360 lines
44 KiB
XML
360 lines
44 KiB
XML
<document id="5A5B328DA34BF5216B4CE9611BF7DB8C" ID-DOI="10.5281/zenodo.3699463" ID-GBIF-Dataset="8a1b577e-5fce-489a-9777-32c9564a26b2" ID-ISSN="1942-1354" ID-Zenodo-Dep="3699463" ID-ZooBank="938220AB-7950-4850-88DC-3C3D8AB72C89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583500003806" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="DB4387E17134FFA1FF4108260CD3F820" docLanguage="en" docName="InsectaMundi.0624.1-13.pdf.imf" docOrigin="Insecta Mundi 2018 (624)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nigidius MacLeay 1819" docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="277AFF997137FFA7FF810A620E3FFFA9" masterDocTitle="Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="2" updateTime="1698725587556" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="B55A09A20715A2E0D9D58CC0F5FDF2C2" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1B3109C39DB618A0A92733BC869FE405">
|
||
<mods:title id="F163843258E91DDDAD54F8F6C7594C5D">Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="F6080BB58C5ABD3E213F1D93DD325141" type="personal">
|
||
<mods:role id="BA5C63E4612ED3342DDC43AEF03559AE">
|
||
<mods:roleTerm id="097FE965C2BD2A5C4DFDA07C0D962F70">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B3EFF4294BD3709EE3B7576359AB50C0">Paulsen, M. J.</mods:namePart>
|
||
<mods:affiliation id="1B4FF715714DFC322BAA5579A00B6A77">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
|
||
<mods:nameIdentifier id="E5756AB644004C0A39B8B3480167EB72" type="email">mjpaulsen@unl.edu</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="70D8E2E8A9266B4AD1F3D8B7180A61FC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="FFDA097ED5CEE3BEBFC55D7D6D06DC40" type="host">
|
||
<mods:titleInfo id="138F9A6F2EFDB789C4ADE67C8172D530">
|
||
<mods:title id="19E9E8066D93EE7B57090EEA00544F54">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="6675E0100A0E9FB830C3C22EE70A9F69">
|
||
<mods:date id="681A052381C304AF1CE549BF935610DC">2018</mods:date>
|
||
<mods:detail id="31745156FC2B0E8F8A5FB888AF5AFE71" type="pubDate">
|
||
<mods:number id="B4CA821E34602BA1298D4B90E52C31A3">2018-04-27</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="87B3DF234FD7664C4141CDE81CD25BA3" type="volume">
|
||
<mods:number id="9DE928367905CBA4775F0C21C8C442EB">2018</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2DE3F8CAA97913C28053E9CE6E4DD0CE" type="issue">
|
||
<mods:number id="DD24998C0C84DA4F87EBB9CF28668290">624</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="6578EB06C8A051F92D6510735CE3E1D0" unit="page">
|
||
<mods:start id="1F72CE6ACB0E41BC9BAD649C3D0E906A">1</mods:start>
|
||
<mods:end id="41E133DE666147459E277B89F40C2404">13</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="3129BFFD31FFCDC19180C8E2969D85FA">journal article</mods:classification>
|
||
<mods:identifier id="ECA4CC639DC0E4EDB754A3240120CC64" type="DOI">10.5281/zenodo.3699463</mods:identifier>
|
||
<mods:identifier id="837761631A31283AC2BF6AA9D1E18951" type="GBIF-Dataset">8a1b577e-5fce-489a-9777-32c9564a26b2</mods:identifier>
|
||
<mods:identifier id="7D80A9598B34324F6B0D87BD97235D82" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="50584825F7ECAD4C712372A219693445" type="Zenodo-Dep">3699463</mods:identifier>
|
||
<mods:identifier id="4B76F3C4103FEC88CB6DCE5F63E9EE32" type="ZooBank">938220AB-7950-4850-88DC-3C3D8AB72C89</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="DB4387E17134FFA1FF4108260CD3F820" ID-DOI="http://doi.org/10.5281/zenodo.3704534" ID-GBIF-Taxon="162463112" ID-Zenodo-Dep="3704534" LSID="urn:lsid:plazi:treatment:DB4387E17134FFA1FF4108260CD3F820" httpUri="http://treatment.plazi.org/id/DB4387E17134FFA1FF4108260CD3F820" lastPageId="6" lastPageNumber="5" pageId="3" pageNumber="2">
|
||
<subSubSection id="1BF0657C7134FFA4FF4108260FDDFDF5" box="[192,482,579,604]" pageId="3" pageNumber="2" type="nomenclature">
|
||
<paragraph id="535536F77134FFA4FF4108260FDDFDF5" blockId="3.[192,482,579,604]" box="[192,482,579,604]" pageId="3" pageNumber="2">
|
||
<heading id="081D819B7134FFA4FF4108260FDDFDF5" bold="true" box="[192,482,579,604]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
|
||
<emphasis id="619EEAE57134FFA4FF4108260FDDFDF5" bold="true" box="[192,482,579,604]" pageId="3" pageNumber="2">
|
||
<taxonomicName id="94EA4D747134FFA4FF4108260F22FDF5" ID-CoL="643G" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[192,285,580,604]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">obesus</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1BF0657C7134FFA4FF6F08160FD3FB25" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="535536F77134FFA4FF6F08160ABEFC05" blockId="3.[192,1440,628,1164]" pageId="3" pageNumber="2">
|
||
Identification of species within the obesus group has been complicated by erroneous species hypotheses and synonymies. Members of this species group can be readily separated from other Southeast Asian
|
||
<taxonomicName id="94EA4D747134FFA4FE9108D10F45FD62" box="[272,378,691,715]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="619EEAE57134FFA4FE9108D10F45FD62" box="[272,378,691,715]" italics="true" pageId="3" pageNumber="2">Nigidius</emphasis>
|
||
</taxonomicName>
|
||
species by the presence of large, sub-oval lobes of the ocular canthus adjacent to the eyes (
|
||
<figureCitation id="CBD12A727134FFA4FE8408B60F75FD45" box="[261,330,724,748]" captionStart="Figure 1" captionStartId="3.[192,275,1877,1899]" captionTargetBox="[227,1426,1214,1817]" captionTargetId="graphics@3.[227,827,1214,1766]" captionTargetPageId="3" captionText="Figure 1. Species groups of Southeast Asian Nigidius species based on ocular canthus shape. 1a) Nigidius obesus group. 1b) cornutus group. 1c) distinctus group. 1d) laevicollis group." figureDoi="http://doi.org/10.5281/zenodo.3699465" httpUri="https://zenodo.org/record/3699465/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
|
||
). This is in contrast to other Southeast Asian groups that can be generalized as follows: b) the cornutus group, including
|
||
<taxonomicName id="94EA4D747134FFA4FDB208960D03FCA5" authority="MacLeay" authorityName="MacLeay" authorityYear="1819" box="[563,828,756,780]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="cornutus">
|
||
<emphasis id="619EEAE57134FFA4FDB208960CFEFCA2" box="[563,705,756,779]" italics="true" pageId="3" pageNumber="2">N. cornutus</emphasis>
|
||
MacLeay
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="94EA4D747134FFA4FCC608960A71FCA5" authority="Okuda" authorityName="Okuda" authorityYear="2002" box="[839,1102,755,780]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="619EEAE57134FFA4FCC608960DCEFCA2" box="[839,1009,755,779]" italics="true" pageId="3" pageNumber="2">N. sabahensis</emphasis>
|
||
Okuda
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="94EA4D747134FFA4FB1108960B9FFCA2" authority="Van Roon" authorityName="Van Roon" authorityYear="1914" box="[1168,1440,755,779]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="oblongus">
|
||
<emphasis id="619EEAE57134FFA4FB1108960B1FFCA2" box="[1168,1312,755,779]" italics="true" pageId="3" pageNumber="2">N. oblongus</emphasis>
|
||
Van Roon
|
||
</taxonomicName>
|
||
with a shelf-like canthus that is truncate anteriorly; c) the distinctus group, including
|
||
<taxonomicName id="94EA4D747134FFA4FA8009760F39FCE5" authority="Parry" authorityName="Parry" authorityYear="1873" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="distinctus">
|
||
<emphasis id="619EEAE57134FFA4FA8009760B9FFC82" box="[1281,1440,787,811]" italics="true" pageId="3" pageNumber="2">N. distinctus</emphasis>
|
||
Parry
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="94EA4D747134FFA4FEC509560C2AFCE5" authority="Boileau" authorityName="Boileau" authorityYear="1905" box="[324,533,819,844]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lewisi">
|
||
<emphasis id="619EEAE57134FFA4FEC509560F8FFCE2" box="[324,432,819,843]" italics="true" pageId="3" pageNumber="2">N. lewisi</emphasis>
|
||
Boileau
|
||
</taxonomicName>
|
||
with a deeply emarginate canthus; and d) the laevicollis group, including numerous species such as
|
||
<taxonomicName id="94EA4D747134FFA4FD9109360D04FCC5" authority="Westwood" authorityName="Westwood" box="[528,827,851,876]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="laevicollis">
|
||
<emphasis id="619EEAE57134FFA4FD9109360C8EFCC2" box="[528,689,851,875]" italics="true" pageId="3" pageNumber="2">N. laevicollis</emphasis>
|
||
Westwood
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="94EA4D747134FFA4FCCB09360A6EFCC5" authority="Boileau" authorityName="Boileau" authorityYear="1902" box="[842,1105,851,876]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="elongatus">
|
||
<emphasis id="619EEAE57134FFA4FCCB09360DD8FCC2" box="[842,999,851,875]" italics="true" pageId="3" pageNumber="2">N. elongatus</emphasis>
|
||
Boileau
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="94EA4D747134FFA4FB1B09360B9FFCC2" authority="Bates" authorityName="Bates" authorityYear="1866" box="[1178,1440,852,876]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="formosanus">
|
||
<emphasis id="619EEAE57134FFA4FB1B09360B6FFCC5" box="[1178,1360,852,876]" italics="true" pageId="3" pageNumber="2">N. formosanus</emphasis>
|
||
Bates
|
||
</taxonomicName>
|
||
with a posteriorly expanding triangular canthus. In some species (such as
|
||
<taxonomicName id="94EA4D747134FFA4FBC609160B47FC25" authority="Ritsema" authorityName="Ritsema" box="[1095,1400,883,908]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="kinabaluenis">
|
||
<emphasis id="619EEAE57134FFA4FBC609160B36FC22" box="[1095,1289,883,907]" italics="true" pageId="3" pageNumber="2">N. kinabaluenis</emphasis>
|
||
Ritsema
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="94EA4D747134FFA4FA0209160FF3FC05" authority="Ritsema" authorityName="Ritsema" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lichtensteinii">
|
||
<emphasis id="619EEAE57134FFA4FA0209160F61FC02" italics="true" pageId="3" pageNumber="2">N. lichtensteinii</emphasis>
|
||
Ritsema
|
||
</taxonomicName>
|
||
) the canthus is weakly emarginate at the anterior third.
