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<document id="4975C3C311EF1FB4AC199E09FE265AF5" ID-CLB-Dataset="30789" ID-DOI="10.5852/ejt.2018.402" ID-GBIF-Dataset="8f1510f6-1b79-467f-9e8d-f193ac82cc4b" ID-Zenodo-Dep="1169559" ID-ZooBank="A3CE68AA-C5C6-40B7-B57C-6EF6D949149B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518136191129" checkinUser="plazi" docAuthor="Ruiz-C, Raquel I., Román-Valencia, César, Taphorn, Donald C., Buckup, Paulo A. &amp; Ortega, Hernán" docDate="2018" docId="2C0787E7FFC9D432FDA9FAF8FA2FFAB0" docLanguage="en" docName="ejt-402_Ruiz-C_Roman-Valencia.pdf.imf" docOrigin="European Journal of Taxonomy 402" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Astyanax gandhiae Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega 2018, sp. nov." docType="treatment" docVersion="17" lastPageNumber="15" masterDocId="D03EFF9FFFC5D43CFFDCFFD0FE0FFFBE" masterDocTitle="Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species" masterLastPageNumber="45" masterPageNumber="1" originalUpdateDomain="Boston" originalUpdateTime="1547298528796" originalUpdateUser="yanell" pageNumber="13" updateTime="1698520653973" updateUser="ExternalLinkService">
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<mods:title id="8D8D56C4A2B26643D2848D9CCA85691B">Revision of the Astyanax orthodus species-group (Teleostei: Characidae) with descriptions of three new species</mods:title>
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<mods:namePart id="7B7E26080B0DC5DA916B07777FD0FCAF">Ruiz-C, Raquel I.</mods:namePart>
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<mods:namePart id="0175F357F0AE1308A1ABF3DB945BF88B">Román-Valencia, César</mods:namePart>
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<mods:namePart id="EE067066DC10A9B4EDBC9D9ED5A8E817">Taphorn, Donald C.</mods:namePart>
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<mods:namePart id="65A1FE09B6A46155499720658A4EC48C">Buckup, Paulo A.</mods:namePart>
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<mods:title id="4F89E8B578D5A4C9D67683546BDE668E">European Journal of Taxonomy</mods:title>
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<treatment id="2C0787E7FFC9D432FDA9FAF8FA2FFAB0" ID-DOI="http://doi.org/10.5281/zenodo.5622614" ID-GBIF-Taxon="140562874" ID-Zenodo-Dep="5622614" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0" httpUri="http://treatment.plazi.org/id/2C0787E7FFC9D432FDA9FAF8FA2FFAB0" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
<subSubSection id="ECB4657AFFC9D430FDA9FAF8FACCFAD8" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="A41136F1FFC9D430FDA9FAF8FDB0FAFD" blockId="12.[369,1219,1320,1417]" box="[629,959,1320,1347]" pageId="12" pageNumber="13">
<heading id="FF59819DFFC9D430FDA9FAF8FDB0FAFD" box="[629,959,1320,1347]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="13" reason="2">
<taxonomicName id="63AE4D72FFC9D430FDA9FAF8FD52FAFD" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[629,861,1320,1347]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFC9D430FDA9FAF8FD52FAFD" bold="true" box="[629,861,1320,1347]" italics="true" pageId="12" pageNumber="13">Astyanax gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FCB8FAF9FDB0FAFD" box="[868,959,1321,1347]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="A41136F1FFC9D430FEADFA9BFACCFAD8" blockId="12.[369,1219,1320,1417]" box="[369,1219,1355,1382]" pageId="12" pageNumber="13">urn:lsid:zoobank.org:act:DB1A95AA-8E70-40F4-B739-043872A32906</paragraph>
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<subSubSection id="ECB4657AFFC9D430FD68FABEFD70FA37" box="[692,895,1390,1417]" pageId="12" pageNumber="13" type="description">
<paragraph id="A41136F1FFC9D430FD68FABEFD70FA37" blockId="12.[369,1219,1320,1417]" box="[692,895,1390,1417]" pageId="12" pageNumber="13">
<figureCitation id="3C952A74FFC9D430FD68FABEFCF0FA37" box="[692,767,1390,1417]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="12" pageNumber="13">Figs 4</figureCitation>
,
<figureCitation id="3C952A74FFC9D430FCD0FABEFD12FA36" box="[780,797,1390,1416]" captionStart="Fig. 6" captionStartId="14.[189,232,1966,1992]" captionTargetBox="[189,1398,1437,1936]" captionTargetId="figure@14.[189,1398,1437,1936]" captionTargetPageId="14" captionText="Fig. 6. Astyanax gandhiae sp. nov., holotype, 89.4 mm SL, Condorcanqui, Amazonas, Upper Cenepa River drainage (MUSM 46845). Scale bar = 1 cm." httpUri="https://zenodo.org/record/1169571/files/figure.png" pageId="12" pageNumber="13">6</figureCitation>
,
<tableCitation id="E92C034AFFC9D430FCF4FABEFD70FA37" box="[808,895,1390,1417]" captionStart="Table 1" captionStartId="11.[145,214,599,625]" captionTargetPageId="11" captionText="Table 1. Morphometric data for some members of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFCED437FF4DFD87FA92FDCF" pageId="12" pageNumber="13" tableUuid="F0D16679FFCED437FF4DFD87FA92FDCF">Table 1</tableCitation>
</paragraph>
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<subSubSection id="ECB4657AFFC9D430FF61FA60FA09F8E6" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="A41136F1FFC9D430FF61FA60FF39FA74" blockId="12.[189,310,1456,1478]" box="[189,310,1456,1482]" pageId="12" pageNumber="13">
<heading id="FF59819DFFC9D430FF61FA60FF39FA74" box="[189,310,1456,1482]" pageId="12" pageNumber="13">
<emphasis id="96DAEAE3FFC9D430FF61FA60FF39FA74" bold="true" box="[189,310,1456,1482]" pageId="12" pageNumber="13">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFC9D430FF61FA0FFA09F8E6" blockId="12.[189,1399,1502,1881]" pageId="12" pageNumber="13">
<taxonomicName id="63AE4D72FFC9D430FF61FA0FFF93FA46" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[189,412,1502,1529]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFC9D430FF61FA0FFF93FA46" box="[189,412,1502,1529]" italics="true" pageId="12" pageNumber="13">Astyanax gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FE7EFA0FFFF4FA47" box="[418,507,1503,1529]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
is a member of the
<taxonomicName id="63AE4D72FFC9D430FD00FA0EFD4BFA46" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[732,836,1502,1528]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFC9D430FD00FA0EFD4BFA46" box="[732,836,1502,1528]" italics="true" pageId="12" pageNumber="13">orthodus</emphasis>
</taxonomicName>
species-group of
<taxonomicName id="63AE4D72FFC9D430FBCEFA0FFA71FA47" authorityName="Baird &amp; Girard" authorityYear="1854" box="[1042,1150,1503,1529]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
<emphasis id="96DAEAE3FFC9D430FBCEFA0FFA71FA47" box="[1042,1150,1503,1529]" italics="true" pageId="12" pageNumber="13">Astyanax</emphasis>
</taxonomicName>
differing from others members of the group (
<taxonomicName id="63AE4D72FFC9D430FE01F9D2FC66F9A2" authorityName="Eigenmann in Eigenmann &amp; Ogle" authorityYear="1907" box="[477,617,1538,1564]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="orthodus">
<emphasis id="96DAEAE3FFC9D430FE01F9D2FC66F9A2" box="[477,617,1538,1564]" italics="true" pageId="12" pageNumber="13">A. orthodus</emphasis>
</taxonomicName>
,
<emphasis id="96DAEAE3FFC9D430FDA7F9D2FCF1F9A2" box="[635,766,1538,1564]" italics="true" pageId="12" pageNumber="13">A.villwocki</emphasis>
,
<taxonomicName id="63AE4D72FFC9D430FCCCF9D2FD92F9A2" authorityName="Myers" authorityYear="1942" box="[784,925,1538,1564]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="superbus">
<emphasis id="96DAEAE3FFC9D430FCCCF9D2FD92F9A2" box="[784,925,1538,1564]" italics="true" pageId="12" pageNumber="13">A. superbus</emphasis>
</taxonomicName>
,
<taxonomicName id="63AE4D72FFC9D430FC73F9D2FA4DF9A2" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[943,1090,1538,1564]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
<emphasis id="96DAEAE3FFC9D430FC73F9D2FA4DF9A2" box="[943,1090,1538,1564]" italics="true" pageId="12" pageNumber="13">A. bopiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FB90F9D2FAC9F9A2" box="[1100,1222,1538,1564]" pageId="12" pageNumber="13">nom. nov.</taxonomicNameLabel>
,
<taxonomicName id="63AE4D72FFC9D430FB09F9D2FB79F9A2" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[1237,1398,1538,1564]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="boliviensis" status="sp. nov.">
<emphasis id="96DAEAE3FFC9D430FB09F9D2FB79F9A2" box="[1237,1398,1538,1564]" italics="true" pageId="12" pageNumber="13">A. boliviensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FF61F9F5FF12F981" box="[189,285,1573,1599]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="63AE4D72FFC9D430FEF0F9F5FFB2F981" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Torres-Mejia, Hernandez &amp; Senechal" baseAuthorityYear="2012" box="[300,445,1573,1599]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="yariguies" status="comb. nov.">
<emphasis id="96DAEAE3FFC9D430FEF0F9F5FFB2F981" box="[300,445,1573,1599]" italics="true" pageId="12" pageNumber="13">A. yariguies</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FE14F9F5FC43F981" box="[456,588,1573,1599]" pageId="12" pageNumber="13">comb. nov.</taxonomicNameLabel>
and
<taxonomicName id="63AE4D72FFC9D430FD57F9F5FD08F981" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[651,775,1573,1599]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="embera" status="sp. nov.">
<emphasis id="96DAEAE3FFC9D430FD57F9F5FD08F981" box="[651,775,1573,1599]" italics="true" pageId="12" pageNumber="13">A. embera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FCCDF9F5FD60F981" box="[785,879,1573,1599]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
) in lacking a spot on the caudal peduncle. It can be further distinguished from
<emphasis id="96DAEAE3FFC9D430FDA5F998FD08F9DC" box="[633,775,1608,1634]" italics="true" pageId="12" pageNumber="13">A. villwocki</emphasis>
by having a short lateral stripe extending from the caudal spot, that does not extend anteriorly beyond a vertical through the origin of the anal fin (vs lateral stripe extending anteriorly beyond a vertical through anterior origin of the anal fin). It differs from
<taxonomicName id="63AE4D72FFC9D430FF61F962FF40F972" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[189,335,1714,1740]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="bopiensis" status="nom. nov.">
<emphasis id="96DAEAE3FFC9D430FF61F962FF40F972" box="[189,335,1714,1740]" italics="true" pageId="12" pageNumber="13">A. bopiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FE84F962FFC0F972" box="[344,463,1714,1740]" pageId="12" pageNumber="13">nom. nov.</taxonomicNameLabel>
in having fewer teeth that do not cover more than a third of the length of the maxillary (vs a larger number of teeth covering more than two thirds of the maxillary). It differs from
<taxonomicName id="63AE4D72FFC9D430FF61F928FF20F8AC" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[189,303,1784,1810]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="moorii" status="comb. nov.">
<emphasis id="96DAEAE3FFC9D430FF61F928FF20F8AC" box="[189,303,1784,1810]" italics="true" pageId="12" pageNumber="13">A. moorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFC9D430FEE5F928FFB1F8AD" box="[313,446,1784,1811]" pageId="12" pageNumber="13">comb. nov.</taxonomicNameLabel>
for dorsal-fin-hypural distance less than 45% SL(vs more than 50), by dorsalpectoral distance greater than 50% SL (vs less than 50), interorbital distance greater than 32% HL (vs less 32) and by upper jaw length than less 35% HL (vs greater than 40).
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFC9D430FF61F850FC94F855" pageId="12" pageNumber="13" type="etymology">
<paragraph id="A41136F1FFC9D430FF61F850FF4BF824" blockId="12.[189,324,1920,1946]" box="[189,324,1920,1946]" pageId="12" pageNumber="13">
<heading id="FF59819DFFC9D430FF61F850FF4BF824" bold="true" box="[189,324,1920,1946]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="96DAEAE3FFC9D430FF61F850FF4BF824" bold="true" box="[189,324,1920,1946]" pageId="12" pageNumber="13">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFC9D430FF61F87EFC94F855" blockId="12.[189,1399,1966,2028]" pageId="12" pageNumber="13">This species epithet is named in homage to the late Mrs Maria Gandhi Calderon, mother of the first author, and used as a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFC8D431FF61FEDBFFF5FC4D" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="A41136F1FFC8D431FF61FEDBFFA5FE9B" blockId="13.[189,426,267,293]" box="[189,426,267,293]" pageId="13" pageNumber="14">
<heading id="FF59819DFFC8D431FF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="96DAEAE3FFC8D431FF61FEDBFFA5FE9B" bold="true" box="[189,426,267,293]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFC8D431FF36FEEAFF53FEEA" blockId="13.[189,1399,314,412]" box="[234,348,314,340]" pageId="13" pageNumber="14">
<emphasis id="96DAEAE3FFC8D431FF36FEEAFF53FEEA" bold="true" box="[234,348,314,340]" pageId="13" pageNumber="14">Holotype</emphasis>
</paragraph>
<paragraph id="A41136F1FFC8D431FF61FE8EFDC3FE22" blockId="13.[189,1399,314,412]" pageId="13" pageNumber="14">
<materialsCitation id="14C63CACFFC8D431FF61FE8EFDC8FE22" ID-GBIF-Occurrence="1986481911" collectionCode="MUSM" country="Peru" latitude="-3.9708333" location="Condorcanqui Province, Marañon River Basin, upper Cenepa River drainage" longLatPrecision="21" longitude="-78.6875" pageId="13" pageNumber="14" specimenCode="MUSM 46845" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<collectingCountry id="DCB97661FFC8D431FF61FE8EFF02FEC6" box="[189,269,350,376]" name="Peru" pageId="13" pageNumber="14">PERU</collectingCountry>
:
<quantity id="63569B14FFC8D431FEC5FE8DFF8CFEC6" box="[281,387,349,376]" metricMagnitude="-2" metricUnit="m" metricValue="8.94" pageId="13" pageNumber="14" unit="mm" value="89.4">89.4 mm</quantity>
SL, Department of
<collectingRegion id="666AF813FFC8D431FDADFE8EFCFFFEC6" box="[625,752,350,376]" country="Peru" name="Amazonas" pageId="13" pageNumber="14">Amazonas</collectingRegion>
,
<location id="A171602AFFC8D431FD21FE8DFFC2FE25" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FD21FE8DFFC2FE25" country="Peru" latitude="-3.9708333" longLatPrecision="21" longitude="-78.6875" municipality="River Basin" name="Condorcanqui Province, Marañon River Basin, upper Cenepa River drainage" pageId="13" pageNumber="14" stateProvince="Amazonas">Condorcanqui Province, Marañon River Basin, upper Cenepa River drainage</location>
,
<geoCoordinate id="C19A5036FFC8D431FE04FEAFFC55FE25" box="[472,602,383,411]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="15" value="-3.9708333">3°5815″ S</geoCoordinate>
,
<geoCoordinate id="C19A5036FFC8D431FDBAFEAFFCF0FE25" box="[614,767,383,411]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="15" value="-78.6875">78°4115″ W</geoCoordinate>
(
<specimenCode id="F4089E8AFFC8D431FCD2FE51FDCFFE22" box="[782,960,385,412]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 46845</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="A41136F1FFC8D431FF36FE13FF68FE63" blockId="13.[189,1399,451,1011]" box="[234,359,451,477]" pageId="13" pageNumber="14">
<emphasis id="96DAEAE3FFC8D431FF36FE13FF68FE63" bold="true" box="[234,359,451,477]" pageId="13" pageNumber="14">Paratypes</emphasis>
</paragraph>
<paragraph id="A41136F1FFC8D431FF61FE37FFF5FC4D" blockId="13.[189,1399,451,1011]" pageId="13" pageNumber="14">
<materialsCitation id="14C63CACFFC8D431FF61FE37FB7DFD9A" ID-GBIF-Occurrence="1806442544" collectionCode="MUSM" country="Peru" location="Amazon River Basin, Condorcanqui, upper Cenepa River drainage" pageId="13" pageNumber="14" specimenCode="MUSM 20891, MUSM 21278" specimenCount="14" stateProvince="Amazonas" typeStatus="paratype">
<collectingCountry id="DCB97661FFC8D431FF61FE37FF02FDBF" box="[189,269,487,513]" name="Peru" pageId="13" pageNumber="14">PERU</collectingCountry>
: Department of
<collectingRegion id="666AF813FFC8D431FE17FE36FC45FDBE" box="[459,586,486,512]" country="Peru" name="Amazonas" pageId="13" pageNumber="14">Amazonas</collectingRegion>
,
<location id="A171602AFFC8D431FD89FE36FB7CFDBE" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FD89FE36FB7CFDBE" box="[597,1395,486,513]" country="Peru" county="Amazon River" municipality="upper Cenepa River" name="Amazon River Basin, Condorcanqui, upper Cenepa River drainage" pageId="13" pageNumber="14" stateProvince="Amazonas">Amazon River Basin, Condorcanqui, upper Cenepa River drainage</location>
