260 lines
30 KiB
XML
260 lines
30 KiB
XML
<document id="94D08DE3C0014E9D13E0187E359128EC" ID-DOI="10.5281/zenodo.212975" ID-GBIF-Dataset="d10859be-023f-4fca-a164-6836331e5fa6" ID-ISSN="1175-5326" ID-Zenodo-Dep="212975" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460504029448" checkinUser="plazi" docAuthor="Schoville, Sean D." docDate="2012" docId="222787CCFFB1FF98838354918DC436B7" docLanguage="en" docName="zt03412p052.pdf" docOrigin="Zootaxa 3412" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Grylloblatta siskiyouensis Schoville, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="48" masterDocId="DE1EFFB4FFB4FF9E831452078F063171" masterDocTitle="Three new species of Grylloblatta Walker (Insecta: Grylloblattodea: Grylloblattidae), from southern Oregon and northern California" masterLastPageNumber="52" masterPageNumber="42" pageNumber="47" updateTime="1698312492540" updateUser="plazi">
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<mods:titleInfo id="340B654224C69D78D62F81344E32016D">
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<mods:title id="DF00E9DE36F605811887EB78B12420A3">Three new species of Grylloblatta Walker (Insecta: Grylloblattodea: Grylloblattidae), from southern Oregon and northern California</mods:title>
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</mods:titleInfo>
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<mods:name id="F2C4A8F0EC1062E03EBA293FC0BA7241" type="personal">
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<mods:roleTerm id="D9D8DC4AD70DA37DA3F961E246EEE200">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B4167FD3F0B22C447B2070DA49E27A62">Schoville, Sean D.</mods:namePart>
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<mods:typeOfResource id="137EA6E43EAE55D14A7B1699F738BB86">text</mods:typeOfResource>
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<mods:title id="78A174540C8E3F574AF3706B102C62F9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A5D9952C0ED4AC9FF70A972C2BACC31C">
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<mods:date id="60DDA6E73E1A5196876402E7BD5D1A29">2012</mods:date>
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<mods:detail id="FE82FE481D49ADCE6EE52E0216D8E5A4" type="volume">
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<mods:number id="CF5C67EC5E758886B76D7BF1E6CAE756">3412</mods:number>
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<mods:start id="FFE532737DED33BE3C5EB3D6B66BFFD3">42</mods:start>
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<mods:classification id="D13F7CA6F7B9C5A097BB73E7721110A4">journal article</mods:classification>
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<mods:identifier id="DE2D4B79837066F36FAB85B6075AA650" type="DOI">10.5281/zenodo.212975</mods:identifier>
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<mods:identifier id="746F5143DBB27B65DBB624E3891B3AAC" type="GBIF-Dataset">d10859be-023f-4fca-a164-6836331e5fa6</mods:identifier>
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<mods:identifier id="73DEAD4E1A45968B9717B8F9A47680B9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="ADD278586FFB701D76C23CF4C56DC40B" type="Zenodo-Dep">212975</mods:identifier>
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<treatment id="222787CCFFB1FF98838354918DC436B7" ID-DOI="http://doi.org/10.5281/zenodo.5673769" ID-GBIF-Taxon="119522303" ID-Zenodo-Dep="5673769" LSID="urn:lsid:plazi:treatment:222787CCFFB1FF98838354918DC436B7" httpUri="http://treatment.plazi.org/id/222787CCFFB1FF98838354918DC436B7" lastPageId="6" lastPageNumber="48" pageId="5" pageNumber="47">
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<subSubSection id="E2946551FFB1FF9B838354918E5F37A0" pageId="5" pageNumber="47" type="nomenclature">
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<paragraph id="AA3136DAFFB1FF9B838354918D6537C1" blockId="5.[151,611,1686,1745]" box="[151,611,1686,1712]" pageId="5" pageNumber="47">
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<heading id="F17981B6FFB1FF9B838354918D6537C1" bold="true" box="[151,611,1686,1712]" fontSize="11" level="1" pageId="5" pageNumber="47" reason="1">
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<emphasis id="98FAEAC8FFB1FF9B838354918D6537C1" bold="true" box="[151,611,1686,1712]" pageId="5" pageNumber="47">
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<taxonomicName id="6D8E4D59FFB1FF9B838354918ECB37C1" ID-CoL="3HG2V" box="[151,461,1686,1712]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