|
||
</paragraph>
|
||
<paragraph id="535536F77134FFA4FF6C09D60FD3FB25" blockId="3.[192,1440,628,1164]" pageId="3" pageNumber="2">Within the obesus species group, species can be distinguished by the shape of the flagellum of the male genitalia, pronotal punctation, and the form of the anterolateral projection of the pronotum. Although useful for species diagnosis, the latter character is informative only for larger specimens because in all species the smaller individuals have a less well-developed projection. The form of the mandibles is conserved in these species, and partially sexually dimorphic. Large males can be sexed easily due to the broad lobe-like tooth near the base of the mandible. In females and small males, the tooth is much narrower and sub-acute.</paragraph>
|
||
</subSubSection>
|
||
<caption id="0795667F7134FFA4FF410D370DFDF82E" ID-DOI="http://doi.org/10.5281/zenodo.3699465" ID-Zenodo-Dep="3699465" httpUri="https://zenodo.org/record/3699465/files/figure.png" pageId="3" pageNumber="2" startId="3.[192,275,1877,1899]" targetBox="[227,1426,1214,1817]" targetPageId="3">
|
||
<paragraph id="535536F77134FFA4FF410D370DFDF82E" blockId="3.[192,1440,1877,1928]" pageId="3" pageNumber="2">
|
||
<emphasis id="619EEAE57134FFA4FF410D370F12F8C3" bold="true" box="[192,301,1877,1899]" pageId="3" pageNumber="2">Figure 1.</emphasis>
|
||
Species groups of Southeast Asian
|
||
<taxonomicName id="94EA4D747134FFA4FD300D370D2EF8C3" box="[689,785,1877,1898]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="619EEAE57134FFA4FD300D370D2EF8C3" box="[689,785,1877,1898]" italics="true" pageId="3" pageNumber="2">Nigidius</emphasis>
|
||
</taxonomicName>
|
||
species based on ocular canthus shape.
|
||
<emphasis id="619EEAE57134FFA4FB470D370AD3F8C2" bold="true" box="[1222,1260,1877,1899]" pageId="3" pageNumber="2">1a)</emphasis>
|
||
<taxonomicName id="94EA4D747134FFA4FB730D370B9FF8C2" authorityName="Parry" authorityYear="1864" box="[1266,1440,1877,1899]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="obesus">Nigidius obesus</taxonomicName>
|
||
group.
|
||
<emphasis id="619EEAE57134FFA4FE8C0D100F0BF821" bold="true" box="[269,308,1906,1928]" pageId="3" pageNumber="2">1b)</emphasis>
|
||
cornutus group.
|
||
<emphasis id="619EEAE57134FFA4FE6E0D100C2CF821" bold="true" box="[495,531,1906,1928]" pageId="3" pageNumber="2">1c)</emphasis>
|
||
distinctus group.
|
||
<emphasis id="619EEAE57134FFA4FD5B0D100D3EF821" bold="true" box="[730,769,1906,1928]" pageId="3" pageNumber="2">1d)</emphasis>
|
||
laevicollis group.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0795667F7133FFA3FF410FE90D8FFA0B" ID-DOI="http://doi.org/10.5281/zenodo.3699467" ID-Zenodo-Dep="3699467" box="[192,944,1419,1442]" httpUri="https://zenodo.org/record/3699467/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,275,1419,1441]" targetBox="[228,1407,238,1367]" targetPageId="4">
|
||
<paragraph id="535536F77133FFA3FF410FE90D8FFA0B" blockId="4.[192,944,1419,1442]" box="[192,944,1419,1442]" pageId="4" pageNumber="3">
|
||
<emphasis id="619EEAE57133FFA3FF410FE90F11FA09" bold="true" box="[192,302,1419,1441]" pageId="4" pageNumber="3">Figure 2.</emphasis>
|
||
<taxonomicName id="94EA4D747133FFA3FEB40FE90C13FA08" authority="Parry" authorityName="Parry" authorityYear="1864" box="[309,556,1419,1441]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57133FFA3FEB40FE90FDCFA09" box="[309,483,1419,1440]" italics="true" pageId="4" pageNumber="3">Nigidius obesus</emphasis>
|
||
Parry
|
||
</taxonomicName>
|
||
, lectotype male, with labels (inset).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1BF0657C7133FFA2FF6C0FB20BB5F9D6" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="535536F77133FFA3FF6C0FB20DC2F880" blockId="4.[192,1440,1488,1929]" pageId="4" pageNumber="3">
|
||
The first species of the group was described from “
|
||
<collectingRegion id="912EF8157133FFA3FCBC0FB30DA5FA40" box="[829,922,1489,1513]" country="Malaysia" name="Pulau Pinang" pageId="4" pageNumber="3">Penang</collectingRegion>
|
||
,
|
||
<collectingCountry id="2BFD76677133FFA3FC250FB20A37FA41" box="[932,1032,1488,1512]" name="Malaysia" pageId="4" pageNumber="3">Malacca</collectingCountry>
|
||
” [West
|
||
<collectingCountry id="2BFD76677133FFA3FBE20FB20AE8FA41" box="[1123,1239,1488,1512]" name="Malaysia" pageId="4" pageNumber="3">Malaysia</collectingCountry>
|
||
], by
|
||
