147 lines
17 KiB
XML
147 lines
17 KiB
XML
<document id="19E1735658FC90DC466AD702C9B48724" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="2CC8B30BDD2C8AAC82039D5B0BD65E70" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus benoiti Guignot 1953" docType="treatment" docVersion="5" lastPageNumber="140" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="139" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="0E3D0DC2AC456D31D786BF26205296EF">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="DDE7C2D9C972E53C48751CCC021D331C">Bistroem, Olof</mods:namePart>
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<mods:namePart id="BE86C60F0637CA0747DC73D6FA8A5BD7">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="76E657AFA2330C28BC7E8731BDF9C2D7">Bergsten, Johannes</mods:namePart>
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<mods:date id="4EA2CCA87C1495D5767A4D855B46084B">2015</mods:date>
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<mods:number id="8A85BF4FCE55C8A1BC465AF803999B6F">542</mods:number>
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<mods:classification id="37B75FE410FF592ABE2BC5196E089405">journal article</mods:classification>
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<mods:identifier id="B68CCEA7DE9C3F542259244E2D392A61" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="2CC8B30BDD2C8AAC82039D5B0BD65E70" ID-GBIF-Taxon="127862767" LSID="urn:lsid:plazi:treatment:2CC8B30BDD2C8AAC82039D5B0BD65E70" httpUri="http://treatment.plazi.org/id/2CC8B30BDD2C8AAC82039D5B0BD65E70" lastPageId="139" lastPageNumber="140" pageId="138" pageNumber="139">
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<subSubSection id="632F80089D347D51831BC1833BB74400" pageId="138" pageNumber="139" type="multiple">
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<paragraph id="958F1015DC9A568F166CC032CB3F5CD3" pageId="138" pageNumber="139">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="572CCB75DEA5D6A6007B7580E848D7ED" pageId="138" pageNumber="139" type="nomenclature">
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<paragraph id="72EB3B563D0BCF9416EBF8F0F206F7E5" pageId="138" pageNumber="139">
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<taxonomicName id="33583179D230263CB648C71E4511D2C3" ID-CoL="3RM5B" authority="Guignot, 1953" authorityName="Guignot" authorityYear="1953" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="138" pageNumber="139" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti Guignot, 1953</taxonomicName>
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Figs 106-107, 295-296, 447, 555
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</paragraph>
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</subSubSection>
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<subSubSection id="ED7543F9C210C52D5B8808AEC35396A3" pageId="139" pageNumber="140" type="reference_group">
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<paragraph id="E8A242D4477BAED6D78F87379FA1383D" pageId="139" pageNumber="140">
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<taxonomicName id="1DC679993AD8BA10CD7A5B95EFCA411C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">
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<pageBreakToken id="D0A682DC370B6D4BCAF5093AD266FB7D" pageId="139" pageNumber="140" start="start">Laccophilus</pageBreakToken>
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benoiti
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</taxonomicName>
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<bibRefCitation id="D5B8233FDF880F9DD63920344CB9957D" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="257" pageNumber="258" pagination="233 - 237" title="Hydrocanthares captures a la lumiere dans le Haut-Katanga." volume="47" year="1953 b">Guignot 1953b</bibRefCitation>
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||
: 234, 236 (original description, discussion, faunistics);
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<bibRefCitation id="38547F5C563B06391C14A24515A1B67D" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="257" pageNumber="258" pagination="181 - 189" title="Hydrocanthares captures a la lumiere dans le Haut-Katanga (3 e serie)." volume="51" year="1955 c">Guignot 1955c</bibRefCitation>
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: 182, 188 (faunistics, female description);
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<bibRefCitation id="AC0279ACAE04D126819FD665B6CA7D38" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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||
: 579, 581, 584 (description, faunistics);
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<bibRefCitation id="1FC3F3598CD64EF1962D5F5856E88D3C" author="Rocchi, S" journalOrPublisher="Roma" pageId="262" pageNumber="263" pagination="11 - 33" title="Contributo alla conoscenza dei ditiscidi di Guinea con descrizione di duo nuove specie (Coleoptera, Dytiscidae)." volumeTitle="Fragmenta Entomologica" year="2000">Rocchi 2000</bibRefCitation>
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: 25 (discussion);
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<bibRefCitation id="CEA9BCC68B62D852D997C47EAEADE959" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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||
: 240 (catalogue, faunistics);
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<bibRefCitation id="CCC59860D2F276376CD4B9651AEFC006" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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||
: 209 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="E6A9E4332311F5C4539FAE8FBD1E6040" pageId="139" pageNumber="140" type="type locality">
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<paragraph id="A078FF23DC133F4EB43B606EE70F5BD7" pageId="139" pageNumber="140">Type locality.</paragraph>