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<emphasis id="98FAEAC8FFB1FF9B838354918ECB37C1" bold="true" box="[151,461,1686,1712]" italics="true" pageId="5" pageNumber="47">Grylloblatta siskiyouensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="83C957B3FFB1FF9B82C054918D6537C1" box="[468,611,1686,1712]" pageId="5" pageNumber="47" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="AA3136DAFFB1FF9B838354BE8E5F37A0" blockId="5.[151,611,1686,1745]" box="[151,345,1721,1745]" pageId="5" pageNumber="47">
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(
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<figureCitation id="32B52A5FFFB1FF9B838B54BE8FEF37A0" box="[159,233,1721,1745]" captionStart="FIGURE 1" captionStartId="2.[151,250,1409,1432]" captionTargetBox="[182,1410,627,1381]" captionTargetId="figure@2.[176,1411,625,1388]" captionTargetPageId="2" captionText="FIGURE 1. Differences in size and coloration observed in type specimens. A) Grylloblatta oregonensis n. sp. B) G. siskiyouensis n. sp. C) G. m a r m o re u s n. sp." httpUri="https://zenodo.org/record/212976/files/figure.png" pageId="5" pageNumber="47">Figs. 1</figureCitation>
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B, 2C–D)
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFB1FF98838355058FD8306D" lastPageId="6" lastPageNumber="48" pageId="5" pageNumber="47" type="materials_examined">
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<paragraph id="AA3136DAFFB1FF9B838355058FE236DD" blockId="5.[151,1437,1793,2036]" pageId="5" pageNumber="47">
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<emphasis id="98FAEAC8FFB1FF9B838355058E42366B" bold="true" box="[151,324,1793,1818]" pageId="5" pageNumber="47">
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<typeStatus id="75358878FFB1FF9B838355058FD6366B" box="[151,208,1794,1818]" pageId="5" pageNumber="47">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="75358878FFB1FF9B825A55068EB03668" box="[334,438,1793,1817]" pageId="5" pageNumber="47" type="holotype">Holotype</typeStatus>
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female in ethanol, deposited in
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<collectionCode id="CC9FAE1FFFB1FF9B803055068C5B3668" LSID="urn:lsid:biocol.org:col:34700" box="[804,861,1793,1817]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="47">CAS</collectionCode>
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, labeled: “
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<collectingCountry id="D299764AFFB1FF9B80C055068B083668" box="[980,1038,1793,1817]" name="United States of America" pageId="5" pageNumber="47">USA</collectingCountry>
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OR: Josephine Co., Oregon Caves N.M., Big Tree Loop Trail
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<geoCoordinate id="CFBA501DFFB1FF9B82DC55228D2F364C" box="[456,553,1829,1853]" direction="north" orientation="latitude" pageId="5" pageNumber="47" precision="55" value="42.098">N42.098</geoCoordinate>
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<geoCoordinate id="CFBA501DFFB1FF9B812455228DA8364C" box="[560,686,1829,1853]" direction="west" orientation="longitude" pageId="5" pageNumber="47" precision="55" value="-123.407">W-123.407</geoCoordinate>
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WGS84, Note: near water station
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<date id="DE30101AFFB1FF9B872455228BB5364C" box="[1072,1203,1829,1853]" pageId="5" pageNumber="47" value="2007-03-06">06-III-2007</date>
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9:15pm AY SDS10- 004” [white label]/ “
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<typeStatus id="75358878FFB1FF9B8295554E8D113610" box="[385,535,1865,1889]" pageId="5" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="6D8E4D59FFB1FF9B8134554E8CBE3610" authority="Schoville" authorityName="Schoville" box="[544,952,1865,1889]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="5" pageNumber="47" phylum="Arthropoda" rank="species" species="siskiyouensis">
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<emphasis id="98FAEAC8FFB1FF9B8134554E8C403610" box="[544,838,1865,1889]" italics="true" pageId="5" pageNumber="47">Grylloblatta siskiyouensis</emphasis>
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Schoville
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</taxonomicName>
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det. S.D. Schoville 2011” [red label]; Big Tree Trail of Oregon Caves National Monument,
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<geoCoordinate id="CFBA501DFFB1FF9B81C055778C7536F9" box="[724,883,1901,1928]" direction="north" orientation="latitude" pageId="5" pageNumber="47" precision="55" value="42.098">42.098o North</geoCoordinate>
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and -
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<geoCoordinate id="CFBA501DFFB1FF9B80A255778B5C36F9" box="[950,1114,1901,1928]" direction="west" orientation="longitude" pageId="5" pageNumber="47" precision="55" value="-123.407">123.407o West</geoCoordinate>
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, Josephine County, Oregon,
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<collectingCountry id="D299764AFFB1FF9B838355938FE236DD" box="[151,228,1940,1964]" name="United States of America" pageId="5" pageNumber="47">U.S.A.</collectingCountry>
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</paragraph>
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<paragraph id="AA3136DAFFB1FF9883D355B08FD8306D" blockId="5.[151,1437,1793,2036]" lastBlockId="6.[151,1437,151,1991]" lastPageId="6" lastPageNumber="48" pageId="5" pageNumber="47">
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<emphasis id="98FAEAC8FFB1FF9B83D355B08E9E36A1" bold="true" box="[199,408,1975,2000]" pageId="5" pageNumber="47">Other specimens.</emphasis>
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Three juveniles in ethanol, deposited in
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<collectionCode id="CC9FAE1FFFB1FF9B807855BF8CA336A1" LSID="urn:lsid:biocol.org:col:34700" box="[876,933,1976,2000]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="47">CAS</collectionCode>
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, labeled: “
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<collectingCountry id="D299764AFFB1FF9B870A55BF8B5E36A1" box="[1054,1112,1976,2000]" name="United States of America" pageId="5" pageNumber="47">USA</collectingCountry>
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OR: Josephine Co., Oregon Caves N.M., Blind Leads Cave,
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<geoCoordinate id="CFBA501DFFB1FF9B82EB55DB8D663685" box="[511,608,2012,2036]" direction="north" orientation="latitude" pageId="5" pageNumber="47" precision="55" value="42.098">N42.098</geoCoordinate>
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||
<geoCoordinate id="CFBA501DFFB1FF9B817355DB8DE23685" box="[615,740,2012,2036]" direction="west" orientation="longitude" pageId="5" pageNumber="47" precision="55" value="-123.407">W-123.407</geoCoordinate>
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WGS84,
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<date id="DE30101AFFB1FF9B804655DB8CDE3685" box="[850,984,2012,2036]" pageId="5" pageNumber="47" value="2010-03-20">20-III-2010</date>
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. Note: twilight zone under rocks” [white label]/ “
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<taxonomicName id="6D8E4D59FFB2FF9883E2529F8D9431DE" authority="Schoville" authorityName="Schoville" box="[246,658,151,176]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis">
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||
<emphasis id="98FAEAC8FFB2FF9883E2529F8D1B31C1" box="[246,541,152,176]" italics="true" pageId="6" pageNumber="48">Grylloblatta siskiyouensis</emphasis>
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Schoville
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</taxonomicName>
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||
det. S.D. Schoville 2011” [white label]. Two juveniles in ethanol, deposited in
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<collectionCode id="CC9FAE1FFFB2FF98823D52BB8E6431A5" LSID="urn:lsid:biocol.org:col:34700" box="[297,354,188,212]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="6" pageNumber="48">CAS</collectionCode>
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||
, labeled: “
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<collectingCountry id="D299764AFFB2FF9882CF52BB8D1331A5" box="[475,533,188,212]" name="United States of America" pageId="6" pageNumber="48">USA</collectingCountry>
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OR: Josephine Co., Oregon Caves N.M.,
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<geoCoordinate id="CFBA501DFFB2FF9880E352BB8B5E31A5" box="[1015,1112,188,212]" direction="north" orientation="latitude" pageId="6" pageNumber="48" precision="55" value="42.098">N42.098</geoCoordinate>
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||
<geoCoordinate id="CFBA501DFFB2FF98877552BB8BD931A5" box="[1121,1247,188,212]" direction="west" orientation="longitude" pageId="6" pageNumber="48" precision="55" value="-123.407">W-123.407</geoCoordinate>
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WGS84,