<bibRefCitation id="377B4B067133FFA3FA8B0FB30B9FFA41" author="Parry" box="[1290,1440,1488,1513]" firstAuthor="Parry" pageId="4" pageNumber="3" pagination="1 - 113" refId="ref6956" refString="Parry, F. J. S. 1864. A catalogue of lucanoid Coleoptera; with illustrations and descriptions of various new and interesting species. Transactions of the Entomological Society of London, Third Series 2: 1 - 113." type="book chapter" year="1864">Parry (1864)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="94EA4D747133FFA3FF600F930F6CF9A1" authorityName="Parry" authorityYear="1864" box="[225,339,1520,1544]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57133FFA3FF600F930F6CF9A1" box="[225,339,1520,1544]" italics="true" pageId="4" pageNumber="3">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CBD12A727133FFA3FEE30F920F97F9A0" box="[354,424,1520,1545]" captionStart="Figure 2" captionStartId="4.[192,275,1419,1441]" captionTargetBox="[228,1407,238,1367]" captionTargetId="figure@4.[213,1415,221,1373]" captionTargetPageId="4" captionText="Figure 2. Nigidius obesus Parry, lectotype male, with labels (inset)." figureDoi="http://doi.org/10.5281/zenodo.3699467" httpUri="https://zenodo.org/record/3699467/files/figure.png" pageId="4" pageNumber="3">Fig. 2</figureCitation>
|
||
), and the species was subsequently illustrated by
|
||
<bibRefCitation id="377B4B067133FFA3FB820F920AE9F9A0" author="Westwood" box="[1027,1238,1520,1545]" firstAuthor="Westwood" pageId="4" pageNumber="3" pagination="357 - 363" refId="ref7044" refString="Westwood, J. O. 1874. Illustrations of several additional species of Lucanidae in the collection of Major F. J. Sidney Parry. Transactions of the Entomological Society of London 1874 (3): 357 - 363." type="journal article" year="1874">Westwood (1874)</bibRefCitation>
|
||
. Since that time, most specimens from
|
||
<collectingCountry id="2BFD76677133FFA3FE4B0C720C05F981" box="[458,570,1552,1576]" name="Malaysia" pageId="4" pageNumber="3">Malaysia</collectingCountry>
|
||
and
|
||
<collectingCountry id="2BFD76677133FFA3FDF80C720CCEF981" box="[633,753,1552,1576]" name="Indonesia" pageId="4" pageNumber="3">Indonesia</collectingCountry>
|
||
in this group were identified as
|
||
<taxonomicName id="94EA4D747133FFA3FB010C730ACCF981" authorityName="Parry" authorityYear="1864" box="[1152,1267,1552,1576]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57133FFA3FB010C730ACCF981" box="[1152,1267,1552,1576]" italics="true" pageId="4" pageNumber="3">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="377B4B067133FFA3FA850C730BA3F980" author="Gestro" box="[1284,1436,1553,1577]" firstAuthor="Gestro" pageId="4" pageNumber="3" pagination="303 - 340" refId="ref6741" refString="Gestro, R. 1881. Enumerazione dei lucanidi raccolti nell'Arcipelage Malese e nella Papuasia dai signori G. Doria, O. Beccari e L. M. D'Albertis. Annali del Museo Civico di Storia Naruraledi Genova 16: 303 - 340." type="journal article" year="1881">Gestro 1881</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="377B4B067133FFA3FF410C520F61F9E1" author="Benesh" box="[192,350,1584,1608]" firstAuthor="Benesh" pageId="4" pageNumber="3" pagination="1 - 178" refId="ref6480" refString="Benesh, B. 1960. Lucanidea (sic). Coleopterorum Catalogus, Supplementa (Second Edition) 8: 1 - 178." type="journal article" year="1960">Benesh 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="377B4B067133FFA3FEE80C530C2FF9E0" author="Bomans" box="[361,528,1585,1609]" firstAuthor="Bomans" pageId="4" pageNumber="3" pagination="137 - 152" refId="ref6562" refString="Bomans, H. E. 1991. Inventaire provisoire de Coleopteres Lucanidae de Thailande et description de trois nouvelles especes. (74 eme contribution a l'etude de Coleopteres Lucanides). Lambillionea 91 (2): 137 - 152." type="journal article" year="1991">Bomans 1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="377B4B067133FFA3FD9B0C520D57F9E0" author="Mizunuma and Nagai" box="[538,872,1584,1609]" firstAuthor="Mizunuma" pageId="4" pageNumber="3" refId="ref6882" refString="Mizunuma, T., and S. Nagai. 1994. The Lucanid beetles of the world. Mushi-sha Iconographic series of Insects, Volume 1. H. Fujita Ed.; Tokyo, Japan. 1338 p." type="book" year="1994">Mizunuma and Nagai 1994</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="377B4B067133FFA3FCFB0C530A85F9E0" author="Bomans and Benoit" box="[890,1210,1584,1609]" firstAuthor="Bomans" pageId="4" pageNumber="3" pagination="3 - 106" refId="ref6640" refString="Bomans, H. E., and P. Benoit. 2007. Inventaire provisoire commente des Coleopteres Lucanides d'Indonesie et de Malaisie. Lambillionea 107 Supplement 2: 3 - 106." type="journal article" year="2007">Bomans and Benoit (2007)</bibRefCitation>
|
||
gave an exceptionally wide distribution for
|
||
<emphasis id="619EEAE57133FFA3FE770C330C4EF9C1" box="[502,625,1616,1640]" italics="true" pageId="4" pageNumber="3">
|
||
<taxonomicName id="94EA4D747133FFA3FE770C330C54F9C1" authorityName="Parry" authorityYear="1864" box="[502,619,1616,1640]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">N. obesus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
encompassing the range of all species in the group, as well as
|
||
<collectingRegion id="912EF8157133FFA3FAE40C330B9FF9C1" box="[1381,1440,1617,1640]" country="Indonesia" name="Jawa Timur" pageId="4" pageNumber="3">Java</collectingRegion>
|
||
and Sumba from which I have not seen specimens.
