218 lines
22 KiB
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218 lines
22 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="1EF929764F0F55C793DB99C6A125930A" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina limitanea Godwin-Austen 1897" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199838993" LSID="urn:lsid:plazi:treatment:1EF929764F0F55C793DB99C6A125930A" httpUri="http://treatment.plazi.org/id/1EF929764F0F55C793DB99C6A125930A" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="1EF92976-4F0F-55C7-93DB-99C6A125930A" authority="Godwin-Austen, 1897" authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">Pupina limitanea Godwin-Austen, 1897</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">Figs 21P-R</figureCitation>
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">, 24D-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina limitaneus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitaneus">Pupina limitaneus</taxonomicName>
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[sic] Godwin-Austen, 1897: 40, pl. 69, fig. 4, 4a, b. Type locality: Eastern frontier of Burmah and Siam; Eastern Shan Plateau [Shan State, Myanmar].
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea" subGenus="Tylotoechus">Pupina (Tylotoechus) limitanea</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 316, 317.
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B60" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca-- III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca- - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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: 196.
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<bibRefCitation author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
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: 39.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Ancey" authorityYear="1904" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="brachysoma">Pupina brachysoma</taxonomicName>
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[non Ancey]-
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B71" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
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: 29, fig. 15f, Nam Ork Roo, Ban Nathong village, Namo District, Oudomxay Province.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
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NHMUK 1903.7.1.2967 (10 shells; Figs
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21P, Q</figureCitation>
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,
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24D, E</figureCitation>
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) from East of Burma & Siam.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2009-10-11" county="Nan Province" location="Pha Tub Cave" municipality="Mueang Nan District" specimenCount="1">
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CUMZ 12043 (
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<specimenCount type="generic">1 specimen</specimenCount>
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in ethanol) from
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<location LSID="urn:lsid:plazi:treatment:1EF929764F0F55C793DB99C6A125930A:13A26A52CCC18CC1E085F140C08F9780" county="Nan Province" municipality="Mueang Nan District" name="Pha Tub Cave">Pha Tub Cave</location>
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,
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<collectingMunicipality>Mueang Nan District</collectingMunicipality>
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,
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<collectingCounty>Nan Province</collectingCounty>
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,
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<collectingDate value="2009-10-11">11 Oct. 2009</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2017-01-19" county="Nan Province" location="Luang Sakoen Cave" municipality="Song Khwae District" specimenCount="1">
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CUMZ 12171 (1 shell) from
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<location LSID="urn:lsid:plazi:treatment:1EF929764F0F55C793DB99C6A125930A:DB3656BD1D4F7BE6EF02FAC1FA560A75" county="Nan Province" municipality="Song Khwae District" name="Luang Sakoen Cave">Luang Sakoen Cave</location>
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,
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<collectingMunicipality>Song Khwae District</collectingMunicipality>
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,
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<collectingCounty>Nan Province</collectingCounty>
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,
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<collectingDate value="2017-01-19">19 Jan. 2017</collectingDate>
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</materialsCitation>
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.
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<materialsCitation collectingDate="2015-01-18" county="Mae Hong Son Province" location="Mae Lana" municipality="Pang Mapha District" specimenCount="1">
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CUMZ 12044 (2 shells; Figs
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21R</figureCitation>
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,
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<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24F</figureCitation>
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) from
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<location LSID="urn:lsid:plazi:treatment:1EF929764F0F55C793DB99C6A125930A:846BE2BCB3C789447397EF9F385DA9A3" county="Mae Hong Son Province" municipality="Pang Mapha District" name="Mae Lana">Mae Lana</location>
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junction,
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<collectingMunicipality>Pang Mapha District</collectingMunicipality>
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,
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<collectingCounty>Mae Hong Son Province</collectingCounty>
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,
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<collectingDate value="2015-01-18">18 Jan. 2015</collectingDate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell ovate; last whorl ca. 60% of shell height. Apertural lip highly thickened, not expanded. Both parietal and columellar teeth fin-shaped and very thickened; parietal tooth always covering posterior canal; columellar tooth either next to or covering slit-like anterior canal.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1897" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="limitanea">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina limitanea</emphasis>
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</taxonomicName>
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is most similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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in shell shape, but differs in having parietal and columellar teeth and apertural lip thickened, and a shorter last whorl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Eastern Myanmar, Laos (
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<bibRefCitation author="Godwin-Austen, HH" journalOrPublisher="Journal of Malacology" pageId="0" pageNumber="1" refId="B51" refString="Godwin-Austen, HH, 1897. Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1-46." title="Land and freshwater mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burma, Pegu, Tenasserim, Malaya Peninsula, Ceylon and other islands of the Indian Ocean. Vol. 2, Part 7: 1 - 46." year="1897">Godwin-Austen 1897</bibRefCitation>
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;
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B71" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. 2019</bibRefCitation>
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), and Nan Province, northern Thailand.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specimen of
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="brachysoma">
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<emphasis italics="true" pageId="0" pageNumber="1">P. brachysoma</emphasis>
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</taxonomicName>
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from Oudomxay Province, Laos figured in
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.834.28800" author="Inkhavilay, K" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 166" refId="B71" refString="Inkhavilay, K, Sutcharit, C, Bantaowong, U, Chanabun, R, Siriwut, W, Srisonchai, R, Pholyotha, A, Jirapatrasilp, P, Panha, S, 2019. Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys 834: 1 - 166, DOI: https://doi.org/10.3897/zookeys.834.28800" title="Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda)." url="https://doi.org/10.3897/zookeys.834.28800" volume="834" year="2019">Inkhavilay et al. (2019</bibRefCitation>
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: fig. 15f) is different from the type materials of
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="brachysoma">
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<emphasis italics="true" pageId="0" pageNumber="1">P. brachysoma</emphasis>
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</taxonomicName>
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(see below) in having a thick and large parietal tooth covering the posterior canal, whereas
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="brachysoma">
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<emphasis italics="true" pageId="0" pageNumber="1">P. brachysoma</emphasis>
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</taxonomicName>
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has a sharp triangular parietal tooth which is not thickened, making the posterior canal visible. Therefore, the specimen from Oudomxay Province, Laos is herein identified as
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. limitanea" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="limitanea">
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<emphasis italics="true" pageId="0" pageNumber="1">P. limitanea</emphasis>
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</taxonomicName>
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of the
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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species group, whereas
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. brachysoma" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="brachysoma">
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<emphasis italics="true" pageId="0" pageNumber="1">P. brachysoma</emphasis>
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</taxonomicName>
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belongs to the
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. aureola" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="aureola">
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<emphasis italics="true" pageId="0" pageNumber="1">P. aureola</emphasis>
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</taxonomicName>
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species group.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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As this species is highly similar to
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
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<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
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</taxonomicName>
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, more sampling of this species, with both morphometric and molecular phylogenetic analyses, are needed to resolve the relationship between these two species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |