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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.82.67973" ID-GBIF-Dataset="c41f2af2-9b31-4886-8e5f-3d56121c97ba" ID-PMC="PMC8367933" ID-Pensoft-Pub="1314-4049-82-57" ID-Pensoft-UUID="DF020A7A4C63518BBE6CC027E3D000E8" ID-PubMed="34408538" ModsDocID="1314-4049-82-57" checkinTime="1628596712451" checkinUser="pensoft" docAuthor="Ren, Guang-Cong, Wanasinghe, Dhanushka N., Monkai, Jutamart, Mortimer, Peter E., Hyde, Kevin D., Xu, Jian-Chu, Pang, Aimin &amp; Gui, Heng" docDate="2021" docId="586F538B54BB5D95A10A50FE2A43AEE0" docLanguage="en" docName="MycoKeys 82: 57-79" docOrigin="MycoKeys 82" docPubDate="2021-08-09" docSource="http://dx.doi.org/10.3897/mycokeys.82.67973" docTitle="Hermatomyces jinghaensis G. C. Ren &amp; K. D. Hyde 2021, sp. nov." docType="treatment" docVersion="4" id="DF020A7A4C63518BBE6CC027E3D000E8" lastPageNumber="57" masterDocId="DF020A7A4C63518BBE6CC027E3D000E8" masterDocTitle="Novel saprobic Hermatomyces species (Hermatomycetaceae, Pleosporales) from China (Yunnan Province) and Thailand" masterLastPageNumber="79" masterPageNumber="57" pageNumber="57" updateTime="1668136700934" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Novel saprobic Hermatomyces species (Hermatomycetaceae, Pleosporales) from China (Yunnan Province) and Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Guang-Cong</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9923-2626</mods:nameIdentifier>
<mods:affiliation>Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Guiyang Nursing Vocational College, Guiyang 550081, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wanasinghe, Dhanushka N.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1759-3933</mods:nameIdentifier>
<mods:affiliation>Center for Mountain futures, Kunming Institute of Botany, Chinese Academy of Sciences, Honghe County 654400, Yunnan, China &amp; CIFOR-ICRAF China Program, World Agroforestry (ICRAF), Kunming 650201, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Monkai, Jutamart</mods:namePart>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mortimer, Peter E.</mods:namePart>
<mods:affiliation>Center for Mountain futures, Kunming Institute of Botany, Chinese Academy of Sciences, Honghe County 654400, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hyde, Kevin D.</mods:namePart>
<mods:affiliation>School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand &amp; Guiyang Nursing Vocational College, Guiyang 550081, Guizhou, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Jian-Chu</mods:namePart>
<mods:affiliation>Center for Mountain futures, Kunming Institute of Botany, Chinese Academy of Sciences, Honghe County 654400, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pang, Aimin</mods:namePart>
<mods:affiliation>Science and Technology on Aerospace Chemical Power Laboratory, Hubei Institute of Aerospace Chemotechnology, Xiangyang, 441003, Hubei, China</mods:affiliation>
<mods:nameIdentifier type="email">ppam@tom.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gui, Heng</mods:namePart>
<mods:affiliation>Center for Mountain futures, Kunming Institute of Botany, Chinese Academy of Sciences, Honghe County 654400, Yunnan, China</mods:affiliation>
<mods:nameIdentifier type="email">guiheng@mail.kib.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-09</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>79</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.82.67973</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.82.67973</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-82-57</mods:identifier>
<mods:identifier type="Pensoft-UUID">DF020A7A4C63518BBE6CC027E3D000E8</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="184008557" LSID="urn:lsid:plazi:treatment:586F538B54BB5D95A10A50FE2A43AEE0" httpUri="http://treatment.plazi.org/id/586F538B54BB5D95A10A50FE2A43AEE0" lastPageNumber="57" pageId="0" pageNumber="57">
<subSubSection pageId="0" pageNumber="57" type="nomenclature">
<paragraph pageId="0" pageNumber="57">
<taxonomicName LSID="586F538B-54BB-5D95-A10A-50FE2A43AEE0" authority="G. C. Ren &amp; K. D. Hyde" authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Hermatomycetaceae" genus="Hermatomyces" higherTaxonomySource="treatment-meta" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis" status="sp. nov.">Hermatomyces jinghaensis G.C. Ren &amp; K.D. Hyde</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="57">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="description">
<paragraph pageId="0" pageNumber="57">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Hermatomyces jinghaensis (HKAS 112167, holotype) a, b sporodochia on natural substrate c vertical section of sporodochium d conidiophores e, f conidiogenous cells g-l cylindrical conidia m-s mature lenticular conidia. Scale bars: 50 μm (c); 30 μm (d); 20 μm (e-r); 30 μm (s)." figureDoi="10.3897/mycokeys.82.67973.figure3" httpUri="https://binary.pensoft.net/fig/575841" pageId="0" pageNumber="57">Figure 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="etymology">
<paragraph pageId="0" pageNumber="57">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="57">
The species epithet &quot;
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
&quot; refers to the location where the species was collected.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="holotype">
<paragraph pageId="0" pageNumber="57">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="57">HKAS 112167.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="description">
<paragraph pageId="0" pageNumber="57">Description.</paragraph>
<paragraph pageId="0" pageNumber="57">
<emphasis italics="true" pageId="0" pageNumber="57">Saprobic</emphasis>
on unidentified woody litter.