|
||
<bibRefCitation id="377B4B067133FFA3FCBA0C120DE4F920" author="Fujita" box="[827,987,1648,1673]" firstAuthor="Fujita" pageId="4" pageNumber="3" refId="ref6710" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo, Japan. 472 p." type="book" year="2010">Fujita (2010)</bibRefCitation>
|
||
limited
|
||
<taxonomicName id="94EA4D747133FFA3FBC20C130A88F921" authorityName="Parry" authorityYear="1864" box="[1091,1207,1648,1672]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57133FFA3FBC20C130A88F921" box="[1091,1207,1648,1672]" italics="true" pageId="4" pageNumber="3">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
to specimens from
|
||
<collectingCountry id="2BFD76677133FFA3FF410CF20F10F901" box="[192,303,1680,1704]" name="Malaysia" pageId="4" pageNumber="3">Malaysia</collectingCountry>
|
||
and Sumatra and treated the Borneo specimens as an undescribed species. However, specimens from Sumatra and Borneo possess distinct male genitalia from those of
|
||
<emphasis id="619EEAE57133FFA3FBAB0CD30A9AF961" box="[1066,1189,1712,1736]" italics="true" pageId="4" pageNumber="3">
|
||
<taxonomicName id="94EA4D747133FFA3FBAB0CD30A9FF961" authorityName="Parry" authorityYear="1864" box="[1066,1184,1712,1736]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">N. obesus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and specimens from these areas can themselves be separated into two distinct species. The true
|
||
<taxonomicName id="94EA4D747133FFA3FBD20CB30AFAF941" authorityName="Parry" authorityYear="1864" box="[1107,1221,1744,1768]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57133FFA3FBD20CB30AFAF941" box="[1107,1221,1744,1768]" italics="true" pageId="4" pageNumber="3">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
is thus far known only from peninsular
|
||
<collectingCountry id="2BFD76677133FFA3FE450C920C0BF8A1" box="[452,564,1776,1800]" name="Malaysia" pageId="4" pageNumber="3">Malaysia</collectingCountry>
|
||
and shares the same gross morphology of the male genitalia as the other mainland Asian species in the group,
|
||
<taxonomicName id="94EA4D747133FFA3FD0C0D730D46F881" authority="Gravely" authorityName="Gravely" box="[653,889,1808,1832]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="dawnae">
|
||
<emphasis id="619EEAE57133FFA3FD0C0D730D2FF881" box="[653,784,1808,1832]" italics="true" pageId="4" pageNumber="3">N. dawnae</emphasis>
|
||
Gravely
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CBD12A727133FFA3FC080D720DD0F880" box="[905,1007,1808,1833]" captionStart="Figures 3–4" captionStartId="5.[192,287,1506,1528]" captionTargetBox="[215,1371,238,1443]" captionTargetId="figure@5.[203,1383,233,1453]" captionTargetPageId="5" captionText="Figures 3–4. Nigidius dawnae Gravely. 3) Major male, dorsal habitus. 4) Lectotype male with labels." figureDoi="http://doi.org/10.5281/zenodo.3699469" httpUri="https://zenodo.org/record/3699469/files/figure.png" pageId="4" pageNumber="3">Fig. 3–4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="535536F77133FFA2FF720D520BB5F9D6" blockId="4.[192,1440,1488,1929]" lastBlockId="5.[192,1440,1572,1929]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
|
||
Specimens from Borneo and
|
||
<collectingRegion id="912EF8157133FFA3FDE00D530CF2F8E0" box="[609,717,1841,1865]" country="Indonesia" name="Sumatera Utara" pageId="4" pageNumber="3">Sumatra</collectingRegion>
|
||
share similarities in the overall male genitalic form but comprise two distinct species based on obvious differences in the pronotal punctation and the form of the anterolateral margin of the pronotum. In the Sumatran species, the punctation of the pronotum is more uniformly distributed and coarser. The pronota of the Borneo species (and both mainland Asian species) show clear subfoveate areas of coarse punctures juxtaposed with almost impunctate areas with only fine, shallow punctures, and the anterolateral marginal area is more strongly produced as well.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0795667F7132FFA2FF410F800B1CFA51" ID-DOI="http://doi.org/10.5281/zenodo.3699469" ID-Zenodo-Dep="3699469" box="[192,1315,1506,1528]" httpUri="https://zenodo.org/record/3699469/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,287,1506,1528]" targetBox="[215,1371,238,1443]" targetPageId="5">
|
||
<paragraph id="535536F77132FFA2FF410F800B1CFA51" blockId="5.[192,1315,1506,1528]" box="[192,1315,1506,1528]" pageId="5" pageNumber="4">
|
||
<emphasis id="619EEAE57132FFA2FF410F800F6BFA5E" bold="true" box="[192,340,1506,1528]" pageId="5" pageNumber="4">Figures 3–4.</emphasis>
|
||
<taxonomicName id="94EA4D747132FFA2FEDA0F800C43FA51" authority="Gravely." authorityName="Gravely." box="[347,636,1506,1528]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="dawnae">
|
||
<emphasis id="619EEAE57132FFA2FEDA0F800C28FA5E" box="[347,535,1506,1527]" italics="true" pageId="5" pageNumber="4">Nigidius dawnae</emphasis>
|
||
Gravely.
|
||
</taxonomicName>
|
||
<emphasis id="619EEAE57132FFA2FD030F800CA6FA51" bold="true" box="[642,665,1506,1528]" pageId="5" pageNumber="4">3)</emphasis>
|
||
Major male, dorsal habitus.