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<date id="DE30101AFFB2FF98864552BB8FD53186" pageId="6" pageNumber="48" value="2010-03-20">20-III- 2010</date>
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. Note: on snow surface” [white label]/ “
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<taxonomicName id="6D8E4D59FFB2FF98818152E78B2F3186" authority="Schoville" authorityName="Schoville" box="[661,1065,223,248]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis">
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<emphasis id="98FAEAC8FFB2FF98818152E78CBE3189" box="[661,952,224,248]" italics="true" pageId="6" pageNumber="48">Grylloblatta siskiyouensis</emphasis>
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Schoville
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</taxonomicName>
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det. S.D. Schoville 2011” [white label].
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</paragraph>
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</subSubSection>
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<subSubSection id="E2946551FFB2FF9883D3532F8DA63015" pageId="6" pageNumber="48" type="etymology">
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<paragraph id="AA3136DAFFB2FF9883D3532F8DA63015" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
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<emphasis id="98FAEAC8FFB2FF9883D3532F8E483031" bold="true" box="[199,334,296,320]" pageId="6" pageNumber="48">Etymology.</emphasis>
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The specific epithet,
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<taxonomicName id="6D8E4D59FFB2FF988128532F8DC83031" box="[572,718,296,320]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis">
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<emphasis id="98FAEAC8FFB2FF988128532F8DC83031" box="[572,718,296,320]" italics="true" pageId="6" pageNumber="48">siskiyouensis</emphasis>
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</taxonomicName>
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, is named for the Siskiyou Mountains in which the
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<typeStatus id="75358878FFB2FF988618532F8A3A3031" box="[1292,1340,296,320]" pageId="6" pageNumber="48">type</typeStatus>
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locality, Oregon Caves National Monument, is located.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="E2946551FFB2FF9883D353688EDB332E" pageId="6" pageNumber="48" type="diagnosis">
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<paragraph id="AA3136DAFFB2FF9883D353688EDB332E" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
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||
<emphasis id="98FAEAC8FFB2FF9883D353688E4730F9" bold="true" box="[199,321,367,392]" pageId="6" pageNumber="48">Diagnosis.</emphasis>
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Adult females of this species can be distinguished from all other Nearctic
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<taxonomicName id="6D8E4D59FFB2FF98878053778A1830F9" box="[1172,1310,368,392]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="98FAEAC8FFB2FF98878053778A1830F9" box="[1172,1310,368,392]" italics="true" pageId="6" pageNumber="48">Grylloblatta</emphasis>
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</taxonomicName>
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species by the following combination of character states: color amber brown on dorsum and white ventrally with light amber yellow patches; small body size; more than 30 antennomeres; ratio of antennomere lengths 2 and 1 1.2; head and interocular distance wider than pronotum; compound eyes large; pronotum short and narrow with a notable point formed at posterior edge; ventral valve of ovipositor broad and slightly up-curved; dorsal valve of ovipositor projects beyond ventral valve one-fifth total length; ovipositor nearly as long as cercus; cercomeres short, distally brown and proximally white.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E2946551FFB2FF9883D3506C8CA43501" pageId="6" pageNumber="48" type="description">
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||
<paragraph id="AA3136DAFFB2FF9883D3506C8B923295" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
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||
<emphasis id="98FAEAC8FFB2FF9883D3506C8D4433F5" bold="true" box="[199,578,619,644]" pageId="6" pageNumber="48">
|
||
Description of
|
||
<typeStatus id="75358878FFB2FF98826F506B8EE433F5" box="[379,482,620,644]" pageId="6" pageNumber="48" type="holotype">holotype</typeStatus>
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female.