<emphasis bold="true" pageId="0" pageNumber="57">Sexual morph</emphasis>
Undetermined.
<emphasis bold="true" pageId="0" pageNumber="57">Asexual morph</emphasis>
<emphasis italics="true" pageId="0" pageNumber="57">Colonies</emphasis>
on natural substrate forming sporodochial conidiomata, superficial, scattered, small groups, circular, sterile mycelial outer zone enclosing a black velvety margin, dense, thick, black sporulating center, shiny, glistening, circular or oval, conidia readily liberated when agitated.
<emphasis italics="true" pageId="0" pageNumber="57">Mycelium</emphasis>
superficial, branched, septate, hyaline to pale brown, 2-3
<normalizedToken originalValue="μm">μm</normalizedToken>
wide.
<emphasis italics="true" pageId="0" pageNumber="57">Conidiophores</emphasis>
30-45
<normalizedToken originalValue="×">x</normalizedToken>
2-3
<normalizedToken originalValue="μm">μm</normalizedToken>
, mononematous, cylindrical, straight or flexuous, smooth, pale brown.
<emphasis italics="true" pageId="0" pageNumber="57">Conidiogenous cells</emphasis>
4-6
<normalizedToken originalValue="×">x</normalizedToken>
2-3
<normalizedToken originalValue="μm">μm</normalizedToken>
, monoblastic, integrated, terminal, determinate, often arising directly on the superficial mycelium, cylindrical, ampulliform, hyaline to pale brown, smooth finely verruculose.
<emphasis italics="true" pageId="0" pageNumber="57">Conidia</emphasis>
dimorphic solitary, smooth-walled.
<emphasis italics="true" pageId="0" pageNumber="57">Lenticular conidia</emphasis>
30-40
<normalizedToken originalValue="×">x</normalizedToken>
25-30
<normalizedToken originalValue="μm">μm</normalizedToken>
(x = 37
<normalizedToken originalValue="×">x</normalizedToken>
28
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 20), 21-25
<normalizedToken originalValue="μm">μm</normalizedToken>
thick, thick-walled, circular to oval in front view, smooth, solitary, muriform, central cells brown to dark brown, peripheral cells hyaline to subhyaline, forming a wide and distinct ring, sometimes slightly constricted at septa, obovoid or oblong in lateral view, central cells brown to dark brown, peripheral cells pale brown to brown.
<emphasis italics="true" pageId="0" pageNumber="57">Cylindrical conidia</emphasis>
33-43
<normalizedToken originalValue="μm">μm</normalizedToken>
in length, 11-13
<normalizedToken originalValue="µm">µm</normalizedToken>
wide in broadest part of lower cells (x = 39
<normalizedToken originalValue="×">x</normalizedToken>
12
<normalizedToken originalValue="μm">μm</normalizedToken>
, n = 20), clavate or subcylindrical, straight or flexuous, septate, constricted distinct at the septa, with large guttules, consisting of one or two columns, each column with 6-8 cells, apical cell rectangular to globose, smooth, hyaline, smooth, basal cells acute, rectangular to cylindrical, pale brown.