|
||
<emphasis id="619EEAE57132FFA2FC570F800DD2FA51" bold="true" box="[982,1005,1506,1528]" pageId="5" pageNumber="4">4)</emphasis>
|
||
Lectotype male with labels.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1BF0657C7132FFA1FF6E0CEA0CD3F820" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="535536F77132FFA1FF6E0CEA0CEDF9F1" blockId="5.[192,1440,1572,1929]" lastBlockId="6.[192,1440,1566,1929]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
|
||
While initially considering both species to be undescribed, I researched existing names that might pertain to these two Indonesian species. The Borneo species in the obesus group has not been previously named and it is described and illustrated (
|
||
<figureCitation id="CBD12A727132FFA2FD4B0CA80D30F94B" box="[714,783,1738,1762]" captionStart="Figure 5" captionStartId="6.[192,275,1506,1528]" captionTargetBox="[223,1373,240,1459]" captionTargetId="figure@6.[210,1389,228,1476]" captionTargetPageId="6" captionText="Figure 5. Nigidius gravelyi Paulsen, n. sp., holotype male, with labels (inset)." figureDoi="http://doi.org/10.5281/zenodo.3699471" httpUri="https://zenodo.org/record/3699471/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
|
||
) below. The original description of
|
||
<taxonomicName id="94EA4D747132FFA2FB490CA90B9FF94B" authority="Boileau" authorityName="Boileau" box="[1224,1440,1738,1762]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="helleri">
|
||
<emphasis id="619EEAE57132FFA2FB490CA90B04F94B" box="[1224,1339,1738,1762]" italics="true" pageId="5" pageNumber="4">N. helleri</emphasis>
|
||
Boileau
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CBD12A727132FFA2FF480C890F32F8AA" box="[201,269,1771,1795]" captionStart="Figures 6–7" captionStartId="7.[192,287,1471,1493]" captionTargetBox="[233,1354,216,1395]" captionTargetId="figure@7.[220,1364,210,1415]" captionTargetPageId="7" captionText="Figures 6–7. Nigidius helleri Boileau. 6) Major male, dorsal habitus. 7) Holotype male with labels. Photo courtesy of Antoine Mantilleri, MNHN." figureDoi="http://doi.org/10.5281/zenodo.3699473" httpUri="https://zenodo.org/record/3699473/files/figure.png" pageId="5" pageNumber="4">Fig. 6</figureCitation>
|
||
) from
|
||
<collectingRegion id="912EF8157132FFA2FEDD0C8E0FF7F8AD" box="[348,456,1772,1796]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="4">Sumatra</collectingRegion>
|
||
mentioned its similarity to
|
||
<taxonomicName id="94EA4D747132FFA2FC970C8E0DB7F8AA" authorityName="Parry" authorityYear="1864" box="[790,904,1771,1795]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57132FFA2FC970C8E0DB7F8AA" box="[790,904,1771,1795]" italics="true" pageId="5" pageNumber="4">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
and, critically, the presence of a very prominent lobe on the head that is rounded anteriorly, truncate posteriorly, and perpendicular to the body (
|
||
<bibRefCitation id="377B4B067132FFA2FF490D4C0F51F8EF" author="Boileau" box="[200,366,1838,1862]" firstAuthor="Boileau" pageId="5" pageNumber="4" pagination="47 - 48" refId="ref6538" refString="Boileau, M. H. 1905. Description de Coleopteres nouveaux. Le Naturaliste 27 (431): 47 - 48." type="journal article" year="1905">Boileau 1905</bibRefCitation>
|
||
). This describes the lobed canthus of the obesus group perfectly. I requested an image of the
|
||
<typeStatus id="8C5188557132FFA2FE8F0D2D0F49F8CE" box="[270,374,1871,1895]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="94EA4D747132FFA2FE1D0D320C30F8CE" box="[412,527,1871,1895]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="helleri">
|
||
<emphasis id="619EEAE57132FFA2FE1D0D320C30F8CE" box="[412,527,1871,1895]" italics="true" pageId="5" pageNumber="4">N. helleri</emphasis>
|
||
</taxonomicName>
|
||
from the MNHN (
|
||
<figureCitation id="CBD12A727132FFA2FD720D2D0D05F8CE" box="[755,826,1871,1895]" captionStart="Figures 6–7" captionStartId="7.[192,287,1471,1493]" captionTargetBox="[233,1354,216,1395]" captionTargetId="figure@7.[220,1364,210,1415]" captionTargetPageId="7" captionText="Figures 6–7. Nigidius helleri Boileau. 6) Major male, dorsal habitus. 7) Holotype male with labels. Photo courtesy of Antoine Mantilleri, MNHN." figureDoi="http://doi.org/10.5281/zenodo.3699473" httpUri="https://zenodo.org/record/3699473/files/figure.png" pageId="5" pageNumber="4">Fig. 7</figureCitation>
|
||
), which confirmed that it is the Sumatran species of the obesus group. This is surprising because the name
|
||
<taxonomicName id="94EA4D747132FFA2FC100D130A3AF821" box="[913,1029,1904,1928]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="helleri">
|
||
<emphasis id="619EEAE57132FFA2FC100D130A3AF821" box="[913,1029,1904,1928]" italics="true" pageId="5" pageNumber="4">N. helleri</emphasis>
|
||
</taxonomicName>
|
||
has been traditionally used for a Javan or Sumatran species of the cornutus species group that is present in most collections (
|
||