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</emphasis>
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Antennomere number on right antennae 32 and left antennae (broken) 19. Compound eyes well developed, large and black. Interocular distance slightly larger than pronotum width. Head width slightly greater than head length. Pronotum short and narrow, slightly convex, anterior end wider than posterior end, transverse sulcus at anterior end curving slightly towards the posterior end, a weak mid-dorsal line visible, and a notable point formed at the posterior edge. Leg coloration distinctly yellow compared to dorsal body and abdomen, and brown setae distinctive. Ovipositor nearly as long as cercus, extending to eighth cercomere. Ventral valve of ovipositor broad in the middle and slightly up-curved, roughened distally (possible scarring), with noticeable long setae. Dorsal valves of ovipositor extend beyond ventral valve by one-fifth total length and at midpoint nearly straighten out. Cercal segments short, distally brown and proximally white. Dorsal body color amber brown (55), legs light amber yellow (83), abdomen white laterally and light amber yellow patches ventrally, antennomeres and cercomeres amber brown fading to white at distal ends, ovipositor salmon-orange (50).
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</paragraph>
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<paragraph id="AA3136DAFFB2FF9883D351E88CA43501" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
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||
<emphasis id="98FAEAC8FFB2FF9883D351E88D9D3576" bold="true" box="[199,667,1006,1031]" pageId="6" pageNumber="48">
|
||
Measurements of
|
||
<typeStatus id="75358878FFB2FF98828A51E88D033576" box="[414,517,1007,1031]" pageId="6" pageNumber="48" type="holotype">holotype</typeStatus>
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female (cm)
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</emphasis>
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. BL 1.295, HL 0.294, HW 0.326, EL 0.071, EW 0.0476, ID 0.223, Ant1L 0.0397, Ant2L 0.0476, AntRatio 1.2, PL 0.3176, PW 0.183, FL front 0.333, FL mid 0.397, FL rear 0.508, TL front 0.238, TL mid 0.349, TL rear 0.453, TrL front 0.071, TrL mid 0.103, TrL rear 0.175, PTrL front 0.270, PTrL mid 0.318, PTrL rear 0.318, OvL 0.437, OvW 0.135, CL 0.476.
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="E2946551FFB2FF9883D3567C8DC436B7" pageId="6" pageNumber="48" type="materials_examined">
|
||
<paragraph id="AA3136DAFFB2FF9883D3567C8FCC35C6" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
|
||
<emphasis id="98FAEAC8FFB2FF9883D3567C8D3835E5" bold="true" box="[199,574,1147,1172]" pageId="6" pageNumber="48">Description of other specimens.</emphasis>
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Juveniles are amber brown (55) on the dorsal surface of the head and pronotum.
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||
</paragraph>
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||
<paragraph id="AA3136DAFFB2FF9883D356C58A9B3766" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
|
||
<emphasis id="98FAEAC8FFB2FF9883D356C58E6035AA" bold="true" box="[199,358,1218,1243]" pageId="6" pageNumber="48">Comparisons</emphasis>
|
||
.
|
||
<taxonomicName id="6D8E4D59FFB2FF98826756C58D9035AB" box="[371,662,1218,1242]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFB2FF98826756C58D9035AB" box="[371,662,1218,1242]" italics="true" pageId="6" pageNumber="48">Grylloblatta siskiyouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFB2FF98818856C48DDD35AA" box="[668,731,1219,1243]" pageId="6" pageNumber="48" rank="species">
|
||
<emphasis id="98FAEAC8FFB2FF98818856C48DDD35AA" bold="true" box="[668,731,1219,1243]" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from all
|
||
<taxonomicName id="6D8E4D59FFB2FF98873F56C58BB035AB" box="[1067,1206,1218,1242]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="98FAEAC8FFB2FF98873F56C58BB035AB" box="[1067,1206,1218,1242]" italics="true" pageId="6" pageNumber="48">Grylloblatta</emphasis>
|
||
</taxonomicName>
|
||
species by dark dorsal coloration and dark abdominal color patches, as well as notable point on posterior end of pronotum. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFB2FF988265570E8D103450" box="[369,534,1289,1313]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="oregonensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFB2FF988265570E8D103450" box="[369,534,1289,1313]" italics="true" pageId="6" pageNumber="48">G. oregonensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFB2FF98810A570E8D5B3450" box="[542,605,1289,1313]" pageId="6" pageNumber="48" rank="species">
|
||
<emphasis id="98FAEAC8FFB2FF98810A570E8D5B3450" bold="true" box="[542,605,1289,1313]" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by a smaller adult body size and limb length, notably narrower pronotum and head, antennomere count higher (>30) and ratio of antennomere 2:1 lower (<2.0), dorsal valve of ovipositor thicker and slightly more up-curved. It can be distinguished from
|
||
<emphasis id="98FAEAC8FFB2FF98806057578CC23416" box="[884,964,1359,1383]" italics="true" pageId="6" pageNumber="48">G. ro t h i</emphasis>
|
||
by a smaller body size, higher antennomere count (>30), and long, setaceous ovipositor. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFB2FF9880E757748B6F34FB" box="[1011,1129,1394,1418]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="sculleni">
|
||
<emphasis id="98FAEAC8FFB2FF9880E757748B6F34FB" box="[1011,1129,1394,1418]" italics="true" pageId="6" pageNumber="48">G. sculleni</emphasis>
|
||
</taxonomicName>
|
||
by a cercus nearly equal in length to ovipositor. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFB2FF9881D257918C3F34DF" box="[710,825,1430,1454]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="barberi">
|
||
<emphasis id="98FAEAC8FFB2FF9881D257918C3F34DF" box="[710,825,1430,1454]" italics="true" pageId="6" pageNumber="48">G. barberi</emphasis>
|
||
</taxonomicName>
|
||
by more reddish dorsal color, narrower head and pronotum, fewer antennomeres, a cercus nearly equal in length to ovipositor, and a less up-curved dorsal valve of ovipositor. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFB2FF98812457DA8DA23485" box="[560,676,1501,1524]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="gurneyi">
|
||
<emphasis id="98FAEAC8FFB2FF98812457DA8DA23485" box="[560,676,1501,1524]" italics="true" pageId="6" pageNumber="48">G. gurneyi</emphasis>
|
||
</taxonomicName>
|
||
in having fewer antennomeres and ovipositor nearly equal in length to cercus. It can be distinguished from
|
||
<taxonomicName id="6D8E4D59FFB2FF98815C54078DD73766" box="[584,721,1535,1559]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="chandleri">
|
||
<emphasis id="98FAEAC8FFB2FF98815C54078DD73766" box="[584,721,1535,1559]" italics="true" pageId="6" pageNumber="48">G. chandleri</emphasis>
|
||
</taxonomicName>
|
||
by fewer antennomeres and broader ventral valve of ovipositor.