</paragraph>
<caption doi="10.3897/mycokeys.82.67973.figure3" httpUri="https://binary.pensoft.net/fig/575841" pageId="0" pageNumber="57" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="57">
<emphasis bold="true" pageId="0" pageNumber="57">Figure 3.</emphasis>
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
(HKAS 112167, holotype)
<emphasis bold="true" pageId="0" pageNumber="57">a, b</emphasis>
sporodochia on natural substrate
<emphasis bold="true" pageId="0" pageNumber="57">c</emphasis>
vertical section of sporodochium
<emphasis bold="true" pageId="0" pageNumber="57">d</emphasis>
conidiophores
<emphasis bold="true" pageId="0" pageNumber="57">e, f</emphasis>
conidiogenous cells
<emphasis bold="true" pageId="0" pageNumber="57">g-l</emphasis>
cylindrical conidia
<emphasis bold="true" pageId="0" pageNumber="57">m-s</emphasis>
mature lenticular conidia. Scale bars: 50
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="57">c</emphasis>
); 30
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="57">d</emphasis>
); 20
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="57">e-r</emphasis>
); 30
<normalizedToken originalValue="μm">μm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="57">s</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="distribution">
<paragraph pageId="0" pageNumber="57">Known host and distribution.</paragraph>
<paragraph pageId="0" pageNumber="57">Unidentified woody litter (China)</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="materials_examined">
<paragraph pageId="0" pageNumber="57">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="57">
<materialsCitation collectingDate="2019-12-19" collectorName="D. N. Wanasinghe" country="China" county="Xishuangbanna Dai Autonomous Prefecture" location="Jingha" longitude="101.0935" municipality="Jinghong" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
,
<collectingRegion country="China" name="Yunnan">Yunnan Province</collectingRegion>
,
<collectingCounty>Xishuangbanna Dai Autonomous Prefecture</collectingCounty>
,
<collectingMunicipality>Jinghong</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:586F538B54BB5D95A10A50FE2A43AEE0:EC0932602C5EE2DEB76FBFE508457664" country="China" county="Xishuangbanna Dai Autonomous Prefecture" longitude="101.0935" municipality="Jinghong" name="Jingha" stateProvince="Yunnan Province">Jingha</location>
(21°78.06'N,
<geoCoordinate degrees="101" direction="east" minutes="05.61" orientation="longitude" precision="9" value="101.0935">101°05.61'E</geoCoordinate>
), on unidentified woody litter,
<collectingDate value="2019-12-19">19 December 2019</collectingDate>
,
<collectorName>D.N. Wanasinghe</collectorName>
, DW57 (HKAS 112167,
<emphasis bold="true" italics="true" pageId="0" pageNumber="57">
<typeStatus>holotype</typeStatus>
</emphasis>
), no living culture
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="57" type="notes">
<paragraph pageId="0" pageNumber="57">Notes.</paragraph>
<paragraph pageId="0" pageNumber="57">
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
is introduced as a new species based on its distinct morphology and the phylogenetic results of a combined LSU, ITS,
<emphasis italics="true" pageId="0" pageNumber="57">tub</emphasis>
2,
<emphasis italics="true" pageId="0" pageNumber="57">tef</emphasis>
1-α and
<emphasis italics="true" pageId="0" pageNumber="57">rpb</emphasis>
2 dataset.
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
nested with
<taxonomicName class="Dothideomycetes" family="Hermatomycetaceae" kingdom="Fungi" lsidName="H. clematidis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="clematidis">
<emphasis italics="true" pageId="0" pageNumber="57">H. clematidis</emphasis>
</taxonomicName>
and
<taxonomicName class="Dothideomycetes" family="Hermatomycetaceae" kingdom="Fungi" lsidName="H. trangensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="trangensis">
<emphasis italics="true" pageId="0" pageNumber="57">H. trangensis</emphasis>
</taxonomicName>
in a strongly supported monophyletic group (99% ML, 100% MP, 1.00 PP; Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Phylogenetic RAxML tree based on analysis of a combined LSU, ITS, tub 2, tef 1 - α and rpb 2 and dataset. Bootstrap support values for ML and MP equal to or higher than 75 % and Bayesian PP equal to or greater than 0.95 are shown at nodes. Hyphens (--) represent support values less than 75 % / 0.95 BYPP. The ex-type strains are in bold and the new isolate in this study is in blue bold. The tree is rooted with Anteaglonium globosum (ANM 925.2) and A. parvulum (MFLUCC 14 - 0821). The scale bar represents the expected number of nucleotide substitutions per site." figureDoi="10.3897/mycokeys.82.67973.figure1" httpUri="https://binary.pensoft.net/fig/575839" pageId="0" pageNumber="57">1</figureCitation>
).
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
is characterized by both lenticular and cylindrical conidia.
<taxonomicName authorityName="G. C. Ren &amp; K. D. Hyde" authorityYear="2021" class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces jinghaensis</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Dothideomycetes" family="Hermatomycetaceae" kingdom="Fungi" lsidName="H. clematidis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="clematidis">
<emphasis italics="true" pageId="0" pageNumber="57">H. clematidis</emphasis>
</taxonomicName>
in having cylindrical conidia with one or two columns, each of which has 6-8 cells with large guttules, while the latter has 5-6 cells for each column conidia.
<taxonomicName class="Dothideomycetes" family="Lophiotremataceae" genus="Hermatomyces" kingdom="Fungi" lsidName="Hermatomyces trangensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="trangensis">
<emphasis italics="true" pageId="0" pageNumber="57">Hermatomyces trangensis</emphasis>
</taxonomicName>
differs from
<taxonomicName class="Dothideomycetes" family="Hermatomycetaceae" kingdom="Fungi" lsidName="H. jinghaensis" order="Pleosporales" pageId="0" pageNumber="57" phylum="Ascomycota" rank="species" species="jinghaensis">
<emphasis italics="true" pageId="0" pageNumber="57">H. jinghaensis</emphasis>
</taxonomicName>
in having only lenticular conidia.
</paragraph>
</subSubSection>
</treatment>
</document>