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<mods:title id="CB7D8914D36CF80381DB1EA7CA5F7768">The systematics of the pseudoscorpion genus Indohya (Pseudoscorpiones: Hyidae) in Australia</mods:title>
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<mods:namePart id="56936B706454624286E5B7AB96B94937">Abrams, Kym M.</mods:namePart>
<mods:affiliation id="F90A56244BF87B171F6960B1C5E40770">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Department of Water and Environmental Regulation, Locked Bag 10, Joondalup DC, Western Australia 6919, Australia https: // orcid. org / 0000 - 0003 - 4543 - 9255</mods:affiliation>
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<mods:affiliation id="AC4CAE3118F7F66AF2FBF1B33A741DC3">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Adjunct, School of Biological Sciences, University of Western Australia, Crawley, Western Australia 6009, Australia &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552</mods:affiliation>
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<mods:affiliation id="76466704A37568FB331ED6FF132BFA6C">Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Present address: Biologic Environmental Survey, East Perth, Western Australia 6004, Australia https: // orcid. org / 0000 - 0001 - 7108 - 0552 &amp; Collections &amp; Research, Western Australian Museum, 49 Kew Street, Welshpool, Western Australia 6106, Australia mark. harvey @ museum. wa. gov. au; https: // orcid. org / 0000 - 0003 - 1482 - 0109 https: // orcid. org / 0009 - 0004 - 5278 - 4238 &amp; Subterranean Research and Groundwater Ecology (SuRGE) Group, Trace and Environmental DNA (TrEnD) Laboratory, School of Molecular and Life Sciences, Curtin University, Perth, Western Australia 6102, Australia https: // orcid. org / 0000 - 0002 - 0349 - 3803 Corresponding author</mods:affiliation>
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<taxonomicName id="4EA24D21FF900B42AFB0FD07FD90F6ED" authority="Harvey &amp; Burger" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[151,586,654,681]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="BBD6EAB0FF900B42AFB0FD07FEBCF6ED" bold="true" box="[151,358,654,681]" italics="true" pageId="106" pageNumber="107">Indohya karenae</emphasis>
Harvey &amp; Burger
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,
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(
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,
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<taxonomicName id="4EA24D21FF900B42AFB0FC92FECBF777" box="[151,273,795,819]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BBD6EAB0FF900B42AFB0FC92FF30F777" box="[151,234,795,819]" italics="true" pageId="106" pageNumber="107">Indohya</emphasis>
sp.
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:
<bibRefCitation id="ED334B53FF900B42AE3CFC92FE2EF777" author="Murienne, J. &amp; Harvey, M. S. &amp; Giribet, G." box="[283,500,795,819]" pageId="106" pageNumber="107" pagination="170 - 184" refId="ref84587" refString="Murienne, J., Harvey, M. S. &amp; Giribet, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49, 170 - 184. https: // doi. org / 10.1016 / j. ympev. 2008.06.002" type="journal article" year="2008">
Murienne
<emphasis id="BBD6EAB0FF900B42AEA3FC95FE6DF777" box="[388,439,795,819]" italics="true" pageId="106" pageNumber="107">et al.</emphasis>
2008
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: figs. 35.
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<emphasis id="BBD6EAB0FF900B42AFB0FCE8FE5AF73F" bold="true" box="[151,384,865,891]" pageId="106" pageNumber="107">Material examined.</emphasis>
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female
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.
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:
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:
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, plateau above
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,
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, sifted litter,
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<emphasis id="BBD6EAB0FF900B42AB48FC0CFA88F7DB" box="[1135,1362,901,927]" italics="true" pageId="106" pageNumber="107">Corymbia clavigera</emphasis>
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group tree,
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,
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(
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).
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</paragraph>
<paragraph id="891D36A2FF900B42AFE0FC47FEABF048" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
<materialsCitation id="39CA3CFFFF900B42AFE0FC47FEB7F048" collectingDate="2007-03-19" collectionCode="WAM" collectorName="J. M. Waldock &amp; K. L. Edward" country="Australia" latitude="-14.436556" location="Northeast Beach" longLatPrecision="1" longitude="124.98908" municipality="South Maret Island" pageId="106" pageNumber="107" specimenCode="T82510, T153117" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
<emphasis id="BBD6EAB0FF900B42AFE0FC47FEE3F7A3" box="[199,313,974,999]" italics="true" pageId="106" pageNumber="107">
<typeStatus id="56198800FF900B42AFE0FC47FEE3F7A3" box="[199,313,974,999]" pageId="106" pageNumber="107" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="F1B57632FF900B42AE6EFC44FE36F7A3" box="[329,492,973,999]" name="Australia" pageId="106" pageNumber="107">AUSTRALIA</collectingCountry>
:
<emphasis id="BBD6EAB0FF900B42AEDCFC47FD1CF7A3" box="[507,710,973,999]" italics="true" pageId="106" pageNumber="107">
<collectingRegion id="4B66F840FF900B42AEDCFC47FD1CF7A3" box="[507,710,973,999]" country="Australia" name="Western Australia" pageId="106" pageNumber="107">Western Australia</collectingRegion>
</emphasis>
: 1 tritonymph, 1 protonymph, collected with holotype (
<collectionCode id="EFB3AE67FF900B42AA71FC44FA47F7A3" box="[1366,1437,973,999]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="106" pageNumber="107" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF900B42AFB0FC78FF2BF048" box="[151,241,1009,1036]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="106" pageNumber="107">T82510</specimenCode>
,
<specimenCode id="D9049ED9FF900B42AFDBFC78FEBFF048" box="[252,357,1009,1036]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="106" pageNumber="107">T153117</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="891D36A2FF900B42AFE0FB9CFD40F017" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
<materialsCitation id="39CA3CFFFF900B42AFE0FB9CFD4CF017" collectingDate="2007-03-19" collectionCode="WAM" collectorName="J. M. Waldock &amp; K. L. Edward" country="Australia" latitude="-14.436556" location="Northeast Beach" longLatPrecision="1" longitude="124.98908" municipality="South Maret Island" pageId="106" pageNumber="107" specimenCode="T83090, T129138" specimenCount="1" stateProvince="Western Australia">
<emphasis id="BBD6EAB0FF900B42AFE0FB9CFEB7F06B" box="[199,365,1045,1071]" italics="true" pageId="106" pageNumber="107">Other material</emphasis>
.
<collectingCountry id="F1B57632FF900B42AE5FFB9CFDC1F06B" box="[376,539,1045,1071]" name="Australia" pageId="106" pageNumber="107">AUSTRALIA</collectingCountry>
:
<emphasis id="BBD6EAB0FF900B42AD03FB9FFD30F06B" box="[548,746,1045,1071]" italics="true" pageId="106" pageNumber="107">
<collectingRegion id="4B66F840FF900B42AD03FB9FFD30F06B" box="[548,746,1045,1071]" country="Australia" name="Western Australia" pageId="106" pageNumber="107">Western Australia</collectingRegion>
</emphasis>
: 2 nymphs, collected with holotype (
<collectionCode id="EFB3AE67FF900B42ABA6FB9CFB12F06B" box="[1153,1224,1045,1071]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="106" pageNumber="107" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF900B42ABEBFB9CFAFCF06B" box="[1228,1318,1045,1071]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="106" pageNumber="107">T83090</specimenCode>
,
<specimenCode id="D9049ED9FF900B42AA08FB9CFA42F06B" box="[1327,1432,1045,1071]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="106" pageNumber="107">T129138</specimenCode>
; specimens destroyed during DNA extraction)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF900B42AFE0FBD4FAD0F10B" pageId="106" pageNumber="107" type="diagnosis">
<paragraph id="891D36A2FF900B42AFE0FBD4FCA0F0FB" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AFE0FBD4FE98F033" bold="true" box="[199,322,1117,1143]" pageId="106" pageNumber="107">Diagnosis.</emphasis>
<taxonomicName id="4EA24D21FF900B42AE6DFBD4FDD2F033" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[330,520,1117,1143]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="BBD6EAB0FF900B42AE6DFBD4FDD2F033" box="[330,520,1117,1143]" italics="true" pageId="106" pageNumber="107">Indohya karenae</emphasis>
</taxonomicName>
is an epigean species with two pairs of eyes and 14 setae on the carapace, thus resembling
<taxonomicName id="4EA24D21FF900B42AE3BFB0BFE5FF0DF" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[284,389,1153,1179]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="boltoni">
<emphasis id="BBD6EAB0FF900B42AE3BFB0BFE5FF0DF" box="[284,389,1153,1179]" italics="true" pageId="106" pageNumber="107">I. boltoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4EA24D21FF900B42AEB2FB0BFE34F0DF" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[405,494,1153,1179]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="cardo">
<emphasis id="BBD6EAB0FF900B42AEB2FB0BFE34F0DF" box="[405,494,1153,1179]" italics="true" pageId="106" pageNumber="107">I. cardo</emphasis>
</taxonomicName>
and
<taxonomicName id="4EA24D21FF900B42AD0FFB0BFD71F0DF" box="[552,683,1153,1179]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="juliannae">
<emphasis id="BBD6EAB0FF900B42AD0FFB0BFD71F0DF" box="[552,683,1153,1179]" italics="true" pageId="106" pageNumber="107">I. juliannae</emphasis>
</taxonomicName>
. It differs from them all by the robust chelal hand [hand (without pedicel) 1.39 (
<specimenCount id="9FA4FD2BFF900B42AE1CFB2CFE94F0FB" box="[315,334,1189,1215]" pageId="106" pageNumber="107" type="female"></specimenCount>
), vs. 1.541.92 (Ô), 1.541.86 (
<specimenCount id="9FA4FD2BFF900B42AD9CFB2CFD17F0FB" box="[699,717,1189,1215]" pageId="106" pageNumber="107" type="female"></specimenCount>
) in the others].
</paragraph>
<paragraph id="891D36A2FF900B42AFE0FB43FAD0F10B" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
It also differs from all other
<taxonomicName id="4EA24D21FF900B42AD35FB40FD12F0A7" box="[530,712,1225,1251]" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AD35FB40FDB4F0A7" box="[530,622,1225,1251]" italics="true" pageId="106" pageNumber="107">Indohya</emphasis>
species
</taxonomicName>
for which sequence data are available by five synapomorphies in
<emphasis id="BBD6EAB0FF900B42AF91FB67FF3CF14C" box="[182,230,1262,1288]" italics="true" pageId="106" pageNumber="107">COI</emphasis>
mtDNA: at base 48 there is a substitution to A; at base 350 there is a substitution to A; at base 351 there is a substitution to G; at base 479 there is a substitution to G; and at base 553 there is a substitution to C. The two sequenced specimens differ from all other sequenced specimens of
<taxonomicName id="4EA24D21FF900B42ACADFABCFC3CF10B" box="[906,998,1333,1359]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="106" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="BBD6EAB0FF900B42ACADFABCFC3CF10B" box="[906,998,1333,1359]" italics="true" pageId="106" pageNumber="107">Indohya</emphasis>
</taxonomicName>
by 12.729.6% (Table 2).
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF900B43AFE0FAD0FCF6F3B1" lastPageId="107" lastPageNumber="108" pageId="106" pageNumber="107" type="description">
<paragraph id="891D36A2FF900B42AFE0FAD0FDD5F1D3" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AFE0FAD0FE6BF137" bold="true" box="[199,433,1369,1395]" pageId="106" pageNumber="107">Description (adult).</emphasis>
<emphasis id="BBD6EAB0FF900B42AE9EFAD0FDC8F137" box="[441,530,1369,1395]" italics="true" pageId="106" pageNumber="107">Colour:</emphasis>
mostly brown-yellow; carapace, cheliceral fingers, pedipalpal trochanter, femur and patella darker reddish brown.
</paragraph>
<paragraph id="891D36A2FF900B42AFE0FA2BFAE0F1FF" blockId="106.[151,1437,865,2007]" box="[199,1338,1441,1468]" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AFE0FA2BFE4AF1FF" box="[199,400,1441,1468]" italics="true" pageId="106" pageNumber="107">Setae and cuticle:</emphasis>
setae long, mostly straight and acicular; most cuticular surfaces smooth and glossy.
</paragraph>
<paragraph id="891D36A2FF900B42AFE0FA4CFC0AF20F" blockId="106.[151,1437,865,2007]" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AFE0FA4CFEE4F19B" box="[199,318,1477,1503]" italics="true" pageId="106" pageNumber="107">Chelicera:</emphasis>
surface smooth; hand with 7 (
<specimenCount id="9FA4FD2BFF900B42ADB0FA4CFD73F19B" box="[663,681,1477,1503]" pageId="106" pageNumber="107" type="female"></specimenCount>
) setae, movable finger with 1 sub-medial seta, all setae acuminate; galea present, long, slender and slightly curved; fixed finger with 12 (
<specimenCount id="9FA4FD2BFF900B42AC89FA60FC1AF247" box="[942,960,1513,1539]" pageId="106" pageNumber="107" type="female"></specimenCount>
) small teeth; movable finger with 7 small (
<specimenCount id="9FA4FD2BFF900B42AF85F984FF6EF263" box="[162,180,1549,1575]" pageId="106" pageNumber="107" type="female"></specimenCount>
) teeth, each approximately same size; with 2 dorsal and 1 ventral lyrifissures; lamina exterior absent; rallum with 7 blades, all blades with anterior spinules, basal blade shorter than others.
</paragraph>
<paragraph id="891D36A2FF900B43AFE0F9DCFD2DF4BD" blockId="106.[151,1437,865,2007]" lastBlockId="107.[151,1437,151,2037]" lastPageId="107" lastPageNumber="108" pageId="106" pageNumber="107">
<emphasis id="BBD6EAB0FF900B42AFE0F9DCFE69F22B" box="[199,435,1621,1648]" italics="true" pageId="106" pageNumber="107">
Pedipalp (
<figureCitation id="11992A27FF900B42AE1AF9DFFE7EF22B" box="[317,420,1621,1648]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="106" pageNumber="107">Fig. 60B</figureCitation>
):
</emphasis>
robust; trochanter smooth, 2.33 (
<specimenCount id="9FA4FD2BFF900B42AC08F9DCFC9BF22B" box="[815,833,1621,1647]" pageId="106" pageNumber="107" type="female"></specimenCount>
) ×; femur broadened distally, retrolateral face lightly granulate, without dorsal tactile seta, with 3 stout retrolateral setae near base, 3.97 (
<specimenCount id="9FA4FD2BFF900B42AB1FF9F0FB90F2D7" box="[1080,1098,1657,1683]" pageId="106" pageNumber="107" type="female"></specimenCount>
) ×; patella broadened distally, prolateral face lightly granulate, pedicel not strongly pronounced but basal portion slimmer than distal portion, with 4 lyrifissures situated basally on dorsal surface, including 3 long and 1 short lyrifissure, 2.48 (
<specimenCount id="9FA4FD2BFF900B42AB86F948FB69F29F" box="[1185,1203,1729,1755]" pageId="106" pageNumber="107" type="female"></specimenCount>
) ×; chela (
<figureCitation id="11992A27FF900B42AA0EF948FA4AF29F" box="[1321,1424,1729,1755]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="106" pageNumber="107">Fig. 60C</figureCitation>
): hand robust, with prolateral and retrolateral faces lightly granulate, chela (with pedicel) 3.13 (
<specimenCount id="9FA4FD2BFF900B42AB9EF96CFB11F2BB" box="[1209,1227,1765,1791]" pageId="106" pageNumber="107" type="female"></specimenCount>
) ×, chela (without pedicel) 2.93 (
<specimenCount id="9FA4FD2BFF900B42AE19F880FE8AF367" box="[318,336,1801,1827]" pageId="106" pageNumber="107" type="female"></specimenCount>
) ×, hand (without pedicel) 1.39 (
<specimenCount id="9FA4FD2BFF900B42ADF7F880FD38F367" box="[720,738,1801,1827]" pageId="106" pageNumber="107" type="female"></specimenCount>
) × longer than broad, movable finger 1.09 (
<specimenCount id="9FA4FD2BFF900B42ABF9F880FB2AF367" box="[1246,1264,1801,1827]" pageId="106" pageNumber="107" type="female"></specimenCount>
) × longer than hand (without pedicel). Fixed finger with 8 trichobothria,
<emphasis id="BBD6EAB0FF900B42AC3EF8A4FCE9F303" box="[793,819,1837,1863]" italics="true" pageId="106" pageNumber="107">eb</emphasis>
,
<emphasis id="BBD6EAB0FF900B42AC66F8A4FCBCF303" box="[833,870,1837,1863]" italics="true" pageId="106" pageNumber="107">esb</emphasis>
and
<emphasis id="BBD6EAB0FF900B42ACBCF8A4FC66F303" box="[923,956,1837,1863]" italics="true" pageId="106" pageNumber="107">isb</emphasis>
in straight row at base of finger,
<emphasis id="BBD6EAB0FF900B42AA0EF8A4FAE5F303" box="[1321,1343,1837,1863]" italics="true" pageId="106" pageNumber="107">ib</emphasis>
situated dorsally at the end of the chelal hand,
<emphasis id="BBD6EAB0FF900B42AD62F8D8FDBCF32F" box="[581,614,1873,1899]" italics="true" pageId="106" pageNumber="107">isb</emphasis>
at the basal end of the fixed finger,
<emphasis id="BBD6EAB0FF900B42ACDBF8D8FBC8F32F" box="[1020,1042,1873,1899]" italics="true" pageId="106" pageNumber="107">ib</emphasis>
situated on dorsum of chelal hand,
<emphasis id="BBD6EAB0FF900B42AFB0F8FFFF7DF3CB" box="[151,167,1910,1935]" italics="true" pageId="106" pageNumber="107">it</emphasis>
closer to
<emphasis id="BBD6EAB0FF900B42AE3DF8FFFEF4F3CB" box="[282,302,1910,1935]" italics="true" pageId="106" pageNumber="107">et</emphasis>
than to
<emphasis id="BBD6EAB0FF900B42AEA8F8FFFE74F3CB" box="[399,430,1910,1935]" italics="true" pageId="106" pageNumber="107">est</emphasis>
,
<emphasis id="BBD6EAB0FF900B42AE99F8FFFE08F3CB" box="[446,466,1910,1935]" italics="true" pageId="106" pageNumber="107">et</emphasis>
slightly distal to
<emphasis id="BBD6EAB0FF900B42ADBBF8FFFD76F3CB" box="[668,684,1910,1935]" italics="true" pageId="106" pageNumber="107">it</emphasis>
; movable finger with 4 trichobothria,
<emphasis id="BBD6EAB0FF900B42AB44F8FCFBA6F3CB" box="[1123,1148,1909,1935]" italics="true" pageId="106" pageNumber="107">sb</emphasis>
much closer to
<emphasis id="BBD6EAB0FF900B42AA11F8FCFA9EF3CB" box="[1334,1348,1909,1935]" italics="true" pageId="106" pageNumber="107">b</emphasis>
than to
<emphasis id="BBD6EAB0FF900B42AFB0F813FF70F3F7" box="[151,170,1946,1971]" italics="true" pageId="106" pageNumber="107">st</emphasis>
,
<emphasis id="BBD6EAB0FF900B42AF9EF813FF16F3F7" box="[185,204,1946,1971]" italics="true" pageId="106" pageNumber="107">st</emphasis>
situated much closer to
<emphasis id="BBD6EAB0FF900B42AEC4F813FE31F3F7" box="[483,491,1946,1971]" italics="true" pageId="106" pageNumber="107">t</emphasis>
than to
<emphasis id="BBD6EAB0FF900B42AD6DF810FDB9F3F7" box="[586,611,1945,1971]" italics="true" pageId="106" pageNumber="107">sb</emphasis>
, ratio
<emphasis id="BBD6EAB0FF900B42AD89F810FD32F3F7" box="[686,744,1945,1972]" italics="true" pageId="106" pageNumber="107">sbst</emphasis>
/
<emphasis id="BBD6EAB0FF900B42ADC8F810FCFEF3F7" box="[751,804,1945,1972]" italics="true" pageId="106" pageNumber="107">sbb</emphasis>
= 1.79 (
<specimenCount id="9FA4FD2BFF900B42ACAEF810FC46F3F7" box="[905,924,1945,1971]" pageId="106" pageNumber="107" type="female"></specimenCount>
),
<emphasis id="BBD6EAB0FF900B42AC92F813FC67F3F7" box="[949,957,1946,1971]" italics="true" pageId="106" pageNumber="107">t</emphasis>
acuminate. Both fingers straight in lateral view. Fixed finger smooth, movable finger with several retrolateral granulations basally. Chelal teeth (
<figureCitation id="11992A27FF900B42AA21F834FA4EF393" box="[1286,1428,1981,2007]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="106" pageNumber="107">Figs. 60D, E</figureCitation>
) juxtadentate, fixed finger with 48 (
<specimenCount id="9FA4FD2BFF910B43AD04FF1EFDEFF4F5" box="[547,565,151,177]" pageId="107" pageNumber="108" type="female"></specimenCount>
) teeth, teeth 1, 30, 32, 34, 36 acute, remainder truncate or rounded, movable finger with 19, low, barely discernible teeth. Venom apparatus present only in movable chelal finger, venom duct long, terminating in nodus ramosus between
