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<document id="8A9AEE7F719B02AC85FA47D730E88AA6" ID-DOI="10.5281/zenodo.5353544" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353544" ID-ZooBank="772BDDF2-AD14-4950-B3B3-D6A2C9E403C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630210667410" checkinUser="felipe" docAuthor="Gordon, Robert D., F, Guillermo González &amp; Hanley, Guy A." docDate="2020" docId="03D687B4FF9A6A17FF0AFE7BFA9AFA2B" docLanguage="en" docName="0766.pdf" docOrigin="Insecta Mundi 2020 (766)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Calloeneis johnnie Gordon and Hanley 2020, new species" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFEFFFCCFF976A1AFFCAFF83FFB6FF8F" masterDocTitle="South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="12" updateTime="1699187120983" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="16EE02597194773480B51BC0462E520C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0ECCEDE8E9FE268B329037E4B3A5DE91">
<mods:title id="D15FBE20E2805D8BA25AC41E8BB8729B">South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)</mods:title>
</mods:titleInfo>
<mods:name id="C44C0D884E53CA271F5AC334656623E2" type="personal">
<mods:role id="B7CA481AD4CDF344E935FA04A1A20F25">
<mods:roleTerm id="00B90E599C9E70BFDA743CF13B9F1BFB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D85B05CF5107EA19A715E2E945FAFD6E">Gordon, Robert D.</mods:namePart>
<mods:affiliation id="0EB4395D7798709451A8F58DC9D74E38">Northern Plains Entomology PO Box 65 Willow City, ND 58384, USA</mods:affiliation>
</mods:name>
<mods:name id="01C94812B439DFBA72EB5657787DE1C8" type="personal">
<mods:role id="1934057FC88E8D99D0EC78C682F7A572">
<mods:roleTerm id="BACEDAB7F300BA2C8C1681EDF6C0B768">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CA662B6DD0EEDBC1D9B995ED3191AB61">F, Guillermo González</mods:namePart>
<mods:affiliation id="4BB73EDE0ED81C4DDCF9DC3059CC4F15">Nocedal 6455 La Reina Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="AB29B083008B19FBB0264B774C1AA2E2" type="personal">
<mods:role id="870338B2B298437B19ED9F121A37254A">
<mods:roleTerm id="9018B1AB259AF6C6BAE9EEC80A130224">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E1C99B52B26DF64B983434DFFA630A9">Hanley, Guy A.</mods:namePart>
<mods:affiliation id="A128F74DCB59EF731321294E5F310DB4">Northern Plains Entomology Minot, ND</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8F1C708C315FB4F0C31AB571C0CC89E4">text</mods:typeOfResource>
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<mods:titleInfo id="78B107533A2C1D5CB0CC3DC6D8D70E00">
<mods:title id="981849B0DE1271BB62097919DF101473">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="6109597241D6B26CE1A953E6D7565BF9">
<mods:date id="EB64E63258A9E233B6CCC8C8A9AFB49D">2020</mods:date>
<mods:detail id="7CDEE9F34785B8CB594DD720AF8BCED7" type="pubDate">
<mods:number id="DEF59DA2436EC53ED38E03815B1BF768">2020-05-29</mods:number>
</mods:detail>
<mods:detail id="43892365EDF71988C9C764A0CC3F551A" type="volume">
<mods:number id="DE25AE30878EC1503276C950594B0451">2020</mods:number>
</mods:detail>
<mods:detail id="5D34AA3BF66AFBBC24D1C45BECF7468F" type="issue">
<mods:number id="DB8C6E66F00E6E97234273561E30DE43">766</mods:number>
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<mods:start id="FEAA5C291B64CCB2D5E5A1BD888700E1">1</mods:start>
<mods:end id="B1E46B308BBED9F8E28B2507FA02188E">25</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="542E3CC843C5DB157CEF66E73A9626B7">journal article</mods:classification>
<mods:identifier id="82022C4F58B397206CBBD2BAFCBC7932" type="DOI">10.5281/zenodo.5353544</mods:identifier>
<mods:identifier id="4DC0970AF323450324D1E8EA6BE4394F" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="523B9F2BC5147C44563835583CF275A1" type="Zenodo-Dep">5353544</mods:identifier>
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</mods:mods>
<treatment id="03D687B4FF9A6A17FF0AFE7BFA9AFA2B" LSID="urn:lsid:plazi:treatment:03D687B4FF9A6A17FF0AFE7BFA9AFA2B" httpUri="http://treatment.plazi.org/id/03D687B4FF9A6A17FF0AFE7BFA9AFA2B" lastPageNumber="12" pageId="13" pageNumber="12">
<subSubSection id="C3656529FF9A6A17FF0AFE7BFC05FD9F" box="[192,947,503,528]" pageId="13" pageNumber="12" type="nomenclature">
<paragraph id="8BC036A2FF9A6A17FF0AFE7BFC05FD9F" blockId="13.[192,947,503,528]" box="[192,947,503,528]" pageId="13" pageNumber="12">
<heading id="D08881CEFF9A6A17FF0AFE7BFC05FD9F" bold="true" box="[192,947,503,528]" fontSize="10" level="2" pageId="13" pageNumber="12" reason="3">
<emphasis id="B90BEAB0FF9A6A17FF0AFE7BFC05FD9F" bold="true" box="[192,947,503,528]" pageId="13" pageNumber="12">
13.
<taxonomicName id="4C7F4D21FF9A6A17FF24FE74FCB4FD9F" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2020" box="[238,770,503,528]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="johnnie" status="sp. nov.">
<emphasis id="B90BEAB0FF9A6A17FF24FE74FE5AFD80" bold="true" box="[238,492,503,527]" italics="true" pageId="13" pageNumber="12">Calloeneis johnnie</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A23857CBFF9A6A17FCC5FE7BFC05FD9F" box="[783,947,504,528]" pageId="13" pageNumber="12" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9A6A17FF0AFDABFCC0FB6C" pageId="13" pageNumber="12" type="description">
<paragraph id="8BC036A2FF9A6A17FF0AFDABFB7DFBCD" blockId="13.[191,1442,552,1090]" pageId="13" pageNumber="12">
<emphasis id="B90BEAB0FF9A6A17FF0AFDABFDF6FDCF" bold="true" box="[192,576,552,576]" pageId="13" pageNumber="12">
Description. Male
<typeStatus id="54C48800FF9A6A17FE0AFDABFD8CFDCF" box="[448,570,552,576]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="4C879B47FF9A6A17FD61FDABFCBDFDB0" box="[683,779,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="13" pageNumber="12" unit="mm" value="1.8">1.8 mm</quantity>
, width
<quantity id="4C879B47FF9A6A17FCA3FDABFC7CFDB0" box="[873,970,552,576]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="12" unit="mm" value="1.6">1.6 mm</quantity>
; dorsal surface somewhat alutaceous; head strongly alutaceous, pronotum, elytron with faint trace of microsculpture. Color reddish brown; head yellow; pronotum with lateral 1/3 yellow, median 1/3 brown; elytron reddish brown medially, anterior and lateral borders dark brown (
<figureCitation id="13442A27FF9A6A17FD74FD0BFCA3FD2F" box="[702,789,648,672]" captionStart="Figures 54–68" captionStartId="24.[192,287,1678,1700]" captionTargetBox="[210,1444,233,1610]" captionTargetId="figure-97@24.[192,1445,217,1656]" captionTargetPageId="24" captionText="Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule." figureDoi="http://doi.org/10.5281/zenodo.5353556" httpUri="https://zenodo.org/record/5353556/files/figure.png" pageId="13" pageNumber="12">Fig. 54</figureCitation>
); venter yellow except pro-, meso-, and metasternum brown; epipleuron reddish yellow; abdomen brownish yellow. Head with punctures not visible; pronotal punctures small, separated by a diameter or less; elytral punctures larger than on pronotum, separated by up to twice a diameter; prosternal, mesosternal punctures fine, barely visible; metasternal punc- tures small in median 1/3, separated by a diameter or less, punctures absent in lateral 1/3; abdominal ventrites 1, 2 with fine punctures separated by up to three times a diameter medially, ventrites 2–4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head with frons widened from vertex to clypeal apex, slightly wider than eye, clypeal apex arcuate (
<figureCitation id="13442A27FF9A6A17FBC2FCEAFBE9FC0E" box="[1032,1119,873,897]" captionStart="Figures 54–68" captionStartId="24.[192,287,1678,1700]" captionTargetBox="[210,1444,233,1610]" captionTargetId="figure-97@24.[192,1445,217,1656]" captionTargetPageId="24" captionText="Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule." figureDoi="http://doi.org/10.5281/zenodo.5353556" httpUri="https://zenodo.org/record/5353556/files/figure.png" pageId="13" pageNumber="12">Fig. 55</figureCitation>
); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 2/3 distance to rear margin of ventrite. Genitalia with penis guide 3/4 as long as paramere, slender, slightly oval, tapered to abrupt apex in apical 1/4; paramere slender, weakly curved, apex rounded (
<figureCitation id="13442A27FF9A6A17FDFAFB89FD07FBAD" box="[560,689,1034,1058]" captionStart="Figures 54–68" captionStartId="24.[192,287,1678,1700]" captionTargetBox="[210,1444,233,1610]" captionTargetId="figure-97@24.[192,1445,217,1656]" captionTargetPageId="24" captionText="Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule." figureDoi="http://doi.org/10.5281/zenodo.5353556" httpUri="https://zenodo.org/record/5353556/files/figure.png" pageId="13" pageNumber="12">Fig. 56, 57</figureCitation>
); penis slender, apex attenuate, basal capsule wide, flat, inner arm straight, slender, apically truncated, outer arm wide, apically rounded (
<figureCitation id="13442A27FF9A6A17FBADFBA9FB09FBCD" box="[1127,1215,1066,1090]" captionStart="Figures 54–68" captionStartId="24.[192,287,1678,1700]" captionTargetBox="[210,1444,233,1610]" captionTargetId="figure-97@24.[192,1445,217,1656]" captionTargetPageId="24" captionText="Figures 54–68. Calloeneis spp. 54–58) Calloeneis johnnie. 54) Habitus. 55) Frons. 56) Penis guide ventral. 57) Penis guide lateral. 58) Penis. 59–63) Calloeneis robyn. 59) Habitus. 60) Frons. 61) Penis guide ventral. 62) Penis guide lateral.63) Penis.64–68) Calloeneis francis. 64) Habitus.65) Frons.66) Penis guide ventral.67) Penis guide lateral. 68) Spermathecal capsule." figureDoi="http://doi.org/10.5281/zenodo.5353556" httpUri="https://zenodo.org/record/5353556/files/figure.png" pageId="13" pageNumber="12">Fig. 58</figureCitation>
).
</paragraph>
<paragraph id="8BC036A2FF9A6A17FF0AFBD8FC5AFB1D" blockId="13.[192,1440,1115,1171]" pageId="13" pageNumber="12">
<emphasis id="B90BEAB0FF9A6A17FF0AFBD8FE9AFBFC" bold="true" box="[192,300,1115,1139]" pageId="13" pageNumber="12">Female.</emphasis>
Similar to male except head reddish brown, pronotum entirely dark brown. Genitalia with ramus of spermathecal capsule bulbous, cornu curved, apex acute.
</paragraph>
<paragraph id="8BC036A2FF9A6A17FF0AFB28FCC0FB6C" blockId="13.[192,1440,1195,1251]" pageId="13" pageNumber="12">
<emphasis id="B90BEAB0FF9A6A17FF0AFB28FEFAFB4C" bold="true" box="[192,332,1195,1219]" pageId="13" pageNumber="12">Variation.</emphasis>
Length
<quantity id="4C879B47FF9A6A17FE7DFB28FDD1FB4D" box="[439,615,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="13" pageNumber="12" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7 to 1.8 mm</quantity>
, Size of median reddish-brown area on elytra varies from wide to almost entirely absent, rendering elytra nearly all black.
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9A6A17FF0AFB78FDB4FADB" pageId="13" pageNumber="12" type="materials_examined">
<paragraph id="8BC036A2FF9A6A17FF0AFB78FDB4FADB" blockId="13.[192,1440,1275,1364]" pageId="13" pageNumber="12">
<emphasis id="B90BEAB0FF9A6A17FF0AFB78FE3DFA9C" bold="true" box="[192,395,1275,1299]" pageId="13" pageNumber="12">Type material.</emphasis>
<materialsCitation id="3B173CFFFF9A6A17FE5FFB78FB05FABC" collectingDate="1966-10" collectionCode="BMNH" collectorName="Pres &amp; Comm Inst Ent BM" country="Trinidad and Tobago" location="Brigand Hill" municipality="Pred. on Pseudaulacaspis" pageId="13" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C48800FF9A6A17FE5FFB78FDB5FA9C" box="[405,515,1275,1299]" pageId="13" pageNumber="12" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F3687632FF9A6A17FD93FB78FD5FFA9C" box="[601,745,1275,1299]" name="Trinidad and Tobago" pageId="13" pageNumber="12">TRINIDAD</collectingCountry>
,
<location id="8EA06079FF9A6A17FD3DFB78FC20FA9C" LSID="urn:lsid:plazi:treatment:03D687B4FF9A6A17FF0AFE7BFA9AFA2B:8EA06079FF9A6A17FD3DFB78FC20FA9C" box="[759,918,1275,1299]" country="Trinidad and Tobago" municipality="Pred. on Pseudaulacaspis" name="Brigand Hill" pageId="13" pageNumber="12">Brigand Hill</location>
,
<date id="FFC11062FF9A6A17FC6EFB78FC4AFA9C" box="[932,1020,1275,1299]" pageId="13" pageNumber="12" value="1966-10">
<collectingDate id="EF85E98AFF9A6A17FC6EFB78FC4AFA9C" box="[932,1020,1275,1299]" pageId="13" pageNumber="12" value="1966-10">X.1966</collectingDate>
</date>
, C.I. E. A. 1306, C.I.B.C., 66/684,
<collectingMunicipality id="6BA4ACD8FF9A6A17FF0AFA98FE4FFABC" box="[192,505,1307,1331]" pageId="13" pageNumber="12">Pred. on Pseudaulacaspis</collectingMunicipality>
pentagona,
<collectorName id="268A5374FF9A6A17FD5AFA9FFD71FABC" box="[656,711,1308,1331]" pageId="13" pageNumber="12">Pres</collectorName>
by
<collectorName id="268A5374FF9A6A17FD39FA98FC52FABC" box="[755,996,1307,1331]" pageId="13" pageNumber="12">Comm Inst Ent BM</collectorName>
1971-1 (
<collectionCode id="ED6EAE67FF9A6A17FB84FA98FB1DFABC" box="[1102,1195,1307,1331]" pageId="13" pageNumber="12">BMNH</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54C48800FF9A6A17FB74FA9FFA8CFABB" box="[1214,1338,1308,1332]" pageId="13" pageNumber="12" type="paratype">Paratypes</typeStatus>
5, same data as
<typeStatus id="54C48800FF9A6A17FEEBFAB8FE3FFADC" box="[289,393,1339,1363]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED6EAE67FF9A6A17FE52FABFFE43FADB" box="[408,501,1340,1364]" pageId="13" pageNumber="12">BMNH</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF9A6A17FF0AFAEFFA9AFA2B" pageId="13" pageNumber="12" type="discussion">
<paragraph id="8BC036A2FF9A6A17FF0AFAEFFA9AFA2B" blockId="13.[192,1442,1388,1444]" pageId="13" pageNumber="12">
<emphasis id="B90BEAB0FF9A6A17FF0AFAEFFEF5FA0B" bold="true" box="[192,323,1388,1412]" pageId="13" pageNumber="12">Remarks.</emphasis>
<taxonomicName id="4C7F4D21FF9A6A17FE86FAEFFD98FA0B" authorityName="Gordon and Hanley" authorityYear="2020" box="[332,558,1388,1412]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="johnnie">
<emphasis id="B90BEAB0FF9A6A17FE86FAEFFD98FA0B" box="[332,558,1388,1412]" italics="true" pageId="13" pageNumber="12">Calloeneis johnnie</emphasis>
</taxonomicName>
does not have a distinctive dorsal color pattern, but the head widened from base to the wide, apically rounded clypeus is thus far unknown elsewhere in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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