:
<specimenCount id="B2A8FD78FFC8D431FF61FDDAFF53FD9A" box="[189,348,522,548]" pageId="13" pageNumber="14" type="generic">14 specimens</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FDF8FDDAFCD1FD9A" box="[548,734,522,548]" metricMagnitude="-2" metricUnit="m" metricValue="8.584999999999999" metricValueMax="10.959999999999999" metricValueMin="6.21" pageId="13" pageNumber="14" unit="mm" value="85.85" valueMax="109.6" valueMin="62.1">62.1109.6 mm</quantity>
SL,
<location id="A171602AFFC8D431FCC4FDDAFDE2FD9A" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FCC4FDDAFDE2FD9A" box="[792,1005,522,548]" country="Peru" county="Amazon River" municipality="upper Cenepa River" name="Quebrada Capitán" pageId="13" pageNumber="14" stateProvince="Amazonas">Quebrada Capitán</location>
(
<specimenCode id="F4089E8AFFC8D431FC21FDDAFAA2FD9A" box="[1021,1197,522,548]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 20891</specimenCode>
,
<specimenCode id="F4089E8AFFC8D431FB65FDDAFB66FD9A" box="[1209,1385,522,548]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21278</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FF61FDFEFA91FDF6" ID-GBIF-Occurrence="1806442496" box="[189,1182,557,584]" collectionCode="MUSM" country="Peru" location="Capitan Quebrada" pageId="13" pageNumber="14" specimenCode="MUSM 21278" specimenCount="2" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FF61FDFEFF41FDF9" box="[189,334,557,584]" pageId="13" pageNumber="14" type="generic">2 specimens</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FDCBFDFDFCCCFDF6" box="[535,707,557,584]" metricMagnitude="-2" metricUnit="m" metricValue="6.24" metricValueMax="6.87" metricValueMin="5.61" pageId="13" pageNumber="14" unit="mm" value="62.4" valueMax="68.7" valueMin="56.1">56.168.7 mm</quantity>
SL,
<location id="A171602AFFC8D431FD21FDFDFDDCFDF9" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FD21FDFDFDDCFDF9" box="[765,979,557,583]" country="Peru" name="Capitan Quebrada" pageId="13" pageNumber="14" stateProvince="Amazonas">Capitán Quebrada</location>
(
<specimenCode id="F4089E8AFFC8D431FC38FDFEFA99FDF6" box="[996,1174,558,584]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21278</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FB70FDFEFAA3FDD5" ID-GBIF-Occurrence="1806442456" collectionCode="MUSM" country="Peru" elevation="750" location="Quebrada Capitán Ponce" pageId="13" pageNumber="14" specimenCode="MUSM 21287" specimenCount="2" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FB70FDFEFB32FDF9" box="[1196,1341,557,584]" pageId="13" pageNumber="14" type="generic">2 specimens</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FE9FFD81FFE1FDD2" box="[323,494,593,620]" metricMagnitude="-2" metricUnit="m" metricValue="9.15" metricValueMax="9.709999999999999" metricValueMin="8.59" pageId="13" pageNumber="14" unit="mm" value="91.5" valueMax="97.1" valueMin="85.9">85.997.1 mm</quantity>
SL,
<location id="A171602AFFC8D431FDFBFD81FD41FDD2" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FDFBFD81FD41FDD2" box="[551,846,593,620]" country="Peru" name="Quebrada Capitán Ponce" pageId="13" pageNumber="14" stateProvince="Amazonas">Quebrada Capitán Ponce</location>
,
<elevation id="2F83D1C2FFC8D431FC86FD81FDECFDD5" box="[858,995,593,620]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="14" unit="m" value="750.0">
<quantity id="63569B14FFC8D431FC86FD81FDAAFDD2" box="[858,933,593,620]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="13" pageNumber="14" unit="m" value="750.0">750 m</quantity>
a.s.l.
</elevation>
(
<specimenCode id="F4089E8AFFC8D431FC2EFD81FAABFDD5" box="[1010,1188,593,619]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21287</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FB65FD81FDB4FD31" ID-GBIF-Occurrence="1806442537" collectionCode="MUSM" country="Peru" latitude="-3.9708333" location="Condorcanqui Province, Marañon River Basin, upper Cenepa River drainage" longLatPrecision="21" longitude="-78.6875" pageId="13" pageNumber="14" specimenCode="MUSM 21300" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FB65FD81FB31FDD5" box="[1209,1342,593,619]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FE9EFDA4FFA5FD31" box="[322,426,628,655]" metricMagnitude="-2" metricUnit="m" metricValue="8.57" pageId="13" pageNumber="14" unit="mm" value="85.7">85.7 mm</quantity>
SL, collected with holotype (
<specimenCode id="F4089E8AFFC8D431FCDEFDA5FDBBFD31" box="[770,948,629,655]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21300</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FC1BFDA5FAD6FD0C" ID-GBIF-Occurrence="1806442459" collectionCode="MUSM" country="Peru" latitude="-3.7781665" location="Capitan Ponce Bravo Quebrada" longLatPrecision="1" longitude="-78.33462" pageId="13" pageNumber="14" specimenCode="MUSM 21312" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FC1BFDA5FA44FD31" box="[967,1099,629,655]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FAD3FDA4FB78FD31" box="[1295,1399,628,655]" metricMagnitude="-2" metricUnit="m" metricValue="6.54" pageId="13" pageNumber="14" unit="mm" value="65.4">65.4 mm</quantity>
