower laterals 7-11 
<normalizedToken originalValue="×">x</normalizedToken>
3-6 mm, including claw ca. 1 mm, glabrous or inner surface of claw sparsely hairy; stamen filaments 4-12 mm, the vexillary shorter than the lower ones, orange tomentose on lower 
<normalizedToken originalValue="½-¾">1/2-3/4</normalizedToken>
on inner surface; anthers 1-1.5 mm; ovary 2-4 mm long, glabrous or sparsely or partially tomentose; style 4-11 mm; stigma cupular or funnel-shaped, the rim papillate, sometimes only slightly so, 0.5-1 mm wide. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Fruit</emphasis>
indehiscent, coriaceous, elliptic to lunate, subsymmetrical to somewhat asymmetrical, slightly inflated, base cuneate, stipe 2-5 mm, apex acute to beaked, beak 1-10 mm, venation reticulate, prominent, glabrous or very sparsely tomentose, (2-)2.7-7 
<normalizedToken originalValue="×">x</normalizedToken>
2.2-3.7 
<normalizedToken originalValue="×">x</normalizedToken>
0.3-0.8 cm, lacking a wing. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="59">Seeds</emphasis>
1, rarely 2, circular to reniform in outline, flat, ca. 2-2.5 
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2 
<normalizedToken originalValue="×">x</normalizedToken>
0.5-1 cm. (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F5" captionText="Figure 3. A T. sinensis (i) habit (ii) leaves (iii) flowers (photos by Xin-Xin Zhu, Guizhou, China, unvouchered) (iv) habit (v) fruit (R. P. Clark 429, Guangxi, China, (IBK, K )) (vi) leaves (R. P. Clark 415, Guangxi, China (IBK, K) B T. aff. szechuenensis (i) habit (ii) inflorescence and leaves (R. P. Clark 422, Guangxi, China (IBK, K ))." figureDoi="10.3897/phytokeys.205.82300.figure3" httpUri="https://binary.pensoft.net/fig/732137" pageId="0" pageNumber="59">3A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="ecology">
<paragraph pageId="0" pageNumber="59">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="59">Riverbanks, sandy beaches, in and behind sandy parts of mangroves, on chalk rocks and limestone, at low altitude, elevation rarely up to 350 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="phenology">
<paragraph pageId="0" pageNumber="59">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="59">The flowering and fruiting time of this species varies throughout its distribution and may be correlated with latitude or biome as indicated below; however, the periodicity listed below derives in some cases from few records and thus may be incomplete.</paragraph>
<paragraph pageId="0" pageNumber="59">China, Japan: Flowering February-April, fruiting April-October;</paragraph>
<paragraph pageId="0" pageNumber="59">Bangladesh, India, Myanmar: Flowering August-December, fruiting December-October;</paragraph>
<paragraph pageId="0" pageNumber="59">Thailand, Vietnam: Flowering January-June, fruiting January-June;</paragraph>
<paragraph pageId="0" pageNumber="59">Malaysia, Indonesia, Palau Islands: Flowering February-December, fruiting February-January;</paragraph>
<paragraph pageId="0" pageNumber="59">Philippines: Flowering December-March, fruiting May-December;</paragraph>
<paragraph pageId="0" pageNumber="59">New Guinea: Flowering January-November, fruiting February-November;</paragraph>
<paragraph pageId="0" pageNumber="59">New Hebrides, Solomon Islands: Flowering February-December, fruiting February-December;</paragraph>
<paragraph pageId="0" pageNumber="59">New Caledonia: Flowering May, fruiting unknown;</paragraph>
<paragraph pageId="0" pageNumber="59">Mauritius: Flowering unknown, fruiting February.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="distribution">
<paragraph pageId="0" pageNumber="59">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="59">
Andaman Islands, Australia, Cambodia, China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hubei, Hunan, Sichuan, Taiwan, Yunnan), India, Indonesia, Japan (Ryukyu Islands), Malaysia, Mauritius, Micronesia, Myanmar, New Caledonia, Papua New Guinea, Philippines, Solomon Islands, Sri Lanka, Thailand, Vanuatu, Vietnam (Map 