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<paragraph id="6191BEE6EB62953EC23E59904F1B1C32" pageId="139" pageNumber="140">Zaire: Elisabethville.</paragraph>
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</subSubSection>
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<subSubSection id="19C98BA7B6AE9FF1417C9D03C9916E85" pageId="139" pageNumber="140" type="type material">
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<paragraph id="ED3DF66762E36B8BCDFEEDE9847ECC7F" pageId="139" pageNumber="140">Type material studied</paragraph>
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<paragraph id="B3B2DF0A1402BCB75610694602C76667" pageId="139" pageNumber="140">
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(1 ex.). Holotype: male: "Holotypus / Coll. Mus. Congo Elisabethville, A la
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<normalizedToken id="6D02840C573E0E0B46AB04E16B5FED5A" originalValue="lumière">lumiere</normalizedToken>
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XI-50/VI-51 Ch. Seydel / Type / R. DET H. 6182 / Guignot det., 1953
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<taxonomicName id="588F7F1F02BC6169736466FB539833AF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti</taxonomicName>
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Guign. Type male" (MRAC; habitus in Fig. 447).
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</paragraph>
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</subSubSection>
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<subSubSection id="59AF46352BE67CE3A0CD4292FE632A9F" pageId="139" pageNumber="140" type="materials_examined">
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<paragraph id="B62968F726207F16380D0BFCF11F50DA" pageId="139" pageNumber="140">Additional material studied</paragraph>
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<paragraph id="1DEBFBDE6394098C4512D32A5385A923" pageId="139" pageNumber="140">
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(1 ex.). Zaire: "Allotypus female / Coll. Mus. Congo Elisabethville (
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<normalizedToken id="E2DAC9D6B52BFA362C301DAA320C8A75" originalValue="à">a</normalizedToken>
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la
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<normalizedToken id="12BF3AF9C98EEBBBC0F58A57173DE6EC" originalValue="lumière">lumiere</normalizedToken>
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) 1-III-52/30-IX-1953 Ch. Seydel / Allotype / R. DET H 6649 ee. / Guignot det., 1954
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<taxonomicName id="57DD11E6EC181460A816EF71C2E4916B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti</taxonomicName>
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Guign. Allotype female" (1 ex. MRAC; not type material). [Comment: attribution of the female specimen to
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<taxonomicName id="12FCDF5BA5E4EA80408E10AB23ED049D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti</taxonomicName>
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is based on
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<normalizedToken id="F1982F8F1ED5BD194021294FBEAA22D8" originalValue="Guignot’s">Guignot's</normalizedToken>
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determination.]
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</paragraph>
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</subSubSection>
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<subSubSection id="34B7ED441E9A8BB08A7517A5AB8A28CF" pageId="139" pageNumber="140" type="diagnosis">
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<paragraph id="2B0A3CB2980741DA979193182E55EE46" pageId="139" pageNumber="140">Diagnosis.</paragraph>
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<paragraph id="D2EFCE57C69CA95DEE2D80279885B923" pageId="139" pageNumber="140">
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<taxonomicName id="2797AACADE0D76236BE7C7119082D998" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti</taxonomicName>
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is characterized by exhibiting inconspicuous external characters but with very peculiar, strongly angled penis, which separates it from all other African
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<taxonomicName id="AF86870E731217ACE9C49504A31494CD" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
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species. Thus far only one male, however, is known and available for study. Comparison with males of
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<taxonomicName id="ACC9A014E33B9549711CFFAC5535F79F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus epinephes" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="epinephes">Laccophilus epinephes</taxonomicName>
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shows that their bodies are externally identical. The unique appearance of penis, being strongly bent, raises suspicion that it is a case of deformation and that
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<taxonomicName id="03B0CAE7807D22F804910613DF298ACE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus benoiti" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="benoiti">Laccophilus benoiti</taxonomicName>
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in fact is conspecific with
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<taxonomicName id="EC227A52C44DDF080F7D677556ECD941" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus epinephes" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="species" species="epinephes">Laccophilus epinephes</taxonomicName>
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. Further study is needed to settle this problem.
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</paragraph>
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</subSubSection>
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<subSubSection id="3A2ECE36AADFF5591D5B79786145FFE6" pageId="139" pageNumber="140" type="description">
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<paragraph id="484077140B6DD334F995EC8DE4043695" pageId="139" pageNumber="140">Description.</paragraph>
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<paragraph id="04DAF64FF3E1AFDAF3E89D2FCC013E8D" pageId="139" pageNumber="140">Body length 4.3-4.4 mm, width 2.1-2.2 mm. Habitus (Fig. 447), distinct colour pattern absent (in female specimen, studied, there is a semi-transverse row of small, pale spots (about five spots/elytron) posterior to foremargin of elytra).</paragraph>
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<paragraph id="2A4406EF7AFE0F6E43EFD30D9954A415" pageId="139" pageNumber="140">Head: Pale ferrugineous. Submat, finely microsculptured; reticulation in part indistinctly double. Large meshes hardly discernible; when discernible large meshes only slightly coarser than fine meshes. At eyes, areas with fine, irregular punctures, the areas which are extended a short distance towards middle of head-disc.</paragraph>
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<paragraph id="A6A9B465990FBDEB61DE1F72BEBC56D4" pageId="139" pageNumber="140">Pronotum: Pale ferrugineous, no distinct colour pattern. Very finely microsculptured; reticulation indistinctly double. Anteriorly and laterally with very fine, sparse and irregular punctures.</paragraph>
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<paragraph id="06C0543A4AB6154B1744222B632FF4C6" pageId="139" pageNumber="140">Elytra: Pale ferrugineous, with extensive and dense but somewhat indistinct, ferrugineous irrorations (Fig. 447). Rather shiny. Very finely microsculptured; reticulation very indistinctly double. Difference between coarser and fine meshes almost non-existent. Discal, dorsolateral and lateral row of punctures very fine and somewhat irregular.</paragraph>
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<paragraph id="07F8837240874E9995F0949FF6CB0AB4" pageId="139" pageNumber="140">Ventral aspect: Pale ferrugineous to ferrugineous, no distinct colour pattern discernible. Almost impunctate. Rather shiny, although very finely, in part indistinctly microsculptured. Semitransverse furrows on metacoxal plates shallow, in part indistinct. Abdomen basally with fine, curved striae. Prosternal process slender, apically pointed. Apical ventrite with a small, asymmetric knob (Fig. 106); ventrite broken.</paragraph>
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<paragraph id="5F467C97F88282FB41DF901E18118F0D" pageId="139" pageNumber="140">Legs: Pale ferrugineous, pro- and mesotarsus slightly enlarged, with suckers.</paragraph>
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<paragraph id="017C5B8B5E5B41B01B41446686791C47" pageId="139" pageNumber="140">
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Male genitalia: Penis strongly modified and different from all other African
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<taxonomicName id="7A3C24014F95E000F59693561F59AFA2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="139" pageNumber="140" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
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species; in lateral aspect penis forms an angle of almost 90° (Figs 295-296).
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</paragraph>
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<paragraph id="279669207EC04C17A9C4D60338E54BBC" pageId="139" pageNumber="140">Female: Externally as male, but elytra basally among irrorations with a semitransverse, irregular row of small pale spots. Apical ventrites simple, without knob (Fig. 107). Pro- and mesotarsus slender.</paragraph>
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</subSubSection>
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<subSubSection id="0404B8764F634173B624EE6A02495481" pageId="139" pageNumber="140" type="distribution">
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<paragraph id="6A91C68C63D0EB23824436A519AC5039" pageId="139" pageNumber="140">Distribution.</paragraph>
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<paragraph id="FD31DC0B2445361F1469FF07DABED1F1" pageId="139" pageNumber="140">Zaire (Fig. 555).</paragraph>
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</subSubSection>
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<subSubSection id="886D611B8BE38E15BB68D5915D56BDCB" pageId="139" pageNumber="140" type="collecting circumstances">
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<paragraph id="2C5782C813934F08CD45321266AB32F8" pageId="139" pageNumber="140">Collecting circumstances.</paragraph>
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<paragraph id="FC24432668C618DC9AF5AF7E4CD7221F" pageId="139" pageNumber="140">Almost unknown, sampled at light.</paragraph>
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</subSubSection>
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</treatment>
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</document> |