SL,
<location id="A171602AFFC8D431FF28FD48FC72FD0C" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FF28FD48FC72FD0C" box="[244,637,664,691]" country="Peru" latitude="-3.7781665" longLatPrecision="1" longitude="-78.33462" name="Capitan Ponce Bravo Quebrada" pageId="13" pageNumber="14" stateProvince="Amazonas">Capitan Ponce Bravo Quebrada</location>
,
<geoCoordinate id="C19A5036FFC8D431FD53FD46FD34FD0C" box="[655,827,662,690]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-3.7781665">3°4641.40″ S</geoCoordinate>
,
<geoCoordinate id="C19A5036FFC8D431FC93FD46FA0CFD0D" box="[847,1027,662,691]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-78.33462">78°204.61″ W</geoCoordinate>
(
<specimenCode id="F4089E8AFFC8D431FBC5FD48FADEFD0C" box="[1049,1233,664,690]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21312</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FB37FD48FB7DFD68" ID-GBIF-Occurrence="1806442462" collectionCode="MUSM" country="Peru" latitude="-3.640261" location="Quebrada Platanal" longLatPrecision="1" longitude="-72.31134" pageId="13" pageNumber="14" specimenCode="MUSM 21372" specimenCount="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FB37FD48FB78FD0C" box="[1259,1399,664,690]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
(sex unknown), 64.0 mm SL,
<location id="A171602AFFC8D431FDEFFD6CFD1DFD68" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FDEFFD6CFD1DFD68" box="[563,786,700,726]" country="Peru" latitude="-3.640261" longLatPrecision="1" longitude="-72.31134" name="Quebrada Platanal" pageId="13" pageNumber="14" stateProvince="Amazonas">Quebrada Platanal</location>
,
<geoCoordinate id="C19A5036FFC8D431FCFEFD6AFDC2FD68" box="[802,973,698,726]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-3.640261">3°3824.94″ S</geoCoordinate>
,
<geoCoordinate id="C19A5036FFC8D431FC02FD6AFAAFFD68" box="[990,1184,698,726]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-72.31134">72°1840.81″ W</geoCoordinate>
(
<specimenCode id="F4089E8AFFC8D431FB68FD6CFB65FD68" box="[1204,1386,700,726]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 21372</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FF61FD0FFF8FFCFF" ID-GBIF-Occurrence="1806442486" collectionCode="MUSM" country="Peru" county="Atalaya province" elevation="248" latitude="-11.237741" location="Lazaro Creek, tributary of Mishahua River, Ucayali River Basin" longLatPrecision="1" longitude="-72.97407" municipality="Sepahua" pageId="13" pageNumber="14" specimenCode="MUSM 35474" specimenCount="11" stateProvince="Ucayali" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FF61FD0FFF57FD47" box="[189,344,735,761]" pageId="13" pageNumber="14" type="generic">11 specimens</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FDCBFD0FFCB0FD44" box="[535,703,735,762]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" metricValueMax="4.93" metricValueMin="3.87" pageId="13" pageNumber="14" unit="mm" value="44.0" valueMax="49.3" valueMin="38.7">38.749.3 mm</quantity>
SL,
<collectingRegion id="666AF813FFC8D431FD2FFD0FFD41FD47" box="[755,846,735,761]" country="Peru" name="Ucayali" pageId="13" pageNumber="14">Ucayali</collectingRegion>
department,
<collectingCounty id="4D704E7DFFC8D431FC3EFD0FFAA3FD47" box="[994,1196,735,761]" pageId="13" pageNumber="14">Atalaya province</collectingCounty>
,
<collectingMunicipality id="4475AC8BFFC8D431FB69FD0FFB14FD47" box="[1205,1307,735,761]" pageId="13" pageNumber="14">Sepahua</collectingMunicipality>
,
<location id="A171602AFFC8D431FAF9FD30FD56FCA3" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FAF9FD30FD56FCA3" country="Peru" county="Atalaya province" latitude="-11.237741" longLatPrecision="1" longitude="-72.97407" municipality="Sepahua" name="Lazaro Creek, tributary of Mishahua River, Ucayali River Basin" pageId="13" pageNumber="14" stateProvince="Ucayali">Lazaro Creek, tributary of Mishahua River, Ucayali River Basin</location>
,
<geoCoordinate id="C19A5036FFC8D431FCB8FCD1FA17FCA3" box="[868,1048,769,797]" direction="south" orientation="latitude" pageId="13" pageNumber="14" precision="1" value="-11.237741">11°1415.87″ S</geoCoordinate>
,
<geoCoordinate id="C19A5036FFC8D431FBF9FCD1FAECFCA3" box="[1061,1251,769,797]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="1" value="-72.97407">72°5826.65″ W</geoCoordinate>
,
<elevation id="2F83D1C2FFC8D431FB32FCD3FB78FCA3" box="[1262,1399,771,797]" metricMagnitude="2" metricUnit="m" metricValue="2.48" pageId="13" pageNumber="14" unit="m" value="248.0">
<quantity id="63569B14FFC8D431FB32FCD3FB36FCA3" box="[1262,1337,771,797]" metricMagnitude="2" metricUnit="m" metricValue="2.48" pageId="13" pageNumber="14" unit="m" value="248.0">248 m</quantity>
a.s.l.
</elevation>
(
<specimenCode id="F4089E8AFFC8D431FF19FCF7FF77FCFF" box="[197,376,806,833]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 35474</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FE52FCF7FAA7FCDA" ID-GBIF-Occurrence="1806442542" collectionCode="MUSM" country="Peru" county="Huacamayo River" location="Huacamayo River, km. 155 on road from Pucallpa to Tingo-Maria, Padre Abad province, Ucayali River Basin" municipality="Pucallpa" pageId="13" pageNumber="14" specimenCode="MUSM 2392" specimenCount="1" stateProvince="Ucayali River" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FE52FCF7FC1AFCFE" box="[398,533,806,833]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FD3DFCF6FD44FCFF" box="[737,843,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" pageId="13" pageNumber="14" unit="mm" value="44.5">44.5 mm</quantity>
SL,
<location id="A171602AFFC8D431FC5BFCF7FDE4FCDA" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FC5BFCF7FDE4FCDA" country="Peru" county="Huacamayo River" municipality="Pucallpa" name="Huacamayo River, km. 155 on road from Pucallpa to Tingo-Maria, Padre Abad province, Ucayali River Basin" pageId="13" pageNumber="14" stateProvince="Ucayali River">Huacamayo River, km. 155 on road from Pucallpa to Tingo-Maria, Padre Abad province, Ucayali River Basin</location>
(
<specimenCode id="F4089E8AFFC8D431FC20FC9AFAAFFCDA" box="[1020,1184,842,868]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 2392</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FB6AFC9AFFD6FC36" ID-GBIF-Occurrence="1806442473" pageId="13" pageNumber="14" specimenCount="1">
<specimenCount id="B2A8FD78FFC8D431FB6AFC9AFB32FCDA" box="[1206,1341,842,868]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
(sex unknown),
<quantity id="63569B14FFC8D431FE9FFCBEFFA3FC36" box="[323,428,878,904]" metricMagnitude="-2" metricUnit="m" metricValue="9.39" pageId="13" pageNumber="14" unit="mm" value="93.9">93.9 mm</quantity>
SL
</materialsCitation>
,
<materialsCitation id="14C63CACFFC8D431FE39FCBEFD52FC12" ID-GBIF-Occurrence="1806442511" collectionCode="MUSM" country="Peru" location="Víbora Creek, tributary of Pisqui River" pageId="13" pageNumber="14" specimenCode="MUSM 46846" specimenCount="2" stateProvince="Loreto" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FE39FCBEFC79FC36" box="[485,630,878,904]" pageId="13" pageNumber="14" type="generic">2 specimens</specimenCount>
(sex unknown) C&amp;S, 59.0
<quantity id="63569B14FFC8D431FC1EFCBDFA25FC36" box="[962,1066,877,904]" metricMagnitude="-2" metricUnit="m" metricValue="6.12" pageId="13" pageNumber="14" unit="mm" value="61.2">61.2 mm</quantity>
SL,
<location id="A171602AFFC8D431FBBFFCBEFF7AFC15" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FBBFFCBEFF7AFC15" country="Peru" county="Vibora Creek" municipality="Pisqui River" name="Víbora Creek, tributary of Pisqui River" pageId="13" pageNumber="14" stateProvince="Loreto">Víbora Creek, tributary of Pisqui River</location>
,
<collectingRegion id="666AF813FFC8D431FE5FFC42FFDBFC12" box="[387,468,914,940]" country="Peru" name="Loreto" pageId="13" pageNumber="14">Loreto</collectingRegion>
,
<location id="A171602AFFC8D431FE3EFC42FC9FFC15" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FE3EFC42FC9FFC15" box="[482,656,913,940]" country="Peru" county="Vibora Creek" municipality="Pisqui River" name="upper Amazon" pageId="13" pageNumber="14" stateProvince="Loreto">upper Amazon</location>
(
<specimenCode id="F4089E8AFFC8D431FD7EFC41FD59FC12" box="[674,854,913,940]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 46846</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="14C63CACFFC8D431FCB0FC41FFF9FC4D" ID-GBIF-Occurrence="1806442509" collectionCode="MUSM" country="Peru" county="Tavara River" location="Sandia, Zona Reservada Tambopata- Candamo" municipality="Quebrada Grande" pageId="13" pageNumber="14" specimenCode="MUSM 46847" specimenCount="3" stateProvince="Puno" typeStatus="paratype">
<specimenCount id="B2A8FD78FFC8D431FCB0FC41FDF1FC15" box="[876,1022,913,939]" pageId="13" pageNumber="14" type="generic">3 specimens</specimenCount>
(sex unknown), 50.0
<quantity id="63569B14FFC8D431FAD0FC41FB78FC12" box="[1292,1399,913,940]" metricMagnitude="-2" metricUnit="m" metricValue="9.86" pageId="13" pageNumber="14" unit="mm" value="98.6">98.6 mm</quantity>
SL,
<locationDeviation id="B5A32FD3FFC8D431FF33FC65FD69FC71" box="[239,870,949,975]" pageId="13" pageNumber="14">
Tavara River
<quantity id="63569B14FFC8D431FE4FFC65FFDFFC71" box="[403,464,949,975]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="km" value="2.0">2 km</quantity>
from mouth of Quebrada Grande
</locationDeviation>
,
<collectingRegion id="666AF813FFC8D431FCA9FC65FDBBFC71" box="[885,948,949,975]" country="Peru" name="Puno" pageId="13" pageNumber="14">Puno</collectingRegion>
,
<location id="A171602AFFC8D431FC1EFC65FF22FC4C" LSID="urn:lsid:plazi:treatment:2C0787E7FFC9D432FDA9FAF8FA2FFAB0:A171602AFFC8D431FC1EFC65FF22FC4C" country="Peru" county="Tavara River" municipality="Quebrada Grande" name="Sandia, Zona Reservada Tambopata- Candamo" pageId="13" pageNumber="14" stateProvince="Puno">Sandia, Zona Reservada Tambopata- Candamo</location>
(
<specimenCode id="F4089E8AFFC8D431FEE1FC08FFE1FC4D" box="[317,494,984,1011]" collectionCode="MUSM" pageId="13" pageNumber="14">MUSM 46847</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFC8D432FF61FBCAFD52FBC4" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="A41136F1FFC8D431FF61FBCAFF41FB8A" blockId="13.[189,334,1050,1076]" box="[189,334,1050,1076]" pageId="13" pageNumber="14">
<heading id="FF59819DFFC8D431FF61FBCAFF41FB8A" bold="true" box="[189,334,1050,1076]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="96DAEAE3FFC8D431FF61FBCAFF41FB8A" bold="true" box="[189,334,1050,1076]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFC8D431FF61FB99FC6DFB4F" blockId="13.[189,1399,1097,1266]" pageId="13" pageNumber="14">Body compressed, greatest body depth at or anterior to dorsal-fin origin. Mouth terminal. Dorsal profile of head straight between snout tip and posterior margin of supraoccipital spine, convex between head and dorsal fin, convex between head and at base of dorsal fin, convex between last dorsal-fin ray to adipose-fin origin. Dorsal and ventral caudal-peduncle margins straight. Ventral profile convex between tip of snout and pelvic-fin insertion.</paragraph>
<paragraph id="A41136F1FFC8D431FF61FAC9FAABFA7F" blockId="13.[189,1398,1305,1474]" pageId="13" pageNumber="14">Premaxillary teeth in two series; outer series with four tricuspid teeth covering three medial most teeth of inner series; inner row with five pentacuspid teeth. Maxilla long, of same width along entire length, with 36 tricuspid teeth set in anterior-most part of ventral margin. Dentary with anterior four teeth pentacuspid, followed laterally by 89 smaller teeth that progressively increase in postero-medial inclination, decreasing from tri- to unicuspid; number of lateral teeth highly variable.</paragraph>
<paragraph id="A41136F1FFC8D431FF61FA39FD1FF942" blockId="13.[189,1399,1513,1789]" pageId="13" pageNumber="14">
Pored scales of lateral line 40(15), 41(3), 42(1) (n = 18), scales from lateral line to origin of dorsal fin 8(18) (n = 18), scales from lateral line to origin of anal fin 7(10), 8(8) (n = 18), scales from lateral line to insertion of pelvic fin 6(11), 7(7) (n = 18). Rays of dorsal fin
<date id="D0101031FFC8D431FC57F9E0FDB3F9F5" box="[907,956,1584,1611]" pageId="13" pageNumber="14">iii 9</date>
, first simple ray small, only visible in cleared and stained specimens, second simple ray about half length of third simple ray. Distal margin of dorsal fin slightly convex. Origin of adipose fin anterior to vertical through insertion of last ray of anal fin. Rays of pectoral fin
<date id="D0101031FFC8D431FE07F94BFC07F90B" box="[475,520,1691,1717]" pageId="13" pageNumber="14" value="2010-01">i 10</date>
ii (11),
<date id="D0101031FFC8D431FDBBF94BFC9BF90B" box="[615,660,1691,1717]" pageId="13" pageNumber="14" value="2012-01">i 12</date>
ii (7). Rays of pelvic fin
<date id="D0101031FFC8D431FC62F94BFDD1F90B" box="[958,990,1691,1717]" pageId="13" pageNumber="14">i 7</date>
(18). Rays of anal fin
<date id="D0101031FFC8D431FB03F94BFB7EF908" box="[1247,1393,1691,1718]" pageId="13" pageNumber="14" value="1925-03-27" valueMax="1925-04-27" valueMin="1925-03-27">iiiiv 2527</date>
; first simple rays only visible in cleared and stained material. Origin of anal fin posterior to vertical through insertion of last ray of dorsal fin (
<tableCitation id="E92C034AFFC8D431FD7BF932FCF0F942" box="[679,767,1762,1788]" captionStart="Table 1" captionStartId="11.[145,214,599,625]" captionTargetPageId="11" captionText="Table 1. Morphometric data for some members of the Astyanax orthodus species-group." httpUri="http://table.plazi.org/id/F0D16679FFCED437FF4DFD87FA92FDCF" pageId="13" pageNumber="14" tableUuid="F0D16679FFCED437FF4DFD87FA92FDCF">Table 1</tableCitation>
).
</paragraph>
<paragraph id="A41136F1FFC8D431FF61F8F4FC8FF83B" blockId="13.[189,1399,1828,1925]" pageId="13" pageNumber="14">Caudal fin with 109 principal rays; dorsal lobe supporting 12(4), procurrent rays; ventral lobe, 10(4) procurrent rays. Upper 10(4) principal caudal-fin rays associated with four dorsal hypurals, next 9(4) associated with three ventral hypurals.</paragraph>
<paragraph id="A41136F1FFC8D432FF61F87DFAEDFEF6" blockId="13.[189,1399,1965,2027]" lastBlockId="14.[189,1399,267,329]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">Total vertebrae 37(4), including those of Weberian apparatus: precaudal centra 17(2) and 18(2), last two without true pleural rib. Caudal centra 19(2), 20 (2). Epineurals 34(2); posterior-most epineural occasionally not reaching anterior surface of urostyle; epipleurals 21(2). Caudal skeleton with seven hypurals. First dorsal three hypurals with swollen anterior margin in contact with urostyle.</paragraph>
<paragraph id="A41136F1FFCBD432FF61FEA0FFE9FE34" blockId="14.[189,1399,368,581]" box="[189,486,368,394]" pageId="14" pageNumber="15">
<heading id="FF59819DFFCBD432FF61FEA0FFE9FE34" bold="true" box="[189,486,368,394]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="96DAEAE3FFCBD432FF61FEA0FFE9FE34" bold="true" box="[189,486,368,394]" pageId="14" pageNumber="15">Pigmentation in alcohol</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFCBD432FF61FE4EFA1DFDFB" blockId="14.[189,1399,368,581]" pageId="14" pageNumber="15">Sides of body yellowish, without reticulated pattern over dorsal region of coelomic cavity, silvery stripe extending from humeral region to base of caudal peduncle, overlain by series of chevron-shaped marks formed by dark lines along myosepta between myotomes extending from dorsal region of coelomic cavity to caudal peduncle; pigmented muscle septae forming chevrons not coinciding with scale rows. Chevrons lacking distal extensions both in juvenile and adult specimens.</paragraph>
<paragraph id="A41136F1FFCBD432FF61FDBDFB78FBB4" blockId="14.[189,1399,621,1034]" pageId="14" pageNumber="15">
Dorsal region of head and body chestnut brown. Sides of cranium and ventral surface of body light brown, not silvery. Melanophores of humeral region forming two (anterior and posterior) humeral spots. The anterior spot formed by two layers of pigment, with brown melanophores distributed in thin superficial layer of epithelium (Layer 1,
<figureCitation id="3C952A74FFCBD432FDCEFD06FC5BFD4E" box="[530,596,726,752]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
) and another deeper layer consisting of dark melanophores (Layer 2,
<figureCitation id="3C952A74FFCBD432FF61FD29FF0EFCAA" box="[189,257,761,788]" captionStart="Fig.1" captionStartId="4.[189,232,1859,1885]" captionTargetBox="[189,1398,540,1837]" captionTargetId="figure@4.[189,1398,540,1837]" captionTargetPageId="4" captionText="Fig.1.A.DistributionofpigmentinthehumeralregionofAstyanaxspecies.B.A.bimaculatus(Linnaeus, 1758).C.A.caucanus(Steindachner,1879).D.A.mexicanus(DeFilippi,1853).E.A.courensisBertaco, Carvalho &amp; Jerep, 2010 (modified from Bertaco et al. 2010). The anterior humeral spot consists of two overlapping layers of pigment identified by numbers 1 and 2 (see Román-Valencia et al. 2015: fig. 1): 1. Diffuse vertical bar. 2. Horizontal spot with concentrated pigments." httpUri="https://zenodo.org/record/1169561/files/figure.png" pageId="14" pageNumber="15">Fig. 1</figureCitation>
). Layer 2 forming polygon-shaped spot made up of two different layers of melanophores that do not precisely overlap, forming vertices on lateral margins, usually resulting in four-sided spot located between third and sixth or seventh scale of lateral series. Posterior humeral spot situated two or three scales posterior to the anterior humeral spot, arc- or sigmoid-shaped, inconspicuous, covering two to three scales above lateral-line. Individual scales on sides of body lacking spots or dots. Spot on caudal peduncle absent. Lateral stripe formed by dispersed brown pigment present only on posterior portion of body, located above posterior two-thirds of anal-fin base and extending on to caudal peduncle. Pectoral, pelvic, dorsal and anal fins hyaline; pigment present on interradial membranes of middle caudal-fin rays.
</paragraph>
<paragraph id="A41136F1FFCBD432FF61FBE1FFBFFBF5" blockId="14.[189,432,1073,1099]" box="[189,432,1073,1099]" pageId="14" pageNumber="15">
<heading id="FF59819DFFCBD432FF61FBE1FFBFFBF5" bold="true" box="[189,432,1073,1099]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="96DAEAE3FFCBD432FF61FBE1FFBFFBF5" bold="true" box="[189,432,1073,1099]" pageId="14" pageNumber="15">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFCBD432FF61FBB0FD52FBC4" blockId="14.[189,861,1120,1146]" box="[189,861,1120,1146]" pageId="14" pageNumber="15">No sexual dimorphism observed, no hooks found on fins.</paragraph>
</subSubSection>
<subSubSection id="ECB4657AFFCBD432FF61FB71FA2FFAB0" pageId="14" pageNumber="15" type="distribution">
<paragraph id="A41136F1FFCBD432FF61FB71FF59FB05" blockId="14.[189,342,1185,1211]" box="[189,342,1185,1211]" pageId="14" pageNumber="15">
<heading id="FF59819DFFCBD432FF61FB71FF59FB05" bold="true" box="[189,342,1185,1211]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="96DAEAE3FFCBD432FF61FB71FF59FB05" bold="true" box="[189,342,1185,1211]" pageId="14" pageNumber="15">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="A41136F1FFCBD432FF61FB00FA2FFAB0" blockId="14.[189,1398,1232,1294]" pageId="14" pageNumber="15">
<taxonomicName id="63AE4D72FFCBD432FF61FB00FFAFFB54" authority="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega, 2018" authorityName="Ruiz-C &amp; Román-Valencia &amp; Taphorn &amp; Buckup &amp; Ortega" authorityYear="2018" box="[189,416,1232,1258]" class="Actinopterygii" family="Characidae" genus="Astyanax" kingdom="Animalia" order="Characiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="gandhiae" status="sp. nov.">
<emphasis id="96DAEAE3FFCBD432FF61FB00FFAFFB54" box="[189,416,1232,1258]" italics="true" pageId="14" pageNumber="15">Astyanax gandhiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8DE95798FFCBD432FE76FB00FC08FB54" box="[426,519,1232,1258]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
is known from the
<collectingRegion id="666AF813FFCBD432FD26FB00FD5AFB54" box="[762,853,1232,1258]" country="Peru" name="Ucayali" pageId="14" pageNumber="15">Ucayali</collectingRegion>
and
<collectingRegion id="666AF813FFCBD432FC4FFB00FA48FB54" box="[915,1095,1232,1258]" country="Peru" name="Madre de Dios" pageId="14" pageNumber="15">Madre de Dios</collectingRegion>
River drainages, Cenepa River, tributary from
<collectingRegion id="666AF813FFCBD432FE64FB23FC2DFAB3" box="[440,546,1267,1293]" country="Brazil" name="Maranhao" pageId="14" pageNumber="15">Marañón</collectingRegion>
River, Upper Amazon Basin,
<collectingCountry id="DCB97661FFCBD432FC5FFB24FDB6FAB0" box="[899,953,1268,1294]" name="Peru" pageId="14" pageNumber="15">Peru</collectingCountry>
(
<figureCitation id="3C952A74FFCBD432FC16FB23FA00FAB3" box="[970,1039,1267,1293]" captionStart="Fig. 4" captionStartId="8.[189,232,2000,2026]" captionTargetBox="[189,1392,764,1921]" captionTargetId="figure@8.[151,1417,755,1932]" captionTargetPageId="8" captionText="Fig. 4.Distribution of speciesof the Astyanaxorthodus species-group(symbol includestypelocalities)." httpUri="https://zenodo.org/record/1169567/files/figure.png" pageId="14" pageNumber="15">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>