<figureCitation captionStart="Map 2" captionStartId="F3" captionText="Map 2. Distribution of T. crista." figureDoi="10.3897/phytokeys.205.82300.map2" httpUri="https://binary.pensoft.net/fig/732135" pageId="0" pageNumber="59">2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="notes">
<paragraph pageId="0" pageNumber="59">Notes.</paragraph>
<paragraph pageId="0" pageNumber="59">
The current authors have not seen any specimens or specimen records from Cambodia (other than a single record in GBIF which is not from a preserved specimen), but 
<bibRefCitation author="Vidal, JE" journalOrPublisher="Current Medical and Drug Research" pageId="0" pageNumber="59" refId="B65" refString="Vidal, JE, Hul Thol, S, 1976. Revision des Caesalpinia asiatiques. Bulletin du Museum National d'histoire naturelle Ser.3: 27-81." title="Revision des Caesalpinia asiatiques. Bulletin du Museum National d'histoire naturelle Ser. 3: 27 - 81." year="1976">Vidal and Hul Thol (1976)</bibRefCitation>
list Cambodia within the range for this species, citing two specimens from Kampot Province (
<emphasis italics="true" pageId="0" pageNumber="59">Geoffray 62</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">62 bis</emphasis>
) and that information is retained here. As these specimens could not be traced by the current authors, they are not included on the map. See also Nomenclatural notes on 
<taxonomicName class="Magnoliopsida" family="Leguminosae" genus="Caesalpinia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caesalpinia (Ticanto) crista" order="Fabales" pageId="0" pageNumber="59" phylum="Tracheophyta" rank="species" species="crista" subGenus="Ticanto">Caesalpinia (Ticanto) crista</taxonomicName>
, below.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="59" type="materials_examined">
<paragraph pageId="0" pageNumber="59">Selected specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="59">
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Australia">Australia</collectingCountry>
. 
<collectingRegion country="Australia" name="Queensland">Queensland</collectingRegion>
</emphasis>
: Daintree area, Noah Creek, Mangroves, 
<emphasis italics="true" pageId="0" pageNumber="59">J.J. Wieringa 4199</emphasis>
(WAG). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Bangladesh">Bangladesh</collectingCountry>
.
</emphasis>
<collectingRegion country="Bangladesh" name="Chittagong">Chittagong</collectingRegion>
, 
<emphasis italics="true" pageId="0" pageNumber="59">Flagstaff 255</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="China">China</collectingCountry>
. 
<collectingRegion country="China" name="Chongqing">Chongqing</collectingRegion>
</emphasis>
: Jiulongpo, Jinfeng, Baihe Village, 
<emphasis italics="true" pageId="0" pageNumber="59">Jiulongpo Exped. 500107150402-289LY</emphasis>
(IMC). Mt. Jinyunshan, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.Y. Liu 182996</emphasis>
(IMC). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Fujian">Fujian</collectingRegion>
</emphasis>
: 
<normalizedToken originalValue="Hua’an">Hua'an</normalizedToken>
, Wenhua, 
<emphasis italics="true" pageId="0" pageNumber="59">W.D. Han 20667</emphasis>
(NF). Pinghe, Daxi, Jiangzhai Village, 
<emphasis italics="true" pageId="0" pageNumber="59">H.B. Chen s.n.</emphasis>
(FJSI). Xiamen, Yunding Cliff, 
<emphasis italics="true" pageId="0" pageNumber="59">G.D. Ye 1208</emphasis>
(IBSC). Yunxiao, Huotian, Baihuayang Reservoir, 
<emphasis italics="true" pageId="0" pageNumber="59">G.D. Ye 2482</emphasis>
(FJSI). Zhangzhou, Zhaoan, Wushan, 
<emphasis italics="true" pageId="0" pageNumber="59">
X.F. Zeng 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZXF19839">ZXF19839</accessionNumber>
</emphasis>
(CZH). Zhangzhou, Zhaoan, Wushan, Jinshui Village, 
<emphasis italics="true" pageId="0" pageNumber="59">
X.F. Zeng 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/ZXF41029">ZXF41029</accessionNumber>
</emphasis>
(CZH). 
<normalizedToken originalValue="Zhao’an">Zhao'an</normalizedToken>
, Jinshui Village, 
<emphasis italics="true" pageId="0" pageNumber="59">s. coll.</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">s.n.</emphasis>
(AU). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Guangdong">Guangdong</collectingRegion>
.
</emphasis>
Boluo, Mt. Luofushan, near Damiao, 
<emphasis italics="true" pageId="0" pageNumber="59">Yue78 5714</emphasis>
(IBSC). Dianbai, Luokeng, Mt. Shuangjiling, 
<emphasis italics="true" pageId="0" pageNumber="59">H.G. Ye 6379</emphasis>
(IBSC). Huidong, Pingshan Forest Farm, Mt. Chenshuishan, Zhulian?, 
<emphasis italics="true" pageId="0" pageNumber="59">P.Y. Chen</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">B.H. Chen &amp; G.C. Zhang 46</emphasis>
(IBSC). Jiangmen, Mt. Guifengshan, 
<emphasis italics="true" pageId="0" pageNumber="59">J.Y. Chen 20165220</emphasis>
(SN). Qingyuan, Yangshan, near Qincaitang Reservoir, 
<emphasis italics="true" pageId="0" pageNumber="59">K.W. Jiang KwT033</emphasis>
(NPH). Ruyuan, Daqiao Health Center, back mountain, 
<emphasis italics="true" pageId="0" pageNumber="59">Yue73 1182</emphasis>
(CSFI). Shenzhen, Longgang, 
<normalizedToken originalValue="Nan’ao">Nan'ao</normalizedToken>
, Yangmeikeng, 
<emphasis italics="true" pageId="0" pageNumber="59">S.Z. Zhang</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">L.Q. Li et al. 185</emphasis>
(SZG). 
<normalizedToken originalValue="Xin’an">Xin'an</normalizedToken>
, Ng-tung Shan, 
<emphasis italics="true" pageId="0" pageNumber="59">T.M. Tsui 231</emphasis>
(NAS). Zhaoqing, Mt. Dinghushan, 
<emphasis italics="true" pageId="0" pageNumber="59">S.J. Li 30</emphasis>
(IBSC). 
<emphasis italics="true" pageId="0" pageNumber="59">ibid.</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.Q. Song 2021057</emphasis>
(IBSC). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
</emphasis>
: Liuzhou, Longtan Park, Mt. Jiaodingshan, 
<emphasis italics="true" pageId="0" pageNumber="59">Longtan &amp; Dule Exped. 242</emphasis>
(IBK). Yang-shoh, 
<emphasis italics="true" pageId="0" pageNumber="59">H. Fung 21112</emphasis>
(SYS). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Guizhou">Guizhou</collectingRegion>
</emphasis>
: Tungtze, 
<emphasis italics="true" pageId="0" pageNumber="59">Y. Tsiang 4894</emphasis>
(IBSC). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Hainan">Hainan</collectingRegion>
</emphasis>
: Kan-en, Chim Fung Ling, near Sam Mo Watt Village, 
<emphasis italics="true" pageId="0" pageNumber="59">S.K. Lau 3582</emphasis>
(IBSC). 
<normalizedToken originalValue="Lin’gao">Lin'gao</normalizedToken>
, Maniao, Wende Village, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.X. Li et al. 911</emphasis>
(IBSC). Qionghai, Lehui, near Shuangbang Village, 
<emphasis italics="true" pageId="0" pageNumber="59">Y. Zhong 4472</emphasis>
(IBSC). Wanning, Mt. Dongshan, the second mountain range, 
<emphasis italics="true" pageId="0" pageNumber="59">S.P. Kao 52115</emphasis>
(IBSC). Wenchang, Longlou, nera Mt. Beijianshan, 
<emphasis italics="true" pageId="0" pageNumber="59">G.W. Tang</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.M. Li &amp; J. Li TangGW2525</emphasis>
(IBSC). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Hubei">Hubei</collectingRegion>
</emphasis>
: Shennongjia, Xingshan to Yangri, 
<emphasis italics="true" pageId="0" pageNumber="59">D.G. Zhang ZB130226624</emphasis>
(JIU). Xingshan, Xiakou, Jianyangping, Lifangyan to Huangliang, 
<emphasis italics="true" pageId="0" pageNumber="59">D.G. Zhang zdg4185</emphasis>
(JIU). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Hunan">Hunan</collectingRegion>
</emphasis>
: Jianghua, 
<emphasis italics="true" pageId="0" pageNumber="59">C.J. Qi 3822</emphasis>
(CSFI, IBSC). Xiangxi, Yongshun, Zejia, Donglu Village, 
<emphasis italics="true" pageId="0" pageNumber="59">K.D. Lei ZZ40516121</emphasis>
(JIU). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingRegion country="China" name="Jiangxi">Jiangxi</collectingRegion>
</emphasis>
: 
<normalizedToken originalValue="Ji’an">Ji'an</normalizedToken>
, Suichuan, Daijiapu, Xianmo, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.C. Liu</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">W.J. Xiong</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">F. Ye</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">L. Deng</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">M. Tu</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">X.J. Zhang</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">L. Feng</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="59">Q.Y. Yin &amp; N.N. Liu LXP-13-23687</emphasis>
(SYS). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Taiwan">Taiwan</collectingCountry>
</emphasis>
: 
<collectingRegion country="Taiwan" name="Hsinchu">Hsinchu</collectingRegion>
, Hsienchiaoshih, 
<emphasis italics="true" pageId="0" pageNumber="59">Z.-H. Chen 277</emphasis>
(TAIF). 
<emphasis bold="true" pageId="0" pageNumber="59">Yunnan</emphasis>
: Hekou, Erqu, 
<emphasis italics="true" pageId="0" pageNumber="59">W.X. Liu 277</emphasis>
(HITBC). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Micronesia">Micronesia</collectingCountry>
.
</emphasis>
Caroline Islands, 
<collectingRegion country="Micronesia" name="Yap">Yap</collectingRegion>
Group, Gorror Island, Central Plateau, 
<emphasis italics="true" pageId="0" pageNumber="59">E.Y. Hosaka 3319</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="India">India</collectingCountry>
.
</emphasis>
Kaswar, 
<emphasis italics="true" pageId="0" pageNumber="59">R.J. Bell 7750</emphasis>
(K). Kuppam River, Taliparamba, 
<emphasis italics="true" pageId="0" pageNumber="59">C.A. Barber 8788</emphasis>
(K). North Kanara, 
<emphasis italics="true" pageId="0" pageNumber="59">W.A. Talbot 1256</emphasis>
(K). S. Andaman, 
<emphasis italics="true" pageId="0" pageNumber="59">
Dr 
<normalizedToken originalValue="King’s">King's</normalizedToken>
Collector s.n.
</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Myanmar">Myanmar</collectingCountry>
.
</emphasis>
Myebon, 
<emphasis italics="true" pageId="0" pageNumber="59">H.S. McKee 6069</emphasis>
(K). 
<collectingRegion country="Myanmar" name="Yangon">Rangoon</collectingRegion>
, 
<emphasis italics="true" pageId="0" pageNumber="59">D.R. Khant 1079</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Mauritius">Mauritius</collectingCountry>
.
</emphasis>
The Pouce, 
<emphasis italics="true" pageId="0" pageNumber="59">J. Gueko s.n.</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="New Caledonia">New Caledonia</collectingCountry>
.
</emphasis>
Yate, Touaourou, 
<emphasis italics="true" pageId="0" pageNumber="59">s. coll. s.n.</emphasis>
(K). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Thailand">Thailand</collectingCountry>
.
</emphasis>
Narithiwat: Kulok river mouth next to bridge on road from 
<collectingRegion country="Thailand" name="Tak">Tak</collectingRegion>
Bai to Sungai Ko-lok, 
<emphasis italics="true" pageId="0" pageNumber="59">
P.S. Herendeen &amp; R. Pooma 
<date value="1999-05-01">1-V-1999</date>
-3
</emphasis>
(US). 
<emphasis bold="true" pageId="0" pageNumber="59">
<collectingCountry name="Vanuatu">Vanuatu</collectingCountry>
.
</emphasis>
Aniwa Island, Isavai village, 
<emphasis italics="true" pageId="0" pageNumber="59">P. Curry 1447</emphasis>
(K). Banks Islands, Port Patterson, 
<emphasis italics="true" pageId="0" pageNumber="59">A. Morrison s.n.</emphasis>
(K). New Hebrides, Erromanga, between Nouanko Camp and Ipota, about 
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
E of Ipota, 
<emphasis italics="true" pageId="0" pageNumber="59">P.S. Green RSNH1318</emphasis>
(K). New Hebrides, Port Vila, 
<emphasis italics="true" pageId="0" pageNumber="59">A. Morrison s.n.</emphasis>
(K).
</paragraph>
</subSubSection>
</treatment>
</document>