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFB2FF9883D354258D4E37D7" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
|
||
<emphasis id="98FAEAC8FFB2FF9883D354258EC6374A" bold="true" box="[199,448,1570,1595]" pageId="6" pageNumber="48">Habitat distribution.</emphasis>
|
||
The
|
||
<typeStatus id="75358878FFB2FF9882E854248D2A374A" box="[508,556,1571,1595]" pageId="6" pageNumber="48">type</typeStatus>
|
||
specimen was found walking on snow at night in pine forest. I collected two juvenile specimens on the snow at night on
|
||
<date id="DE30101AFFB2FF98818754418C4F372F" box="[659,841,1606,1630]" pageId="6" pageNumber="48" value="2010-03-20">March 20, 2010</date>
|
||
and three more in the twilight zone of Blind Leads Cave under stones. The adult female specimen examined by
|
||
<bibRefCitation id="CE1F4B2BFFB2FF988052546D8B6837F3" author="Jarvis" box="[838,1134,1642,1666]" pageId="6" pageNumber="48" refString="Jarvis, K. J. & Whiting, M. F. (2006) Phylogeny and biogeography of ice crawlers (Insecta: Grylloblattodea) based on six molecular loci: Designating conservation status for Grylloblattodea species. Molecular Phylogenetics and Evolution, 41, 222 - 237." type="journal article" year="2006">
|
||
<collectingCountry id="D299764AFFB2FF988052546D8C8137F3" box="[838,903,1642,1666]" name="United States Pacific Island Wildlife Refuges" pageId="6" pageNumber="48">Jarvis</collectingCountry>
|
||
and Whiting (2006)
|
||
</bibRefCitation>
|
||
was collected in a cheesebaited trap near the entrance of a cave.
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFB2FF9883D354B58A773663" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
|
||
<emphasis id="98FAEAC8FFB2FF9883D354B58D0E37BA" bold="true" box="[199,520,1714,1739]" pageId="6" pageNumber="48">Geographical distribution.</emphasis>
|
||
<taxonomicName id="6D8E4D59FFB2FF98810754B58C3D37BB" box="[531,827,1714,1738]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFB2FF98810754B58C3D37BB" box="[531,827,1714,1738]" italics="true" pageId="6" pageNumber="48">Grylloblatta siskiyouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFB2FF98805154B48C8E37BA" box="[837,904,1715,1739]" pageId="6" pageNumber="48" rank="species">
|
||
<emphasis id="98FAEAC8FFB2FF98805154B48C8E37BA" bold="true" box="[837,904,1715,1739]" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is presently known only from Oregon Caves National Monument (
|
||
<figureCitation id="32B52A5FFFB2FF98829854D18ED7379F" box="[396,465,1750,1774]" captionStart="FIGURE 3" captionStartId="4.[151,250,1659,1682]" captionTargetBox="[177,1435,193,1608]" captionTargetId="figure@4.[151,1436,193,1638]" captionTargetPageId="4" captionText="FIGURE 3. Geographical distribution of grylloblattid populations (dots) and type localities (crosses and asterisks) in California and Oregon, USA." httpUri="https://zenodo.org/record/212978/files/figure.png" pageId="6" pageNumber="48">Fig. 3</figureCitation>
|
||
). Due to its surface activity, it is potentially more widespread in the Siskiyou Mountains.
|
||
<bibRefCitation id="CE1F4B2BFFB2FF9883CE54FC8E763663" author="Kamp" box="[218,368,1786,1810]" pageId="6" pageNumber="48" refString="Kamp, J. W. (1973) Biosystematics of the Grylloblattodea. Department of Zoology, University of British Columbia, Vancouver, 275 pp." type="book" year="1973">Kamp (1973)</bibRefCitation>
|
||
refers to a grylloblattid population near Mt. Ashland that might represent the same species.
|
||
</paragraph>
|
||
<paragraph id="AA3136DAFFB2FF9883D355198DC436B7" blockId="6.[151,1437,151,1991]" pageId="6" pageNumber="48">
|
||
<emphasis id="98FAEAC8FFB2FF9883D355198D0F3646" bold="true" box="[199,521,1822,1847]" pageId="6" pageNumber="48">Phylogenetic relationships.</emphasis>
|
||
Based on mitochondrial cytochrome oxidase subunit II data,
|
||
<taxonomicName id="6D8E4D59FFB2FF9887DE55188A7A3647" box="[1226,1404,1822,1846]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="siskiyouensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFB2FF9887DE55188A7A3647" box="[1226,1404,1822,1846]" italics="true" pageId="6" pageNumber="48">G. siskiyouensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFB2FF98869155198FBE362A" pageId="6" pageNumber="48" rank="species">
|
||
<emphasis id="98FAEAC8FFB2FF98869155198FBE362A" bold="true" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(GenBank Accession
|
||
<accessionNumber id="B5DDAB39FFB2FF9882A355458D2A362B" box="[439,556,1858,1882]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/JN612961" pageId="6" pageNumber="48">JN612961</accessionNumber>
|
||
) is distinct from (minimum ~10%) other nearby grylloblattids (
|
||
<tableCitation id="E70C0361FFB2FF9887E855458A57362B" box="[1276,1361,1858,1882]" captionStart="TABLE 1" captionStartId="5.[151,235,1016,1037]" captionTargetBox="[151,1436,1126,1580]" captionTargetPageId="5" captionText="TABLE 1. Genetic divergence of Grylloblatta siskiyouensis, G. oregonensis, and G. m a r m o re u s from nearby grylloblattid populations. Values are Kimura two-parameter corrected pairwise genetic distance at the cytochrome oxidase II gene. Intraspecific genetic variability, if available, is shown for each species." httpUri="http://table.plazi.org/id/FEF16652FFB1FF9B838351FF8C63353A" pageId="6" pageNumber="48" tableUuid="FEF16652FFB1FF9B838351FF8C63353A">Table 1</tableCitation>
|
||
). This species is represented by the Oregon Caves sample deposited in GenBank (
|
||
<bibRefCitation id="CE1F4B2BFFB2FF9880FC55618BE9360F" author="Jarvis" box="[1000,1263,1894,1918]" pageId="6" pageNumber="48" refString="Jarvis, K. J. & Whiting, M. F. (2006) Phylogeny and biogeography of ice crawlers (Insecta: Grylloblattodea) based on six molecular loci: Designating conservation status for Grylloblattodea species. Molecular Phylogenetics and Evolution, 41, 222 - 237." type="journal article" year="2006">
|
||
<collectingCountry id="D299764AFFB2FF9880FC55618B2D360F" box="[1000,1067,1894,1918]" name="United States Pacific Island Wildlife Refuges" pageId="6" pageNumber="48">Jarvis</collectingCountry>
|
||
& Whiting 2006
|
||
</bibRefCitation>
|
||
), from which it differs by a single base (0.1%). In a neighbor-joining analysis, it is grouped with
|
||
<taxonomicName id="6D8E4D59FFB2FF988731558C8BCD36D3" box="[1061,1227,1931,1954]" class="Insecta" family="Grylloblattidae" genus="Grylloblatta" kingdom="Animalia" order="Grylloblattodea" pageId="6" pageNumber="48" phylum="Arthropoda" rank="species" species="oregonensis" status="sp. nov.">
|
||
<emphasis id="98FAEAC8FFB2FF988731558C8BCD36D3" box="[1061,1227,1931,1954]" italics="true" pageId="6" pageNumber="48">G. oregonensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="83C957B3FFB2FF9887C5558C8A1636D2" box="[1233,1296,1931,1955]" pageId="6" pageNumber="48" rank="species">
|
||
<emphasis id="98FAEAC8FFB2FF9887C5558C8A1636D2" bold="true" box="[1233,1296,1931,1955]" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<emphasis id="98FAEAC8FFB2FF988653558C8FEF36B6" italics="true" pageId="6" pageNumber="48">G. m a rmoreus</emphasis>
|
||
<emphasis id="98FAEAC8FFB2FF9883FB55A88E2836B6" bold="true" box="[239,302,1967,1991]" pageId="6" pageNumber="48">n. sp.</emphasis>
|
||
in an unresolved polytomy (
|
||
<figureCitation id="32B52A5FFFB2FF98816455A98DB336B7" box="[624,693,1966,1990]" captionStart="FIGURE 4" captionStartId="7.[151,250,1651,1674]" captionTargetBox="[242,1290,187,1620]" captionTargetId="figure@7.[230,1359,167,1634]" captionTargetPageId="7" captionText="FIGURE 4. Relationships of new Grylloblatta species based on mitochondrial cytochrome oxidase subunit II gene data." httpUri="https://zenodo.org/record/212979/files/figure.png" pageId="6" pageNumber="48">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |