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<document id="39D6AE1755BFB6E472F37BDE5316B52B" ID-DOI="10.5281/zenodo.3699463" ID-GBIF-Dataset="8a1b577e-5fce-489a-9777-32c9564a26b2" ID-ISSN="1942-1354" ID-Zenodo-Dep="3699463" ID-ZooBank="938220AB-7950-4850-88DC-3C3D8AB72C89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583500003806" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="DB4387E1713FFFAFFF4109190FCDF911" docLanguage="en" docName="InsectaMundi.0624.1-13.pdf.imf" docOrigin="Insecta Mundi 2018 (624)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nigidius obesus Parry 1864" docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="277AFF997137FFA7FF810A620E3FFFA9" masterDocTitle="Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="7" updateTime="1698725587556" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="30084E0FF472EB746ED230A98A98BAFC">Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
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<mods:namePart id="42ED2DB8DB537926680539868B9CA5BF">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="A8DC1D7ECAE954048885987F004E8BC6">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:title id="BDE4F9CFF011072F6361BDD57D56EBE0">Insecta Mundi</mods:title>
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<mods:date id="D5E826F27B3FB08A58764F40BCCA450C">2018</mods:date>
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<mods:number id="AC77B1733625BA7677ECEC36D5F88C04">2018-04-27</mods:number>
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<mods:number id="B9A53C73B7E1D4377D665C74A5B8AF01">2018</mods:number>
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<treatment id="DB4387E1713FFFAFFF4109190FCDF911" ID-DOI="http://doi.org/10.5281/zenodo.3704542" ID-GBIF-Taxon="162463114" ID-Zenodo-Dep="3704542" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFAFFF4109190FCDF911" httpUri="http://treatment.plazi.org/id/DB4387E1713FFFAFFF4109190FCDF911" lastPageNumber="7" pageId="8" pageNumber="7">
<subSubSection id="1BF0657C713FFFAFFF4109190C5FFC3D" box="[192,608,891,916]" pageId="8" pageNumber="7" type="nomenclature">
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<heading id="081D819B713FFFAFFF4109190C5FFC3D" bold="true" box="[192,608,891,916]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<treatmentCitationGroup id="73FA11D9713FFFAFFF4109190C5FFC3D" box="[192,608,891,916]" pageId="8" pageNumber="7">
<treatmentCitation id="D24B10E6713FFFAFFF4109190C5FFC3D" author="Parry" box="[192,608,891,916]" page="63" pageId="8" pageNumber="7" year="1864">
<taxonomicName id="94EA4D74713FFFAFFF4109190C5FFC3D" ID-CoL="6RY7P" authority="Parry 1864: 63" authorityName="Parry" authorityPageNumber="63" authorityYear="1864" box="[192,608,891,916]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="619EEAE5713FFFAFFF4109190C5FFC3D" bold="true" box="[192,608,891,916]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF4109190FA8FC3A" bold="true" box="[192,407,891,915]" italics="true" pageId="8" pageNumber="7">Nigidius obesus</emphasis>
<bibRefCitation id="377B4B06713FFFAFFE1F091E0C5FFC3D" author="Parry" box="[414,608,892,916]" firstAuthor="Parry" pageId="8" pageNumber="7" pagination="1 - 113" refId="ref6956" refString="Parry, F. J. S. 1864. A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, Third Series 2: 1 - 113." type="book chapter" year="1864">Parry 1864: 63</bibRefCitation>
</emphasis>
</taxonomicName>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713FFFAFFF4109DF0B28FB03" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="535536F7713FFFAFFF4109DF0A46FBF1" blockId="8.[192,1447,956,1113]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF4109DF0FB3FC7D" bold="true" box="[192,396,956,981]" pageId="8" pageNumber="7">Type material.</emphasis>
<materialsCitation id="E3823CAA713FFFAFFE1709DF0CA1FC5F" ID-GBIF-Occurrence="3878839304" collectionCode="NHM" country="Malaysia" location="Penang" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<typeStatus id="8C518855713FFFAFFE1709DF0C2FFC7C" box="[406,528,957,981]" pageId="8" pageNumber="7" type="lectotype">Lectotype</typeStatus>
,
<emphasis id="619EEAE5713FFFAFFD9E09DF0CC0FC7D" bold="true" box="[543,767,956,981]" pageId="8" pageNumber="7">here designated</emphasis>
,
<specimenCount id="45ECFD7E713FFFAFFC9109DF0D73FC7C" box="[784,844,957,981]" pageId="8" pageNumber="7" type="male">male</specimenCount>
(
<collectionCode id="35FBAE32713FFFAFFCDF09DF0D99FC7C" box="[862,934,957,981]" pageId="8" pageNumber="7">NHM</collectionCode>
;
<figureCitation id="CBD12A72713FFFAFFC3409DF0DC1FC7C" box="[949,1022,957,981]" captionStart="Figure 2" captionStartId="4.[192,275,1419,1441]" captionTargetBox="[228,1407,238,1367]" captionTargetId="figure@4.[213,1415,221,1373]" captionTargetPageId="4" captionText="Figure 2. Nigidius obesus Parry, lectotype male, with labels (inset)." figureDoi="http://doi.org/10.5281/zenodo.3699467" httpUri="https://zenodo.org/record/3699467/files/figure.png" pageId="8" pageNumber="7">Fig. 2</figureCitation>
) labeled: a) “85/28; yellow paper, handwritten “
<taxonomicName id="94EA4D74713FFFAFFEEA09BF0F85FC5C" authorityName="Parry" authorityYear="1864" box="[363,442,989,1013]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="619EEAE5713FFFAFFEEA09BF0F85FC5C" box="[363,442,989,1013]" italics="true" pageId="8" pageNumber="7">obesus</emphasis>
</taxonomicName>
/ Parry / Penang”
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFD2A09BC0DF1FC5F" ID-GBIF-Occurrence="3878839302" box="[683,974,990,1014]" collectionCode="BMNH" country="Malaysia" location="Penang" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
b) “
<collectionCode id="35FBAE32713FFFAFFD5509BC0D10FC5F" box="[724,815,990,1014]" pageId="8" pageNumber="7">BMNH</collectionCode>
(E) / 607172”
</materialsCitation>
.
<materialsCitation id="E3823CAA713FFFAFFC5809BC0D3BFBBE" ID-GBIF-Occurrence="2574936459" collectionCode="NHM" pageId="8" pageNumber="7" specimenCount="2" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="8C518855713FFFAFFC5809BC0ABDFC5F" box="[985,1154,990,1014]" pageId="8" pageNumber="7" type="paralectotype">Paralectotype</typeStatus>
<specimenCount id="45ECFD7E713FFFAFFB0809BC0AE4FC5F" box="[1161,1243,990,1014]" pageId="8" pageNumber="7" type="female">female</specimenCount>
(
<collectionCode id="35FBAE32713FFFAFFB6A09BC0B0DFC5F" box="[1259,1330,990,1014]" pageId="8" pageNumber="7">NHM</collectionCode>
) labeled: a) red-bordered circular “Type”; b) “TYPE /
<specimenCount id="45ECFD7E713FFFAFFD4F099D0CC7FBBE" box="[718,760,1023,1047]" pageId="8" pageNumber="7" type="generic">SP.</specimenCount>
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFC90099D0D36FB9E" ID-GBIF-Occurrence="2574936465" collectorName="Tr. Ent. Soc." country="Cambodia" location="Cambodia" pageId="8" pageNumber="7" specimenCount="1" typeStatus="paralectotype">
c) yellow paper, handwritten “
<emphasis id="619EEAE5713FFFAFFB01099D0AC5FBBF" box="[1152,1274,1022,1046]" italics="true" pageId="8" pageNumber="7">
<taxonomicName id="94EA4D74713FFFAFFB01099D0AC9FBBF" authorityName="Parry" authorityYear="1864" box="[1152,1270,1022,1046]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obesus">N. obesus</taxonomicName>
.
</emphasis>
/ Parry, Type / Westw. fig. 5 /
<collectorName id="FE1F5321713FFFAFFE090E420C1BFB91" box="[392,548,1056,1080]" pageId="8" pageNumber="7">Tr. Ent. Soc.</collectorName>
1874 /
<collectingCountry id="2BFD7667713FFFAFFD010E7D0CC0FB9E" box="[640,767,1055,1079]" name="Cambodia" pageId="8" pageNumber="7">Cambodia</collectingCountry>
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFC970E7D0F99FBF1" ID-GBIF-Occurrence="2574936475" collectorName="Tr. Ent. Soc." country="Cambodia" pageId="8" pageNumber="7" specimenCount="1" typeStatus="type">
d) yellow paper, handwritten “Type
<specimenCount id="45ECFD7E713FFFAFFB550E420AC9FB9E" box="[1236,1270,1056,1079]" pageId="8" pageNumber="7" type="generic">sp.</specimenCount>
vid /
<collectorName id="FE1F5321713FFFAFFABC0E420ECEFBF0" pageId="8" pageNumber="7">Tr. Ent. Soc.</collectorName>
73”; e) “85/28”
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFE330E220CF0FBF1" ID-GBIF-Occurrence="3878839303" box="[434,719,1088,1113]" country="Malaysia" location="Penang" pageId="8" pageNumber="7" specimenCount="1" stateProvince="Pulau Pinang" typeStatus="paralectotype">f) “BMNH(E) / 607171”</materialsCitation>
. Type locality: “
<collectingRegion id="912EF815713FFFAFFC160E230DCAFBF0" box="[919,1013,1089,1113]" country="Malaysia" name="Pulau Pinang" pageId="8" pageNumber="7">Penang</collectingRegion>
,
<collectingCountry id="2BFD7667713FFFAFFB830E220A54FBF1" box="[1026,1131,1088,1112]" name="Malaysia" pageId="8" pageNumber="7">Malacca</collectingCountry>
”.
</paragraph>
<paragraph id="535536F7713FFFAFFF410E130B28FB03" blockId="8.[192,1440,1137,1194]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF410E130CEDFB20" bold="true" box="[192,722,1137,1161]" pageId="8" pageNumber="7">
Additional material examined (
<figureCitation id="CBD12A72713FFFAFFDE60E130CFCFB20" box="[615,707,1137,1161]" captionStart="Figure 14" captionStartId="11.[192,275,1877,1899]" captionTargetBox="[236,1395,288,1824]" captionTargetId="figure@11.[236,1395,288,1824]" captionTargetPageId="11" captionText="Figure 14. Distribution of obesus group species: N. obesus (triangles), N. dawnae (squares), N. gravelyi Paulsen, n. sp. (diamonds), N. helleri (circles)." figureDoi="http://doi.org/10.5281/zenodo.3699479" httpUri="https://zenodo.org/record/3699479/files/figure.png" pageId="8" pageNumber="7">Fig. 14</figureCitation>
).
</emphasis>
<materialsCitation id="E3823CAA713FFFAFFD590E130BA3FB20" ID-GBIF-Occurrence="3878839301" box="[728,1436,1137,1162]" collectionCode="NHM, ZMHB, MJPC" country="Malaysia" location="Changkat Menteri" municipality="Batang Padang Jor Camp" pageId="8" pageNumber="7" specimenCount="12" stateProvince="PERAK">
<emphasis id="619EEAE5713FFFAFFD590E130D46FB20" bold="true" box="[728,889,1137,1161]" pageId="8" pageNumber="7">
<collectingCountry id="2BFD7667713FFFAFFD590E130D4BFB20" box="[728,884,1137,1161]" name="Malaysia" pageId="8" pageNumber="7">MALAYSIA</collectingCountry>
:
</emphasis>
<collectingRegion id="912EF815713FFFAFFCFF0E100DDDFB23" box="[894,994,1138,1162]" country="Malaysia" name="Perak" pageId="8" pageNumber="7">PERAK</collectingRegion>
:
<collectingMunicipality id="B331AC8D713FFFAFFC6A0E100B1DFB20" box="[1003,1314,1137,1162]" pageId="8" pageNumber="7">Batang Padang Jor Camp</collectingMunicipality>
(
<specimenCount id="45ECFD7E713FFFAFFAB20E130B7FFB20" box="[1331,1344,1137,1161]" pageId="8" pageNumber="7" type="generic">1</specimenCount>
,
<collectionCode id="35FBAE32713FFFAFFACA0E130BACFB20" box="[1355,1427,1137,1161]" pageId="8" pageNumber="7">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFF410EF00C1EFB03" ID-GBIF-Occurrence="3878839306" box="[192,545,1170,1194]" collectionCode="NHM, ZMHB, MJPC" country="Malaysia" location="Changkat Menteri" municipality="Batang Padang Jor Camp" pageId="8" pageNumber="7" specimenCount="12" stateProvince="PERAK">
<location id="5635602C713FFFAFFF410EF00F9CFB03" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFAFFF4109190FCDF911:5635602C713FFFAFFF410EF00F9CFB03" box="[192,419,1170,1194]" country="Malaysia" municipality="Batang Padang Jor Camp" name="Changkat Menteri" pageId="8" pageNumber="7" stateProvince="PERAK">Changkat Menteri</location>
(
<specimenCount id="45ECFD7E713FFFAFFE350EF00FFBFB03" box="[436,452,1170,1194]" pageId="8" pageNumber="7" type="generic">5</specimenCount>
,
<collectionCode id="35FBAE32713FFFAFFE510EF00C27FB03" box="[464,536,1170,1194]" pageId="8" pageNumber="7">NHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="E3823CAA713FFFAFFDAC0EF00D3CFB03" ID-GBIF-Occurrence="3878839308" box="[557,771,1170,1194]" collectionCode="NHM, ZMHB, MJPC" country="Malaysia" location="Changkat Menteri" municipality="Batang Padang Jor Camp" pageId="8" pageNumber="7" specimenCount="12" stateProvince="PERAK">
<location id="5635602C713FFFAFFDAC0EF00C49FB03" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFAFFF4109190FCDF911:5635602C713FFFAFFDAC0EF00C49FB03" box="[557,630,1170,1194]" country="Malaysia" municipality="Batang Padang Jor Camp" name="Perak" pageId="8" pageNumber="7" stateProvince="PERAK">Perak</location>
(
<specimenCount id="45ECFD7E713FFFAFFD060EF00CA8FB03" box="[647,663,1170,1194]" pageId="8" pageNumber="7" type="generic">5</specimenCount>
,
<collectionCode id="35FBAE32713FFFAFFD250EF00CC4FB03" box="[676,763,1170,1194]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="8" pageNumber="7">ZMHB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E3823CAA713FFFAFFC8F0EF00B2DFB03" ID-GBIF-Occurrence="3878839307" box="[782,1298,1170,1194]" collectionCode="NHM, ZMHB, MJPC" country="Malaysia" location="Changkat Menteri" municipality="Batang Padang Jor Camp" pageId="8" pageNumber="7" specimenCount="12" stateProvince="PERAK">
<collectingRegion id="912EF815713FFFAFFC8F0EF00DB4FB03" box="[782,907,1170,1194]" country="Malaysia" name="Pahang" pageId="8" pageNumber="7">PAHANG</collectingRegion>
:
<location id="5635602C713FFFAFFC140EF00AB3FB03" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFAFFF4109190FCDF911:5635602C713FFFAFFC140EF00AB3FB03" box="[917,1164,1170,1194]" country="Malaysia" municipality="Batang Padang Jor Camp" name="Cameron Highlands" pageId="8" pageNumber="7" stateProvince="PERAK">Cameron Highlands</location>
(
<specimenCount id="45ECFD7E713FFFAFFB1E0EF00A93FB03" box="[1183,1196,1170,1194]" pageId="8" pageNumber="7" type="generic">1</specimenCount>
,
<collectionCode id="35FBAE32713FFFAFFB380EF00B35FB03" box="[1209,1290,1170,1194]" collectionName="MJPC" pageId="8" pageNumber="7">MJPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713FFFAFFF410EA10A4BFA94" pageId="8" pageNumber="7" type="diagnosis">
<paragraph id="535536F7713FFFAFFF410EA10A4BFA94" blockId="8.[192,1440,1219,1342]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF410EA10F70FB72" bold="true" box="[192,335,1219,1243]" pageId="8" pageNumber="7">Diagnosis.</emphasis>
This species can be distinguished from the insular species by the narrower, more convex and less punctate elytral intervals, as well as by details of the male genitalia, which has a more sinuous flagellum with a different apex (
<figureCitation id="CBD12A72713FFFAFFDCD0F670C9BFAB4" box="[588,676,1285,1309]" captionStart="Figures 1013" captionStartId="9.[192,287,1578,1600]" captionTargetBox="[221,1399,218,1550]" captionTargetPageId="9" captionText="Figures 1013. Male genitalia of obesus group species. 10) N. obesus. 11) N. dawnae, arrow indicates small apical lobe. 12) N. gravelyi Paulsen, n. sp. 13) N. helleri, arrow indicates large lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3699477" httpUri="https://zenodo.org/record/3699477/files/figure.png" pageId="8" pageNumber="7">Fig. 10</figureCitation>
). It differs from
<taxonomicName id="94EA4D74713FFFAFFCEF0F670DCEFAB4" box="[878,1009,1285,1309]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713FFFAFFCEF0F670DCEFAB4" box="[878,1009,1285,1309]" italics="true" pageId="8" pageNumber="7">N. dawnae</emphasis>
</taxonomicName>
in the medially impunctate elytral intervals and more oval (less lenticular) punctures laterally on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713FFFAFFF410F350FCDF911" pageId="8" pageNumber="7" type="discussion">
<paragraph id="535536F7713FFFAFFF410F350A02F9DF" blockId="8.[192,1441,1367,1720]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF410F350F7CFAC6" bold="true" box="[192,323,1367,1391]" pageId="8" pageNumber="7">Remarks.</emphasis>
The number of specimens examined by
<bibRefCitation id="377B4B06713FFFAFFCBE0F350DE4FAC6" author="Parry" box="[831,987,1367,1391]" firstAuthor="Parry" pageId="8" pageNumber="7" pagination="1 - 113" refId="ref6956" refString="Parry, F. J. S. 1864. A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, Third Series 2: 1 - 113." type="book chapter" year="1864">Parry (1864)</bibRefCitation>
is not indicated. The label 85/28 on both putative type specimens indicates accession into the NHM from the auction of specimens from the Parry collection via Mr. Janson. The
<typeStatus id="8C518855713FFFAFFD030FFB0CCFFA18" box="[642,752,1433,1457]" pageId="8" pageNumber="7" type="lectotype">lectotype</typeStatus>
male is appropriately labeled as being from Penang.The female has been labeled “
<collectingCountry id="2BFD7667713FFFAFFE720FD80C50FA7B" box="[499,623,1466,1490]" name="Cambodia" pageId="8" pageNumber="7">Cambodia</collectingCountry>
” in the same style, but this locality is not mentioned in the original description. A secondary label on the female specimen alludes to it being a type described in 1873. The species described by
<bibRefCitation id="377B4B06713FFFAFFE3D0F9E0C69F9BD" author="Parry" box="[444,598,1532,1556]" firstAuthor="Parry" pageId="8" pageNumber="7" pagination="335 - 345" refId="ref6999" refString="Parry, F. J. S. 1873. Characters of seven nondescript Lucanoid Coleoptera, and remarks upon the genus Lissotes, Nigidius and Figulus. Transactions of the Royal Entomological Society of London 1873 (3): 335 - 345." type="journal article" year="1873">Parry (1873)</bibRefCitation>
from
<collectingCountry id="2BFD7667713FFFAFFD1F0F990D25F9BA" box="[670,794,1531,1555]" name="Cambodia" pageId="8" pageNumber="7">Cambodia</collectingCountry>
is
<taxonomicName id="94EA4D74713FFFAFFCBC0F9E0A16F9BD" authority="Parry" authorityName="Parry" authorityYear="1873" box="[829,1065,1531,1556]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="distinctus">
<emphasis id="619EEAE5713FFFAFFCBC0F9E0DE6F9BA" box="[829,985,1531,1555]" italics="true" pageId="8" pageNumber="7">N. distinctus</emphasis>
Parry
</taxonomicName>
, from a different species group with a drastically different form of ocular canthus. It is clear from the description of the emarginate lateral margin of the head with two distinct acute angles that the female in question is not
<taxonomicName id="94EA4D74713FFFAFFAA50C5C0BA7F9FC" authorityName="Parry" authorityYear="1864" box="[1316,1432,1597,1621]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="619EEAE5713FFFAFFAA50C5C0BA7F9FC" box="[1316,1432,1597,1621]" italics="true" pageId="8" pageNumber="7">N. obesus</emphasis>
</taxonomicName>
. Thus, the attribution of the species to
<collectingCountry id="2BFD7667713FFFAFFD120C3C0D30F9DF" box="[659,783,1630,1654]" name="Cambodia" pageId="8" pageNumber="7">Cambodia</collectingCountry>
is considered erroneous.
</paragraph>
<paragraph id="535536F7713FFFAFFF6C0C1D0FCDF911" blockId="8.[192,1441,1367,1720]" pageId="8" pageNumber="7">
Specimens studied were from March, October, and September. In general, adults of
<taxonomicName id="94EA4D74713FFFAFFB5B0C1D0B7BF93E" box="[1242,1348,1663,1687]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713FFFAFFB5B0C1D0B7BF93E" box="[1242,1348,1663,1687]" italics="true" pageId="8" pageNumber="7">Nigidius</emphasis>
</taxonomicName>
species can be found year-round.
</paragraph>
</subSubSection>
</treatment>
</document>