<emphasis id="BBD6EAB0FF910B43ADA9FF56FD7DF4BD" box="[654,679,223,249]" italics="true" pageId="107" pageNumber="108">sb</emphasis>
and
<emphasis id="BBD6EAB0FF910B43ADF9FF56FD2BF4BC" box="[734,753,223,248]" italics="true" pageId="107" pageNumber="108">st</emphasis>
.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FE8AFD94F5E8" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FE8AFEA2F559" box="[199,376,259,285]" italics="true" pageId="107" pageNumber="108">Cephalothorax:</emphasis>
carapace (
<figureCitation id="11992A27FF910B43AED9FE8AFDB1F559" box="[510,619,259,285]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="107" pageNumber="108">Fig. 60A</figureCitation>
) sub-rectangular; smooth; anterior margin slightly convex; epistome absent; lateral margins slightly convex; 1.17 (
<specimenCount id="9FA4FD2BFF910B43ADB3FEAFFD7CF504" box="[660,678,294,320]" pageId="107" pageNumber="108" type="female"></specimenCount>
) × longer than broad;
<specimenCount id="9FA4FD2BFF910B43ACB8FEAEFC6EF505" box="[927,948,295,321]" pageId="107" pageNumber="108" type="female"></specimenCount>
with 14 setae arranged 4: 2: 4: 2: 2; without furrows; with two pairs of corneate eyes, situated near anterior margin of carapace. Manducatory process distally triangular, with 2 distal setae; pedipalpal coxa with 6 additional setae. Coxa I with anterior margin not modified; chaetotaxy of coxae IIV:
<specimenCount id="9FA4FD2BFF910B43AEE7FE1BFE0FF5E8" box="[448,469,402,428]" pageId="107" pageNumber="108" type="female"></specimenCount>
, 4: 4: 4: 5.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FE3FFAE0F61A" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FE3FFEDCF58B" box="[199,262,438,463]" italics="true" pageId="107" pageNumber="108">Legs:</emphasis>
femora I and II much longer than patellae I and II, respectively; femora I and II with basal swelling; femora I and II each with 2 slit sensilla, each directed transversely; junction between anterior femora and patellae perpendicular; junction between posterior femora and patellae slightly oblique; femora III and IV much smaller than patellae III and IV, respectively; femur + patella IV 2.43 (
<specimenCount id="9FA4FD2BFF910B43AC7EFDA8FCB1F67F" box="[857,875,545,571]" pageId="107" pageNumber="108" type="female"></specimenCount>
) × longer than broad; metatarsi shorter than tarsi; subterminal tarsal seta acuminate; claws smooth, arolium slightly shorter than claws; arolium not divided.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FDE1FD9AF755" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FDE1FEE0F6C6" box="[199,314,616,642]" italics="true" pageId="107" pageNumber="108">Abdomen:</emphasis>
tergites not divided; tergal chaetotaxy
<specimenCount id="9FA4FD2BFF910B43ADDCFDE0FCCAF6C7" box="[763,784,617,643]" pageId="107" pageNumber="108" type="female"></specimenCount>
, 4: 6: 6: 6: 6: 7: 8: 8: 11: 7 (including 4 tactile setae): 8 (including 4 tactile setae): 2; arranged in single rows; sternal chaetotaxy
<specimenCount id="9FA4FD2BFF910B43ACE7FD05FC0FF6E2" box="[960,981,652,678]" pageId="107" pageNumber="108" type="female"></specimenCount>
, 7: (1) 6 (1): (3) 8 (3): 11: 11: 11: 11: 11: 10: 6 (including 2 tactile setae): 2; setae of anterior genital operculum (sternite II) of
<specimenCount id="9FA4FD2BFF910B43AB44FD39FBA2F68E" box="[1123,1144,688,714]" pageId="107" pageNumber="108" type="female"></specimenCount>
minute; posterior tergites and sternites with several tactile setae; medial sternites without suture line; pleural membrane uniformly granulate; without setae; stigmatic helix present.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FC92FBD2F771" blockId="107.[151,1437,151,2037]" box="[199,1032,795,821]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FC92FEE0F771" box="[199,314,795,821]" italics="true" pageId="107" pageNumber="108">Genitalia:</emphasis>
female: with large gonosac that is covered with scattered pores.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FCB6FEFFF7AC" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FCB6FE41F71D" box="[199,411,831,857]" italics="true" pageId="107" pageNumber="108">Dimensions (mm):</emphasis>
<materialsCitation id="39CA3CFFFF910B43AE8FFCB7FE41F738" collectionCode="WAM" latitude="0.28" longLatPrecision="78" longitude="0.12" pageId="107" pageNumber="108" specimenCode="T126382" specimenCount="1" typeStatus="holotype">
female
<typeStatus id="56198800FF910B43AD27FCB7FDB8F71C" box="[512,610,830,856]" pageId="107" pageNumber="108" type="holotype">holotype</typeStatus>
(
<collectionCode id="EFB3AE67FF910B43AD52FCB7FD66F71C" box="[629,700,830,856]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="107" pageNumber="108" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF910B43ADEFFCB7FCEBF71C" box="[712,817,830,856]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="107" pageNumber="108">T126382</specimenCode>
). Body length (excluding chelicerae) 1.59. Pedipalp: trochanter
<geoCoordinate id="EC965065FF910B43AE35FCEBFE88F738" box="[274,338,866,892]" degrees="0.280" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.28">0.280</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AE7CFCEBFE41F738" box="[347,411,866,892]" degrees="0.120" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.12">0.120</geoCoordinate>
</materialsCitation>
, femur
<geoCoordinate id="EC965065FF910B43AED2FCEBFDEEF739" box="[501,564,866,893]" degrees="0.595" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.595">0.595</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AD1AFCEBFDA4F739" box="[573,638,866,893]" degrees="0.150" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.15">0.150</geoCoordinate>
, patella
<geoCoordinate id="EC965065FF910B43ADF8FCEBFCFAF739" box="[735,800,866,893]" degrees="0.410" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.41">0.410</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AC0EFCEBFCB2F739" box="[809,872,866,893]" degrees="0.165" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.165">0.165</geoCoordinate>
, chela (with pedicel)
<geoCoordinate id="EC965065FF910B43AB45FCEBFB7BF739" box="[1122,1185,866,893]" degrees="0.875" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.875">0.875</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AB8CFCEBFB36F739" box="[1195,1260,866,893]" degrees="0.280" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.28">0.280</geoCoordinate>
, chela (without pedicel) 0.820, hand (without pedicel) 0.390, movable finger length 0.425. Carapace
<geoCoordinate id="EC965065FF910B43AB5BFC0FFB67F7E5" box="[1148,1213,902,929]" degrees="0.450" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.45">0.450</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43ABE2FC0FFADEF7E5" box="[1221,1284,902,929]" degrees="0.385" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.385">0.385</geoCoordinate>
, anterior eye 0.040, posterior eye 0.035. Leg IV: femur + patella
<geoCoordinate id="EC965065FF910B43ADC5FC20FCF8F780" box="[738,802,937,964]" degrees="0.560" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.56">0.560</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AC0CFC20FCB1F780" box="[811,875,937,964]" degrees="0.210" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.21">0.210</geoCoordinate>
, tibia
<geoCoordinate id="EC965065FF910B43AC96FC20FC2BF787" box="[945,1009,937,963]" degrees="0.360" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.36">0.360</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43ACDDFC20FBE0F787" box="[1018,1082,937,963]" degrees="0.010" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.01">0.010</geoCoordinate>
, metatarsus
<geoCoordinate id="EC965065FF910B43ABE1FC20FADFF780" box="[1222,1285,937,964]" degrees="0.175" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.175">0.175</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AA29FC20FA95F780" box="[1294,1359,937,964]" degrees="0.070" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.07">0.070</geoCoordinate>
, tarsus
<geoCoordinate id="EC965065FF910B43AFB0FC44FF02F7AC" box="[151,216,973,1000]" degrees="0.270" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.27">0.270</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AFC6FC44FEFBF7AC" box="[225,289,973,1000]" degrees="0.055" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.055">0.055</geoCoordinate>
.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FC78FCE6F04F" blockId="107.[151,1437,151,2037]" box="[199,828,1009,1035]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FC78FE21F04F" bold="true" box="[199,507,1009,1035]" pageId="107" pageNumber="108">Description (tritonymph).</emphasis>
<emphasis id="BBD6EAB0FF910B43AD25FC78FD80F04F" box="[514,602,1009,1035]" italics="true" pageId="107" pageNumber="108">Colour:</emphasis>
pale yellow-brown.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FB9CFC17F06B" blockId="107.[151,1437,151,2037]" box="[199,973,1045,1071]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FB9CFEE4F06B" box="[199,318,1045,1071]" italics="true" pageId="107" pageNumber="108">Chelicera:</emphasis>
hand with 6 setae, movable finger with 1 sub-medial seta.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FBB1FD3EF0A5" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FBB1FEEFF016" box="[199,309,1080,1106]" italics="true" pageId="107" pageNumber="108">Pedipalp:</emphasis>
trochanter 2.42 ×, femur 3.65 ×, patella 2.50 ×, chela (with pedicel) 2.91 ×, chela (without pedicel) 2.75 ×, hand (without pedicel) 1.27 × longer than broad, movable finger 1.14 × longer than hand (without pedicel). Fixed finger with 7 trichobothria (
<figureCitation id="11992A27FF910B43AD29FB09FDAAF0DE" box="[526,624,1152,1178]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="107" pageNumber="108">Fig. 60F</figureCitation>
),
<emphasis id="BBD6EAB0FF910B43ADA5FB09FD46F0DE" box="[642,668,1152,1178]" italics="true" pageId="107" pageNumber="108">eb</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AD8EFB09FD14F0DE" box="[681,718,1152,1178]" italics="true" pageId="107" pageNumber="108">esb</emphasis>
,
<emphasis id="BBD6EAB0FF910B43ADFCFB08FD20F0DE" box="[731,762,1153,1178]" italics="true" pageId="107" pageNumber="108">est</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AC20FB08FCC1F0DE" box="[775,795,1153,1178]" italics="true" pageId="107" pageNumber="108">et</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AC00FB09FCE7F0DE" box="[807,829,1152,1178]" italics="true" pageId="107" pageNumber="108">ib</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AC6DFB08FCBEF0DE" box="[842,868,1153,1178]" italics="true" pageId="107" pageNumber="108">ist</emphasis>
and
<emphasis id="BBD6EAB0FF910B43ACBFFB08FC72F0DE" box="[920,936,1153,1178]" italics="true" pageId="107" pageNumber="108">it</emphasis>
present; movable finger with 3 trichobothria;
<emphasis id="BBD6EAB0FF910B43AFB0FB2DFF7FF0FA" box="[151,165,1188,1214]" italics="true" pageId="107" pageNumber="108">b</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AF92FB2DFF12F0F9" box="[181,200,1188,1213]" italics="true" pageId="107" pageNumber="108">st</emphasis>
and
<emphasis id="BBD6EAB0FF910B43AE25FB2DFED0F0F9" box="[258,266,1188,1213]" italics="true" pageId="107" pageNumber="108">t</emphasis>
present,
<emphasis id="BBD6EAB0FF910B43AE53FB2DFE5DF0F9" box="[372,391,1188,1213]" italics="true" pageId="107" pageNumber="108">st</emphasis>
much closer to
<emphasis id="BBD6EAB0FF910B43AD66FB2DFD93F0F9" box="[577,585,1188,1213]" italics="true" pageId="107" pageNumber="108">t</emphasis>
than to
<emphasis id="BBD6EAB0FF910B43AD8DFB2DFD62F0FA" box="[682,696,1188,1214]" italics="true" pageId="107" pageNumber="108">b</emphasis>
. Fixed chelal finger with 37 mostly quadrate, juxtadentate teeth, movable chelal finger with 5 low, juxtadentate teeth.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FB62FB5BF16E" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FB62FEA2F141" box="[199,376,1259,1285]" italics="true" pageId="107" pageNumber="108">Cephalothorax:</emphasis>
carapace 1.08 × longer than broad; with 12 setae, arranged 4: 0: 4: 2: 2, including posterior margin with 2 setae; smooth; without furrows; 2 pair of eyes. Coxal chaetotaxy 4: 4: 4: 5.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FABAFB93F109" blockId="107.[151,1437,151,2037]" box="[199,1097,1331,1357]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FABAFEDCF108" box="[199,262,1331,1356]" italics="true" pageId="107" pageNumber="108">Legs:</emphasis>
femur + patella IV 2.37 × longer than broad; metatarsi and tarsi not fused.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FADFFB7FF1D0" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FADFFEE0F134" box="[199,314,1366,1392]" italics="true" pageId="107" pageNumber="108">Abdomen:</emphasis>
tergal chaetotaxy 4: 6: 6: 7: 10: 9: 9: 9: 7: 6 (including 2 tactile setae): 6 (including 4 tactile setae): 2; sternal chaetotaxy 0: (1) 6 (1): (2) 6 (2): 10: 11: 9: 9: 10: 9: 7 (including 4 tactile setae): 2.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0FA16FB15F244" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0FA16FE40F1FC" box="[199,410,1438,1464]" italics="true" pageId="107" pageNumber="108">Dimensions (mm):</emphasis>
<materialsCitation id="39CA3CFFFF910B43AE81FA17FB6EF1FD" box="[422,1204,1438,1465]" collectionCode="WAM" collectorName="Body" latitude="0.205" location="Pedipalp" longLatPrecision="78" longitude="0.085" pageId="107" pageNumber="108" specimenCode="T82510" specimenCount="1">
<collectionCode id="EFB3AE67FF910B43AE81FA17FE30F1FC" box="[422,490,1438,1464]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="107" pageNumber="108" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF910B43AED2FA17FD95F1FD" box="[501,591,1438,1465]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="107" pageNumber="108">T82510</specimenCode>
.
<collectorName id="24575374FF910B43AD79FA17FD41F1FC" box="[606,667,1438,1464]" pageId="107" pageNumber="108">Body</collectorName>
length 1.00.
<location id="8C7D6079FF910B43AC1DFA17FC7AF1FC" LSID="urn:lsid:plazi:treatment:010B87B4FF900B45AFB0FD07FECEF5B1:8C7D6079FF910B43AC1DFA17FC7AF1FC" box="[826,928,1438,1464]" latitude="0.205" longLatPrecision="78" longitude="0.085" name="Pedipalp" pageId="107" pageNumber="108">Pedipalp</location>
: trochanter
<geoCoordinate id="EC965065FF910B43AB0CFA17FBB1F1FD" box="[1067,1131,1438,1465]" degrees="0.205" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.205">0.205</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AB53FA17FB6EF1FD" box="[1140,1204,1438,1465]" degrees="0.085" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.085">0.085</geoCoordinate>
</materialsCitation>
, femur
<geoCoordinate id="EC965065FF910B43AA37FA17FA8AF1FD" box="[1296,1360,1438,1465]" degrees="0.420" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.42">0.420</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AA7EFA17FA42F1FD" box="[1369,1432,1438,1465]" degrees="0.115" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.115">0.115</geoCoordinate>
, patella
<geoCoordinate id="EC965065FF910B43AFCEFA4BFEF3F198" box="[233,297,1474,1500]" degrees="0.300" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.3">0.300</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AE15FA4BFEA8F198" box="[306,370,1474,1500]" degrees="0.120" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.12">0.120</geoCoordinate>
, chela (with pedicel)
<geoCoordinate id="EC965065FF910B43AD43FA48FD7EF19F" box="[612,676,1473,1499]" degrees="0.640" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.64">0.640</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AD8AFA48FD37F19F" box="[685,749,1473,1499]" degrees="0.220" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.22">0.220</geoCoordinate>
,
<materialsCitation id="39CA3CFFFF910B43ADDFFA48FCF2F244" collectionCode="T" latitude="0.335" location="Carapace" longLatPrecision="78" longitude="0.31" pageId="107" pageNumber="108" specimenCount="1">
chela (without pedicel) 0.605, hand (without pedicel) length 0.280, movable finger length 0.320.
<location id="8C7D6079FF910B43AD08FA6CFD4DF1BB" LSID="urn:lsid:plazi:treatment:010B87B4FF900B45AFB0FD07FECEF5B1:8C7D6079FF910B43AD08FA6CFD4DF1BB" box="[559,663,1509,1535]" latitude="0.335" longLatPrecision="78" longitude="0.31" name="Carapace" pageId="107" pageNumber="108">Carapace</location>
<geoCoordinate id="EC965065FF910B43ADB8FA6CFD05F244" box="[671,735,1509,1536]" degrees="0.335" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.335">0.335</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43ADC0FA6CFCF2F244" box="[743,808,1509,1536]" degrees="0.310" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.31">0.310</geoCoordinate>
</materialsCitation>
. Leg IV: femur + patella
<geoCoordinate id="EC965065FF910B43AB66FA6CFB5BF244" box="[1089,1153,1509,1536]" degrees="0.355" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.355">0.355</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43ABADFA6CFB11F244" box="[1162,1227,1509,1536]" degrees="0.150" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.15">0.150</geoCoordinate>
.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F980FCA5F267" blockId="107.[151,1437,151,2037]" box="[199,895,1545,1572]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F980FDDDF267" bold="true" box="[199,519,1545,1571]" pageId="107" pageNumber="108">Description (protonymph).</emphasis>
<emphasis id="BBD6EAB0FF910B43AD29F980FDBCF267" box="[526,614,1545,1571]" italics="true" pageId="107" pageNumber="108">Colour:</emphasis>
very pale, yellow-white.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F9A4FC82F203" blockId="107.[151,1437,151,2037]" box="[199,856,1581,1607]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F9A4FEE4F203" box="[199,318,1581,1607]" italics="true" pageId="107" pageNumber="108">Chelicera:</emphasis>
hand with 4 setae, movable finger without seta.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F9D9FB12F292" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F9D9FEEFF22E" box="[199,309,1616,1642]" italics="true" pageId="107" pageNumber="108">Pedipalp:</emphasis>
trochanter 1.92 ×, femur 3.60 ×, patella 2.13 ×, chela (with pedicel) 3.23 ×, chela (without pedicel) 3.08 ×, hand (without pedicel) 1.38 × longer than broad, movable finger 1.22 × longer than hand (without pedicel). Fixed finger with 3 trichobothria (
<figureCitation id="11992A27FF910B43AD33F911FDA6F2F6" box="[532,636,1688,1714]" captionStart="FIGURE 60" captionStartId="108.[151,250,1854,1878]" captionTargetBox="[158,1436,195,1820]" captionTargetId="figure-19@108.[151,1436,181,1830]" captionTargetPageId="108" captionText="FIGURE 60. Indohya karenae Harvey &amp; Burger, n. sp., holotype female (WAM T126382), unless stated otherwise:A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E)." figureDoi="http://doi.org/10.5281/zenodo.8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="107" pageNumber="108">Fig. 60G</figureCitation>
),
<emphasis id="BBD6EAB0FF910B43ADA8F911FD73F2F6" box="[655,681,1688,1714]" italics="true" pageId="107" pageNumber="108">eb</emphasis>
,
<emphasis id="BBD6EAB0FF910B43AD90F910FD0BF2F6" box="[695,721,1689,1714]" italics="true" pageId="107" pageNumber="108">ist</emphasis>
and
<emphasis id="BBD6EAB0FF910B43AC2FF910FCC6F2F6" box="[776,796,1689,1714]" italics="true" pageId="107" pageNumber="108">et</emphasis>
present; movable finger with 1 trichobothrium,
<emphasis id="BBD6EAB0FF910B43AA12F910FAE7F2F6" box="[1333,1341,1689,1714]" italics="true" pageId="107" pageNumber="108">t</emphasis>
present; fixed chelal finger with 20 low, juxtadentate teeth, quadrate; movable chelal finger with 2 teeth.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F956FDD7F359" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F956FEA2F2BD" box="[199,376,1759,1785]" italics="true" pageId="107" pageNumber="108">Cephalothorax:</emphasis>
0.90 × longer than broad; with 12 setae arranged 4: 0: 4: 2: 2; without furrows; with 2 pairs of eyes. Coxal chaetotaxy 1: 1: 1: 1.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F8A1FD8AF305" blockId="107.[151,1437,151,2037]" box="[199,592,1831,1857]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F8A1FEDCF305" box="[199,262,1832,1857]" italics="true" pageId="107" pageNumber="108">Legs:</emphasis>
metatarsi and tarsi not fused.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F8C2FC93F3CC" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F8C2FEE0F321" box="[199,314,1867,1893]" italics="true" pageId="107" pageNumber="108">Abdomen:</emphasis>
tergal chaetotaxy 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: 2 (including 2 tactile setae): 2; sternal chaetotaxy 0: (0) 2 (0): (1) 2 (1): 4: 4: 4: 4: 4: 4: 4 (including 2 tactile setae): 2.
</paragraph>
<paragraph id="891D36A2FF910B43AFE0F81AFCF6F3B1" blockId="107.[151,1437,151,2037]" pageId="107" pageNumber="108">
<emphasis id="BBD6EAB0FF910B43AFE0F81AFE43F3E9" box="[199,409,1939,1965]" italics="true" pageId="107" pageNumber="108">Dimensions (mm):</emphasis>
<materialsCitation id="39CA3CFFFF910B43AE84F81AFB6FF3E9" box="[419,1205,1938,1965]" collectionCode="WAM" latitude="0.125" longLatPrecision="78" longitude="0.065" pageId="107" pageNumber="108" specimenCode="T153117" specimenCount="1">
<collectionCode id="EFB3AE67FF910B43AE84F81AFE3DF3E9" box="[419,487,1939,1965]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="107" pageNumber="108" type="Museum">WAM</collectionCode>
<specimenCode id="D9049ED9FF910B43AED6F81BFD82F3E9" box="[497,600,1938,1965]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="107" pageNumber="108">T153117</specimenCode>
. Body length 0.69. Pedipalp: trochanter
<geoCoordinate id="EC965065FF910B43AB0BF81BFBB6F3E9" box="[1068,1132,1938,1965]" degrees="0.125" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.125">0.125</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AB52F81BFB6FF3E9" box="[1141,1205,1938,1965]" degrees="0.065" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.065">0.065</geoCoordinate>
</materialsCitation>
, femur
<geoCoordinate id="EC965065FF910B43AA28F81BFA8AF3E9" box="[1295,1360,1938,1965]" degrees="0.270" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.27">0.270</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AA7EF81BFA42F3E9" box="[1369,1432,1938,1965]" degrees="0.075" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.075">0.075</geoCoordinate>
, patella 0.16/0.075, chela (with pedicel)
<geoCoordinate id="EC965065FF910B43AD7BF83FFD46F394" box="[604,668,1974,2000]" degrees="0.420" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.42">0.420</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43AD83F83FFD3EF394" box="[676,740,1974,2000]" degrees="0.130" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.13">0.130</geoCoordinate>
, chela (without pedicel) 0.400, hand (without pedicel) length 0.180, movable finger length 0.220. Carapace
<geoCoordinate id="EC965065FF910B43ADB8F853FD3AF3B1" box="[671,736,2010,2037]" degrees="0.230" direction="north" orientation="latitude" pageId="107" pageNumber="108" precision="55" value="0.23">0.230</geoCoordinate>
/
<geoCoordinate id="EC965065FF910B43ADCFF853FCF2F3B1" box="[744,808,2010,2037]" degrees="0.255" direction="east" orientation="longitude" pageId="107" pageNumber="108" precision="55" value="0.255">0.255</geoCoordinate>
.
</paragraph>
</subSubSection>
<caption id="DDDD662AFF960B44AFB0F8B7FB47F386" ID-DOI="http://doi.org/10.5281/zenodo.8324040" ID-Zenodo-Dep="8324040" httpUri="https://zenodo.org/record/8324040/files/figure.png" pageId="108" pageNumber="109" startId="108.[151,250,1854,1878]" targetBox="[158,1436,195,1820]" targetPageId="108" targetType="figure">
<paragraph id="891D36A2FF960B44AFB0F8B7FB47F386" blockId="108.[151,1437,1854,1987]" pageId="108" pageNumber="109">
<emphasis id="BBD6EAB0FF960B44AFB0F8B7FEC5F312" bold="true" box="[151,287,1854,1878]" pageId="108" pageNumber="109">FIGURE 60.</emphasis>
<taxonomicName id="4EA24D21FF960B44AE04F8B7FD5BF313" authority="Harvey &amp; Burger" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[291,641,1854,1879]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="karenae" status="sp. nov.">
<emphasis id="BBD6EAB0FF960B44AE04F8B7FE16F312" box="[291,460,1854,1878]" italics="true" pageId="108" pageNumber="109">Indohya karenae</emphasis>
Harvey &amp; Burger
</taxonomicName>
,
<emphasis id="BBD6EAB0FF960B44ADAEF8B7FD1BF312" bold="true" box="[649,705,1854,1878]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A0E557CBFF960B44ADAEF8B7FD1BF312" box="[649,705,1854,1878]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype female (WAM T126382), unless stated otherwise: A, carapace, dorsal (distorted); B, right pedipalp, dorsal; C, left chela, retrolateral; D, left movable finger, chelal teeth, retrolateral; E, left fixed finger, chelal teeth, retrolateral; F, right chela, retrolateral, paratype tritonymph (WAM T82510); G, left chela, retrolateral, paratype protonymph (WAM T153117). Scale lines = 0.2 mm (Figs. AC, F, G), 0.1 mm (Figs. D, E).
</paragraph>
</caption>
<subSubSection id="C1B86529FF970B45AFE0FF1FFBC4F5E9" pageId="109" pageNumber="110" type="distribution">
<paragraph id="891D36A2FF970B45AFE0FF1FFC5FF505" blockId="109.[151,1437,150,501]" pageId="109" pageNumber="110">
<emphasis id="BBD6EAB0FF970B45AFE0FF1FFE24F4F5" bold="true" box="[199,510,150,177]" pageId="109" pageNumber="110">Distribution and remarks.</emphasis>
<taxonomicName id="4EA24D21FF970B45AD24FF1EFD64F4F5" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[515,702,151,177]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="BBD6EAB0FF970B45AD24FF1EFD64F4F5" box="[515,702,151,177]" italics="true" pageId="109" pageNumber="110">Indohya karenae</emphasis>
</taxonomicName>
has only been collected from South Maret Island in the Kimberley region of
<collectingRegion id="4B66F840FF970B45AFDAFF32FE18F491" box="[253,450,187,213]" country="Australia" name="Western Australia" pageId="109" pageNumber="110">Western Australia</collectingRegion>
(
<figureCitation id="11992A27FF970B45AEE9FF32FDFFF491" box="[462,549,187,213]" captionStart="FIGURE 6" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[200,1392,188,1917]" captionTargetId="figure-19@22.[194,1394,181,1920]" captionTargetPageId="22" captionText="FIGURE 6. Maps of north-western Australia depicting recorded distributions of the three epigean species of Indohya from the Kimberley:A, epigean species; B, troglobitic species." figureDoi="http://doi.org/10.5281/zenodo.8323897" httpUri="https://zenodo.org/record/8323897/files/figure.png" pageId="109" pageNumber="110">Fig. 6A</figureCitation>
).The island are situated within the Northern Kimberley bioregion.The specimens were collected from leaf litter under a member of the apple gum complex that includes
<taxonomicName id="4EA24D21FF970B45AB73FF56FAEEF4BD" box="[1108,1332,223,249]" class="Insecta" family="Cerambycidae" genus="Corymbia" kingdom="Animalia" order="Coleoptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="clavigera">
<emphasis id="BBD6EAB0FF970B45AB73FF56FAEEF4BD" box="[1108,1332,223,249]" italics="true" pageId="109" pageNumber="110">Corymbia clavigera</emphasis>
</taxonomicName>
.
<taxonomicName id="4EA24D21FF970B45AA67FF56FF2AF559" authorityName="Harvey &amp; Burger" authorityYear="2023" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="BBD6EAB0FF970B45AA67FF56FF2AF559" italics="true" pageId="109" pageNumber="110">Indohya karenae</emphasis>
</taxonomicName>
was included in the molecular phylogeny of pseudoscorpions by
<bibRefCitation id="ED334B53FF970B45ACF1FE8AFB00F559" author="Murienne, J. &amp; Harvey, M. S. &amp; Giribet, G." box="[982,1242,259,285]" pageId="109" pageNumber="110" pagination="170 - 184" refId="ref84587" refString="Murienne, J., Harvey, M. S. &amp; Giribet, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49, 170 - 184. https: // doi. org / 10.1016 / j. ympev. 2008.06.002" type="journal article" year="2008">
Murienne
<emphasis id="BBD6EAB0FF970B45AB6CFE8AFB5EF559" box="[1099,1156,259,285]" italics="true" pageId="109" pageNumber="110">et al.</emphasis>
(2008)
</bibRefCitation>
. The species was previously known by WAM identification code
<taxonomicName id="4EA24D21FF970B45AD89FEAEFCD0F505" authority="PSE" authorityName="PSE" box="[686,778,295,321]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="109" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="BBD6EAB0FF970B45AD89FEAEFCD0F505" box="[686,778,295,321]" italics="true" pageId="109" pageNumber="110">Indohya</emphasis>
</taxonomicName>
PSE155.
</paragraph>
<paragraph id="891D36A2FF970B45AFE0FEC3FBC4F5E9" blockId="109.[151,1437,150,501]" pageId="109" pageNumber="110">
<emphasis id="BBD6EAB0FF970B45AFE0FEC3FE2EF521" bold="true" box="[199,500,330,357]" pageId="109" pageNumber="110">Conservation assessment</emphasis>
.
<taxonomicName id="4EA24D21FF970B45AD22FEC2FD1EF521" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[517,708,331,357]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="BBD6EAB0FF970B45AD22FEC2FD1EF521" box="[517,708,331,357]" italics="true" pageId="109" pageNumber="110">Indohya karenae</emphasis>
</taxonomicName>
has been collected at a single site on South Maret Island. The surface habitat appears to be relatively undisturbed and
<taxonomicName id="4EA24D21FF970B45AC2AFEE6FCA4F5CD" authorityName="Harvey &amp; Burger" authorityYear="2023" box="[781,894,367,393]" class="Arachnida" family="Hyidae" genus="Indohya" kingdom="Animalia" order="Pseudoscorpiones" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="karenae">
<emphasis id="BBD6EAB0FF970B45AC2AFEE6FCA4F5CD" box="[781,894,367,393]" italics="true" pageId="109" pageNumber="110">I. karenae</emphasis>
</taxonomicName>
does not meet the status of threatened using the criteria developed by the International Union for Conservation of Nature (2012).
</paragraph>
</subSubSection>
<subSubSection id="C1B86529FF970B45AFE0FE3EFECEF5B1" pageId="109" pageNumber="110" type="etymology">
<paragraph id="891D36A2FF970B45AFE0FE3EFECEF5B1" blockId="109.[151,1437,150,501]" pageId="109" pageNumber="110">
<emphasis id="BBD6EAB0FF970B45AFE0FE3EFE94F595" bold="true" box="[199,334,439,465]" pageId="109" pageNumber="110">Etymology.</emphasis>
This species is named for our friend and colleague Karen Cullen, one of the collectors of the
<typeStatus id="56198800FF970B45AA4BFE3EFA46F595" box="[1388,1436,439,465]" pageId="109" pageNumber="110">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
</treatment>
</document>