<bibRefCitation id="377B4B067131FFA1FA9B0C7C0FBAF9F1" author="Mizunuma and Nagai" firstAuthor="Mizunuma" pageId="6" pageNumber="5" refId="ref6882" refString="Mizunuma, T., and S. Nagai. 1994. The Lucanid beetles of the world. Mushi-sha Iconographic series of Insects, Volume 1. H. Fujita Ed.; Tokyo, Japan. 1338 p." type="book" year="1994">Mizunuma and Nagai 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="377B4B067131FFA1FE110C220C17F9F1" author="Okuda" box="[400,552,1600,1624]" firstAuthor="Okuda" pageId="6" pageNumber="5" pagination="1 - 4" refId="ref6923" refString="Okuda, N. 2002. A new species of genus Nigidius MacLeay (Coleoptera, Lucanidae), from North Borneo, East Malaysia. Kogane 3: 1 - 4." type="journal article" year="2002">Okuda 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="377B4B067131FFA1FDB50C220CFAF9F1" author="Fujita" box="[564,709,1600,1624]" firstAuthor="Fujita" pageId="6" pageNumber="5" refId="ref6710" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo, Japan. 472 p." type="book" year="2010">Fujita 2010</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="0795667F7131FFA1FF410F800A22FA51" ID-DOI="http://doi.org/10.5281/zenodo.3699471" ID-Zenodo-Dep="3699471" box="[192,1053,1506,1528]" httpUri="https://zenodo.org/record/3699471/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,275,1506,1528]" targetBox="[223,1373,240,1459]" targetPageId="6">
|
||
<paragraph id="535536F77131FFA1FF410F800A22FA51" blockId="6.[192,1053,1506,1528]" box="[192,1053,1506,1528]" pageId="6" pageNumber="5">
|
||
<emphasis id="619EEAE57131FFA1FF410F800F10FA5E" bold="true" box="[192,303,1506,1528]" pageId="6" pageNumber="5">Figure 5.</emphasis>
|
||
<taxonomicName id="94EA4D747131FFA1FEB40F800C6AFA51" authority="Paulsen" authorityName="Paulsen" authorityYear="2018" box="[309,597,1506,1528]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="gravelyi" status="sp. nov.">
|
||
<emphasis id="619EEAE57131FFA1FEB40F800FCCFA5E" box="[309,499,1506,1527]" italics="true" pageId="6" pageNumber="5">Nigidius gravelyi</emphasis>
|
||
Paulsen
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="7AAD579E7131FFA1FDE10F800CA2FA5E" box="[608,669,1506,1527]" pageId="6" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype male, with labels (inset).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="535536F77131FFA1FF690C000CD3F820" blockId="6.[192,1440,1566,1929]" pageId="6" pageNumber="5">
|
||
The second mainland species of this group is from
|
||
<collectingCountry id="2BFD76677131FFA1FCF10C000DD4F9D3" box="[880,1003,1634,1658]" name="Myanmar" pageId="6" pageNumber="5">Myanmar</collectingCountry>
|
||
, northern
|
||
<collectingCountry id="2BFD76677131FFA1FBF30C000ADBF9D3" box="[1138,1252,1634,1658]" name="Thailand" pageId="6" pageNumber="5">Thailand</collectingCountry>
|
||
, and
|
||
<collectingCountry id="2BFD76677131FFA1FAAC0C000B58F9D0" box="[1325,1383,1634,1657]" name="Laos" pageId="6" pageNumber="5">Laos</collectingCountry>
|
||
and was first described as
|
||
<taxonomicName id="94EA4D747131FFA1FE4E0CE60D2CF932" authority="Gravely (1915)" authorityName="Gravely" authorityYear="1915" box="[463,787,1667,1691]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dawnae">
|
||
<emphasis id="619EEAE57131FFA1FE4E0CE60C6DF932" box="[463,594,1667,1691]" italics="true" pageId="6" pageNumber="5">N. dawnae</emphasis>
|
||
<bibRefCitation id="377B4B067131FFA1FDDB0CE10D2CF932" author="Gravely" box="[602,787,1667,1691]" firstAuthor="Gravely" pageId="6" pageNumber="5" pagination="407 - 431" refId="ref6786" refString="Gravely, F. H. 1915. A catalogue of the Lucanidae in the collection of the Indian Museum. Records of the Indian Museum 9 (6): 407 - 431." type="journal article" year="1915">Gravely (1915)</bibRefCitation>
|
||
</taxonomicName>
|
||
. Subsequently it has been identified incorrectly as
|
||
<taxonomicName id="94EA4D747131FFA1FA020CE60FBCF914" authority="de Lisle" authorityName="de Lisle" authorityYear="1974" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="larssoni">
|
||
<emphasis id="619EEAE57131FFA1FA020CE60F1EF914" italics="true" pageId="6" pageNumber="5">N. larssoni</emphasis>
|
||
de Lisle
|
||
</taxonomicName>
|
||
(de
|
||
<bibRefCitation id="377B4B067131FFA1FE330CC70C0FF914" author="Lisle" box="[434,560,1701,1725]" firstAuthor="Lisle" pageId="6" pageNumber="5" pagination="785 - 804" refId="ref6821" refString="Lisle, M. O. de 1974. Troisieme note sur quelques Coleoptera Lucanidae nouveaux ou peu connus. Revue Suisse de Zoologie 80 (4): 785 - 804." type="journal article" year="1974">Lisle 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="377B4B067131FFA1FDBB0CC70CFAF914" author="Fujita" box="[570,709,1701,1725]" firstAuthor="Fujita" pageId="6" pageNumber="5" refId="ref6710" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo, Japan. 472 p." type="book" year="2010">Fujita 2010</bibRefCitation>
|
||
) or
|
||
<taxonomicName id="94EA4D747131FFA1FD710CC40A00F914" authority="(Mizunuma 2000)" baseAuthorityName="Mizunuma" baseAuthorityYear="2000" box="[752,1087,1701,1725]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57131FFA1FD710CC40D5EF914" box="[752,865,1701,1725]" italics="true" pageId="6" pageNumber="5">N. obesus</emphasis>
|
||
(
|
||
<bibRefCitation id="377B4B067131FFA1FCEE0CC70A07F914" author="Mizunuma" box="[879,1080,1701,1725]" firstAuthor="Mizunuma" pageId="6" pageNumber="5" refId="ref6854" refString="Mizunuma, T. 2000. Stag beetles II (Lucanidae). Endless Collection Series 5. Toshitsugu Endo; Tokyo, Japan. 101 p." type="book" year="2000">Mizunuma 2000</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="377B4B067131FFA1FBC90CC40AC3F914" author="Bomans" box="[1096,1276,1701,1725]" firstAuthor="Bomans" pageId="6" pageNumber="5" pagination="326 - 337" refId="ref6605" refString="Bomans, H. E. 1993. Notes diverses sur le genre Nigidius MacLeay et descriptions d'especes nouvelles (Coleoptera, Lucanidae). Lambillionea 93 (3): 326 - 337." type="journal article" year="1993">Bomans (1993)</bibRefCitation>
|
||
synonymized
|
||
<taxonomicName id="94EA4D747131FFA1FF410CAA0F8FF976" authority="de Lisle" authorityName="de Lisle" authorityYear="1974" box="[192,432,1735,1759]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="larssoni">
|
||
<emphasis id="619EEAE57131FFA1FF410CAA0F78F976" box="[192,327,1735,1759]" italics="true" pageId="6" pageNumber="5">N. larssoni</emphasis>
|
||
de Lisle
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="94EA4D747131FFA1FE760CAA0C53F976" baseAuthorityName="Mizunuma" baseAuthorityYear="2000" box="[503,620,1735,1759]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57131FFA1FE760CAA0C53F976" box="[503,620,1735,1759]" italics="true" pageId="6" pageNumber="5">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="377B4B067131FFA1FDFD0CA50D24F976" author="Araya" box="[636,795,1735,1759]" firstAuthor="Araya" pageId="6" pageNumber="5" pagination="18 - 25" refId="ref6449" refString="Araya, K. 2003. Notes on some type specimens of the genus Nigidius from Asia stored in several European Museums. Gekkan Mushi 402: 18 - 25." type="journal article" year="2003">Araya (2003)</bibRefCitation>
|
||
included images of the type material for all three of these available names.
|
||
<bibRefCitation id="377B4B067131FFA1FE630C8B0CBDF8A8" author="Fujita" box="[482,642,1769,1793]" firstAuthor="Fujita" pageId="6" pageNumber="5" refId="ref6710" refString="Fujita, H. 2010. The lucanid beetles of the world. Mushi-Sha's iconographic series of insects, 6. Mushi- Sha; Tokyo, Japan. 472 p." type="book" year="2010">Fujita (2010)</bibRefCitation>
|
||
recalled
|
||
<taxonomicName id="94EA4D747131FFA1FD720C8B0D44F8A8" authorityName="de Lisle" authorityYear="1974" box="[755,891,1769,1793]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="larssoni">
|
||
<emphasis id="619EEAE57131FFA1FD720C8B0D44F8A8" box="[755,891,1769,1793]" italics="true" pageId="6" pageNumber="5">N. larssoni</emphasis>
|
||
</taxonomicName>
|
||
from synonymy without mention of
|
||
<taxonomicName id="94EA4D747131FFA1FABF0C8B0ED4F88B" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dawnae">
|
||
<emphasis id="619EEAE57131FFA1FABF0C8B0ED4F88B" italics="true" pageId="6" pageNumber="5">N. dawnae</emphasis>
|
||
</taxonomicName>
|
||
. Examination of the
|
||
<typeStatus id="8C5188557131FFA1FE760D690C60F88A" box="[503,607,1803,1827]" pageId="6" pageNumber="5" type="holotype">holotype</typeStatus>
|
||
from ZMUC confirmed that
|
||
<taxonomicName id="94EA4D747131FFA1FC3E0D690A79F88B" authorityName="de Lisle" authorityYear="1974" box="[959,1094,1802,1826]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="larssoni">
|
||
<emphasis id="619EEAE57131FFA1FC3E0D690A79F88B" box="[959,1094,1802,1826]" italics="true" pageId="6" pageNumber="5">N. larssoni</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<emphasis id="619EEAE57131FFA1FA950D690B9FF88B" box="[1300,1440,1802,1826]" italics="true" pageId="6" pageNumber="5">
|
||
<taxonomicName id="94EA4D747131FFA1FA950D690BA3F88B" box="[1300,1436,1802,1826]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dawnae">N. dawnae</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Compared to
|
||
<taxonomicName id="94EA4D747131FFA1FEE80D4F0FE1F8ED" baseAuthorityName="Mizunuma" baseAuthorityYear="2000" box="[361,478,1836,1860]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="obesus">
|
||
<emphasis id="619EEAE57131FFA1FEE80D4F0FE1F8ED" box="[361,478,1836,1860]" italics="true" pageId="6" pageNumber="5">N. obesus</emphasis>
|
||
</taxonomicName>
|
||
, the anterolateral pronotal projection is broader and more strongly produced in this species. When compared with similarly sized specimens,
|
||
<taxonomicName id="94EA4D747131FFA1FC5F0D2D0A5CF8CF" box="[990,1123,1870,1894]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="dawnae">
|
||
<emphasis id="619EEAE57131FFA1FC5F0D2D0A5CF8CF" box="[990,1123,1870,1894]" italics="true" pageId="6" pageNumber="5">N. dawnae</emphasis>
|
||
</taxonomicName>
|
||
has the largest, broadest mandibles of any species in the obesus group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |