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<document id="A2486E78AC25F30CD987E920EE1B3FB3" ID-CLB-Dataset="301495" ID-DOI="10.3897/zookeys.1209.127951" ID-GBIF-Dataset="260efa91-98a5-4abe-96de-636bdebff8e8" ID-ZooBank="A07CCEE1-C2D5-4993-9437-55E3040E1F79" ID-publisher-id="127951" URI-arpha="64DE4B4F-BF17-5208-91E2-23C3DD2B612B" URI-zoobank="http://zoobank.org/A07CCEE1-C2D5-4993-9437-55E3040E1F79" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1723647139662" checkinUser="pensoft" docAuthor="Zhang, Ludan, Wang, Ying, Li, Shuqiang, Zhang, Xiaoqing &amp; Yao, Zhiyuan" docDate="2024" docId="1E9C3E64934F5342823C2185B1342211" docLanguage="en" docName="ZooKeys 1209: 315-330" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/127951/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana lingui Zhang, Li &amp; Yao 2024, sp. nov." docType="treatment" docUuid="1B082AB3-F3BF-4B1D-B26E-E6E90BA19883" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="330" masterDocId="64DE4B4FBF17520891E223C3DD2B612B" masterDocTitle="Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China" masterLastPageNumber="330" masterPageNumber="315" pageNumber="315" updateTime="1723650954597" updateUser="admin">
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<mods:title id="20C743A3DEED1D8FD5699FA2B7F9C28D">Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China</mods:title>
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<mods:namePart id="A7CB3B2A6EDC67B9D921DAC9DA636F7F">Zhang, Ludan</mods:namePart>
<mods:nameIdentifier id="7A6C3EA0124037F2CECA2B3DA27CC1E6" type="ORCID">0000-0001-5657-751X</mods:nameIdentifier>
<mods:affiliation id="06596E51C476149B63A208D07F05B0F8">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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<mods:namePart id="1E47E1617384549DAE860EB78A277530">Wang, Ying</mods:namePart>
<mods:affiliation id="6150713290EDA028F7403A474A130D47">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name id="8C59F1F8C0F02861AC51A5B72A775E8D" type="personal">
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<mods:namePart id="3A480B74646CC6CD20126BF7FAA214E7">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="F86818139F856DF315574009ADF8198F" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="BD97CA001A44BB379B61363897F93535">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="6EACD9DA5BFC4312C54FB999294EE35A">Zhang, Xiaoqing</mods:namePart>
<mods:affiliation id="608ABA3838E271614EB5A209627EBDD7">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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</mods:role>
<mods:namePart id="7D85D0B62A6197E717CE296BAE78246F">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="9D719E98FFA5B0313CDAB8517521DF28" type="ORCID">0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="DC2CD7373CDE7AAAEC10B08933E1D9CD">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:part id="83728EA7F028EB8618D2E7B73676A092">
<mods:date id="4E57B4687B99D20458B6C7002E6034A2">2024</mods:date>
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<mods:number id="9CB4D03460C546ED6B9F767A2A6F9FE0">2024-08-14</mods:number>
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<mods:detail id="DC86AF9A43700CD0BC792125AB10CFCA" type="volume">
<mods:number id="06DC5710FCB1DB4D5D5C703DE16FCDA1">1209</mods:number>
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<treatment id="1E9C3E64934F5342823C2185B1342211" ID-DOI="http://doi.org/10.5281/zenodo.13323010" ID-Zenodo-Dep="13323010" ID-arpha="1E9C3E64-934F-5342-823C-2185B1342211" ID-zoobank="https://zoobank.org/1B082AB3-F3BF-4B1D-B26E-E6E90BA19883" LSID="urn:lsid:zoobank.org:act:1B082AB3-F3BF-4B1D-B26E-E6E90BA19883" LSID-ZBK="urn:lsid:zoobank.org:act:1B082AB3-F3BF-4B1D-B26E-E6E90BA19883" httpUri="http://treatment.plazi.org/id/1E9C3E64934F5342823C2185B1342211">
<subSubSection id="8EB438C528D3FB49C402641344B9C297" type="nomenclature">
<paragraph id="4A53DE185DB45A228A4BC6037D374B64">
<taxonomicName id="8489EE5F1BB3DACFAE0FD37E643554F9" authority="Zhang, Li &amp; Yao" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lingui" status="sp. nov.">
<emphasis id="0AF8F4F125449C29BAD1C7E9F63C44E5" italics="true">Belisana lingui</emphasis>
Zhang, Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="6BF5CB10DF1AE8FFAD923DA137A8B3AE" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="01CC0E9F011B6846BF7961BC502FC165" type="description">
<paragraph id="18210AD63F2737A356C082B8E80B6B8C">
<figureCitation id="FE973114A6F25159A1956B4AEB706AED" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">Figs 4</figureCitation>
,
<figureCitation id="6444B95EFBF3053EB35D9B619164E404" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5</figureCitation>
,
<figureCitation id="CB0648073651CC19BAAE4C6CD2232EFE" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 C, D</figureCitation>
,
<figureCitation id="C8D3B4D9CC698EA3E6AE17BE3C70BB01" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG6AE" type="type material">
<paragraph id="2EC9D27E63ED03CDBE0E51F243F53705">
<heading id="1639C7D570DE2779B076F7BBF0CB28F2" reason="title">Type material.</heading>
</paragraph>
<paragraph id="5424512118AEF812BB6543413DAC8477">
<materialsCitation id="76F29BFDAC7DC2F0E9F04F97C392ECE7" collectingDate="2009-07-18" collectionCode="IZCAS" collectorName="Z Yao" country="China" county="Lingui County" elevation="161" latitude="25.21365" location="Yanmendi Village" longLatPrecision="1" longitude="110.200836" municipality="Guilin" specimenCode="Ar 44993" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="62EFE1707007AA96FC73F50D2372E103" bold="true">
<emphasis id="EF149D17ECA469FDD0C290818C2A6072" bold="true" italics="true">
<typeStatus id="0D50DCD94E3F1BCE4462ACB2F25BEA61" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="624D3F1980895B7B4C0575DFFE6FFD3E" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="1A7AFC5750974367A16F9DEE9AA47926" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="0991694410B7C4E2D16572630D8A23A6" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingMunicipality id="37FA2DAE42470BCC5EFCDC9D61B9C18F">Guilin</collectingMunicipality>
,
<collectingCounty id="9E6050CCDD8685FA7520494AE5E45DAE">Lingui County</collectingCounty>
,
<location id="3DA449F0964DA9A0521DFA845C35978A" LSID="urn:lsid:plazi:treatment:1E9C3E64934F5342823C2185B1342211:3DA449F0964DA9A0521DFA845C35978A" country="China" county="Lingui County" latitude="25.21365" longLatPrecision="1" longitude="110.200836" municipality="Guilin" name="Yanmendi Village" stateProvince="Guangxi">Yanmendi Village</location>
,
<location id="172B27C8CDBE49D12909ED4E5EC3EECB" LSID="urn:lsid:plazi:treatment:1E9C3E64934F5342823C2185B1342211:172B27C8CDBE49D12909ED4E5EC3EECB" country="China" county="Lingui County" latitude="25.21365" longLatPrecision="1" longitude="110.200836" municipality="Guilin" name="Shuixianyan Cave" stateProvince="Guangxi">Shuixianyan Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EU6AE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[110.200833,25.213650]}">
<geoCoordinate id="3BC81A9317B20011238097BA3542CF8B" degrees="25" direction="north" minutes="12.819" orientation="latitude" precision="1" value="25.21365">25 ° 12.819 ' N</geoCoordinate>
,
<geoCoordinate id="F49B6F1F907BABDF1319F960383483F1" degrees="110" direction="east" minutes="12.050" orientation="longitude" precision="1" value="110.200836">110 ° 12.050 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="3D6A189A213D95A803CD90D74EC10831" metricMagnitude="2" metricUnit="m" metricValue="1.61" unit="m" value="161.0">
<elevation id="850C4C6A33531750AE0172AE0F3B57F9" metricMagnitude="2" metricUnit="m" metricValue="1.61" unit="m" value="161.0">161 m</elevation>
</quantity>
;
<date id="16FE6ED85EB94FAFFDD286047580F10E" value="2009-07-18">
<collectingDate id="FBDE800F0577474E524D5D4EC0F44494" value="2009-07-18">18 Jul. 2009</collectingDate>
</date>
;
<collectorName id="7B676DDB97697682D63DA721BE0BE5A4">Z Yao</collectorName>
leg.;
<named-content id="5B6721ADCB18F8B883580E7F9F1B5845" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="D12379888CFE573B1B8CB9E875745FE4" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="31502CBF76866AA686EFA59709058085">Ar 44993</specimenCode>
</materialsCitation>
.
<materialsCitation id="075F46A12B9F2E212556CC43BCF497D8" collectingDate="2009-07-18" collectionCode="IZCAS" collectorName="Z Yao &amp; Ar" country="China" county="Lingui County" elevation="161" latitude="25.21365" location="China" longLatPrecision="1" longitude="110.200836" municipality="Guilin" specimenCode="Ar 44994" specimenCount="3" specimenCount-female="3" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="FFDCDCCB62B8390A2DA75A3BBD47ED9F" bold="true">
<emphasis id="9025FE3CC01175754738705C6101B933" bold="true" italics="true">
<typeStatus id="F38EE252B20999A6748917C8650EEA2B" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="663A7ABB559FBDCAB934578A96497188" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="7A5D8D6FF77126033C6A36F390BD2BED" count="3" type="female">3 ♀</specimenCount>
; same data as for holotype;
<named-content id="D5B1FBBCDAEE8469182A7019EAC8F7F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="8B6088ACFD03EE257B620973A00FB1B8" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="CF017574C06AFC272307E6E972502B48">Ar 44994</specimenCode>
–96
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGAAG" type="etymology">
<paragraph id="D76C918ADAF8A1D9B10F2CAEBE6132FF">
<heading id="C1C3D0D86422AD45C6B582E6E5E28E4A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="977DD56CFDD67C7D31F74E76276B46F4">The specific name refers to the type locality; noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0ELAAG" type="diagnosis">
<paragraph id="54813F3A1F0ECE23ACEABF026934E892">
<heading id="A5C5978E305FC51747F14F1F8D32EE37" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="9EB21DEF967EB3D4D97BD42BD52B48D4">
The new species resembles
<taxonomicName id="5FB3C9AAB2F8B543329930B98E72B511" authority="Zhang &amp; Peng, 2011" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="63AE43FC18093066F370F13330326E6D" italics="true">B. galeiformis</emphasis>
<bibRefCitation id="3EFFFF9522DA25460098A65570248FF2" DOI="10.11646/zootaxa.2989.1.2" author="Zhang F &amp; Peng Y" issue="1" journalOrPublisher="Zootaxa" pagination="51-68" refId="B15" refString="Zhang F, Peng Y (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa 2989 (1): 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1. 2" title="Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China." volume="2989" year="2011">Zhang &amp; Peng, 2011</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="98AFC355D6E5A687B63B10F454A34B64" DOI="10.11646/zootaxa.2989.1.2" author="Zhang F &amp; Peng Y" firstAuthor="Zhang" issue="1" journalOrPublisher="Zootaxa" page="52" pagination="51-68" refId="B15" refString="Zhang F, Peng Y (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa 2989 (1): 51–68. https://doi.org/10.11646/zootaxa.2989.1.2" title="Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China." volume="2989" year="2011">Zhang and Peng 2011: 52</bibRefCitation>
, fig. 1 A – F) by having similar bulbal apophyses and male chelicerae (Fig.
<figureCitation id="DCCFF6505D18102B49F2509DDC1973B6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 C, D</figureCitation>
), but can be distinguished by procursus without prolatero-subdistal sclerite (Figs
<figureCitation id="4F638D62DACE2AEFDE398D86A15527A8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 C</figureCitation>
,
<figureCitation id="80C34FBA31E6948158570D26459B55DB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 C</figureCitation>
vs. present in
<taxonomicName id="08327A1E4313579078F55C16F248CB23" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="39CECBFFC2870056C04577350551B468" italics="true">B. galeiformis</emphasis>
</taxonomicName>
), by epigyne with posterior pockets (
<abbrev id="ABBRID0EXBAG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="FCF7B28A3CC64DDFCBAA7E81E2DDBD84" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 A</figureCitation>
,
<figureCitation id="77F4D3C955719CDDE17ED3B7D926397A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 C</figureCitation>
vs. median in
<taxonomicName id="9FA57507359C9BFE39B04577828C2CD2" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="A8A77D8674540A31C4D578948719BD08" italics="true">B. galeiformis</emphasis>
</taxonomicName>
), by pore plates anteriorly narrow and posteriorly wide (
<abbrev id="ABBRID0EOCAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="F054C0DEB4AECBAC86C13AA3C81F1309" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 B</figureCitation>
,
<figureCitation id="580B43170A66214DF8050CA9E9BEA736" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 D</figureCitation>
vs. elliptic in
<taxonomicName id="2AF0AFB8CFE42476394EEABF67C31C69" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="A41A089E58DABA7CD4FE738500111A0E" italics="true">B. galeiformis</emphasis>
</taxonomicName>
), and by dorsal shield of prosoma without marks (Fig.
<figureCitation id="44E1233811D1CBEF9BAA04815C42F445" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 E, G</figureCitation>
vs. with radiating marks in
<taxonomicName id="52CDF44EC1E9E6F66E5C7634A2903DD1" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="galeiformis">
<emphasis id="C6FD1C1AC07811ABDF7068B5B93D5119" italics="true">B. galeiformis</emphasis>
</taxonomicName>
); also distinguished from
<taxonomicName id="06709E28F5EC2CDF374430E59B34ECDF" authority="Zhang, Chen &amp; Zhu, 2008" authorityName="Zhang, Chen &amp; Zhu" authorityYear="2008" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongle">
<emphasis id="0B14BE56728F388FF5578FBED712A3A9" italics="true">B. tongle</emphasis>
<bibRefCitation id="0AD962F6AA3D6B5AF866F44AA6D23166" author="Zhang Y &amp; Chen H &amp; Zhu M" journalOrPublisher="Acta Zootaxonomica Sinica" pagination="654-656" refId="B16" refString="Zhang Y, Chen H, Zhu M (2008) A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae). Acta Zootaxonomica Sinica 33: 654 - 656." title="A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae)." volume="33" year="2008">Zhang, Chen &amp; Zhu, 2008</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F062E1BD2E75A74150864FC4311886F9" author="Zhang Y &amp; Chen H &amp; Zhu M" etAl="et al." firstAuthor="Zhang" journalOrPublisher="Acta Zootaxonomica Sinica" page="654" pagination="654-656" refId="B16" refString="Zhang Y, Chen H, Zhu M (2008) A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae). Acta Zootaxonomica Sinica 33: 654–656." title="A new troglophilous Belisana spider from Guangxi, China (Araneae, Pholcidae)." volume="33" year="2008">Zhang et al. 2008: 654</bibRefCitation>
, figs 1–5) by procursus without prolatero-subdistal sclerite and ventro-subdistal apophysis (Figs
<figureCitation id="46F3F9FA88ED17778F68C9FA8E00B768" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 C</figureCitation>
,
<figureCitation id="82112C5D2CFB50308F4BB720842E4F28" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 C</figureCitation>
vs. present in
<taxonomicName id="206B7B3F9B208D737D35235614EB27C8" authorityName="Zhang, Chen &amp; Zhu" authorityYear="2008" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tongle">
<emphasis id="5170BFA9F5C23D43A2DC086F3538038D" italics="true">B. tongle</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="7BEFD44D395CE46357D75AF6B5C4F801" ID-DOI="10.3897/zookeys.1209.127951.figure4" ID-arpha="93E5845D-2D6E-57F2-A3AE-380DD10F8256" httpUri="https://binary.pensoft.net/fig/1113528" startId="F4">
<paragraph id="3258EDF72A6E371EF355FEDB221A3C43">
<label id="5FBE33E73E6FCFF9D2997F3E1739CC5A">Figure 4.</label>
</paragraph>
<paragraph id="06C5E912B1C1237ED5E6611B33687DF4">
<taxonomicName id="ADE8DEC8D66B27B3D74458C38AC38ABB" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lingui" status="sp. nov.">
<emphasis id="25A0940166FD3846B9C6370859BAEB19" italics="true">Belisana lingui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CFCB28FE8E45568C711E1C87B56C60F6" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="6F99C56FFBA865F3E35DB0DB9E69B107" bold="true">A, B</emphasis>
palp (
<emphasis id="5D35B72993EED3CA33E08425CA5B8CAB" bold="true">A</emphasis>
prolateral view
<emphasis id="E57A1812892B0C3F23C3D8188695911B" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="0D67FC789DFE6124D96D37CF7B0F7CCC" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="8278810D8ECA0C4C7E142021FFD827B8" bold="true">C</emphasis>
prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis
<emphasis id="27B8908627A3085A22F8400152C78FE6" bold="true">D</emphasis>
retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap,
<abbrev id="ABBRID0EVFAG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 mm (
<emphasis id="169066409CD0C09EBC326E7B598A64D3" bold="true">A, B</emphasis>
); 0.02 mm (
<emphasis id="575750476C556C344E6FAC1F62E1D1B7" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="C02750C480375E3B16C0500E4592D05A" ID-DOI="10.3897/zookeys.1209.127951.figure5" ID-arpha="D36D7AEF-3B84-5F4D-880D-264D2E592D92" httpUri="https://binary.pensoft.net/fig/1113529" startId="F5">
<paragraph id="2F9AD1217CE9BF153967640B62DB4B67">
<label id="D759C06453A41D39CB484B44D3A96A96">Figure 5.</label>
</paragraph>
<paragraph id="3192E70C033A383AAAA5F984168C5276">
<taxonomicName id="029862123C278BC71025A220FE09ECD3" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lingui" status="sp. nov.">
<emphasis id="AC393F41E44490432DB4A792BE31C376" italics="true">Belisana lingui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="20EA401F0264B30D2C33D4FE414A3D45" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="C0800B42636CB6D6ACD36E57AA98AF27" bold="true">C – F</emphasis>
) and paratype female (
<emphasis id="7564779D0CFF299D3C0B983A8A93CD93" bold="true">A, B, G, H</emphasis>
)
<emphasis id="84DAE8C7EE8E087A461A82CDDD22E93B" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="7F1E9156CF08D9308CB7A1E75D358D02" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="B2250DA1F3D5F0930D776EFD7A9F6CFA" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="7B4E1EA2E638EC538FE7EE50152E5588" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="FECE47F5EB1AF3757494F001359C40EC" bold="true">E – H</emphasis>
habitus (
<emphasis id="344C37B1C1C6D77B3309B2010DFCB7DD" bold="true">E, G</emphasis>
dorsal view
<emphasis id="6117317A0224EB18E4EAF9E1BBBB03A4" bold="true">F</emphasis>
lateral view
<emphasis id="4BBA73847B65DEA442FF7A104DB39F7D" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EJHAG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0ENHAG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0ERHAG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0EVHAG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0EZHAG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0E4HAG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.05 mm (
<emphasis id="9F817F77DA507AB9FEF8A41A3FCBCEFC" bold="true">A – D</emphasis>
); 0.30 mm (
<emphasis id="51A74C230341E72DC1F7C4788AEBE519" bold="true">E – H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EKIAG" type="description">
<paragraph id="1C02C873C792DE3E448F3291D2ED1FAF">
<heading id="4E1D3186576578CFB43CF711B22D7731" reason="title">Description.</heading>
</paragraph>
<paragraph id="52AA0136B8C5B335DCF94747895B9BC0">
<emphasis id="29473F865A199040244AA533B7A5E9FC" bold="true">Male</emphasis>
(
<emphasis id="7B9FB469A0A950A4FCA221119AE62E6F" bold="true" italics="true">
<typeStatus id="E60467D77A0D371EDB15AF661C9D8EF0" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.51 (1.59 with clypeus), prosoma 0.56 long, 0.59 wide, opisthosoma 0.95 long, 0.84 wide. Leg I: 20.66 (5.26, 0.33, 5.19, 7.98, 1.90), leg II: 14.16 (3.88, 0.33, 3.64, 5.00, 1.31), leg III: – (2.53, 0.30, 2.25, 3.00, –), leg IV: 11.97 (3.60, 0.30, 3.23, 3.76, 1.08); tibia I
<abbrev id="ABBRID0EVIAG" xlink_title="length/diameter">L / d</abbrev>
: 80. Eye interdistances and diameters:
<abbrev id="ABBRID0EZIAG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E4IAG" xlink_title="posterior median eye">PME</abbrev>
0.12,
<abbrev id="ABBRID0EBJAG" xlink_title="posterior median eye">PME</abbrev>
0.05,
<abbrev id="ABBRID0EFJAG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EJJAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0ENJAG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.49 / 0.44. Habitus as in Fig.
<figureCitation id="18BA2151495E19E493C2124769E43F35" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 E, F</figureCitation>
. Dorsal shield of prosoma and sternum yellowish, without marks. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Eyes without pigments, but apparently with small lenses. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0EVJAG" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="E0F4389D0A1743C81439003B47A45DA0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 D</figureCitation>
) and pair of distal apophyses (
<abbrev id="ABBRID0E4JAG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="811FA9E2C4EDC374CE9D68643F9DEE04" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 D</figureCitation>
; distance between tips: 0.22). Palp as in Fig.
<figureCitation id="2364D57088A4FB048BF812FFB64E8021" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 A, B</figureCitation>
; trochanter with ventral apophysis (arrow
<quantity id="0C78AB576C17DA000064458139291E87" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Fig.
<figureCitation id="C9CA0F1F88AD98B27BDAABC4604087AE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 B</figureCitation>
); femur with small retrolatero-proximal protrusion (arrow
<quantity id="E3DE1BCAA27A7B4014C58980B91FE0C1" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Fig.
<figureCitation id="10BC10E9E6C45C7D19BCADF7D1E9FB7F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 B</figureCitation>
); procursus simple proximally but complex distally, with ventro-subdistal membranous lamella (arrow
<quantity id="DB1F45283B08D57E7EE81528C5C9BBEB" metricMagnitude="-2" metricUnit="m" metricValue="2.54" unit="in" value="1.0">1 in</quantity>
Figs
<figureCitation id="B1E11F86FB538E32933A6502CE686E98" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 C</figureCitation>
,
<figureCitation id="046F13763B69F7C30BAFF8EE736CEC46" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 C</figureCitation>
), spine-shaped distal apophysis (tip broken; arrow
<quantity id="BF3C20FA71DA3A5C9CC121862CB4CDEC" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
Figs
<figureCitation id="3476B5BD33909ADED1A1840C78A2DF66" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 C</figureCitation>
,
<figureCitation id="53A196AD18E8229EC7CDDF7787C41185" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 C</figureCitation>
), retrolatero-subdistal membranous lamella (arrow in Figs
<figureCitation id="C68E333B44BB98458A37F65EEEAA51F7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 D</figureCitation>
,
<figureCitation id="689720915C5CA19105618BECE0B0C34E" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 D</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="C9294D95865CF89C01257377F119E3D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Belisana lingui sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at distal apophysis D retrolateral view, arrow points at retrolatero-subdistal membranous lamella). Abbreviations: f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure4" httpUri="https://binary.pensoft.net/fig/1113528">4 D</figureCitation>
,
<figureCitation id="F64ED77A17C8AB8B3C114F3AEC087EBB" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 D</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0ERLAG" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="63D03CDCBF363966080B37B4F26F017F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="0C832CFB7B692C786939C0554A530434" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 C</figureCitation>
). Retrolateral trichobothria on tibia I at 4 % proximally; legs with short vertical setae on metatarsi; tarsus I with 17 distinct pseudosegments.
</paragraph>
<paragraph id="70D12DC284B8100F50137850466865B6">
<emphasis id="DB31CAFB92CEBC1AE7857DB85F369B63" bold="true">Female</emphasis>
(
<emphasis id="DC84D8FC5488F6B2CFAA4FE132847FD6" bold="true" italics="true">
<typeStatus id="9AC46A4EE543A526AB1ACCEB49FC5565" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="18EF5A3CED3D2AEEE349A8B46E966739" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="BBD8F8BEC01BA5D03402B9ABE2C68651" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="97D186B9F39A18AD69C8B5885A1C9347">Ar 44994</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="ECAF6EE880239424683A4505FA340CC8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 G, H</figureCitation>
. Total length 2.00 (2.09 with clypeus), prosoma 0.58 long, 0.63 wide, opisthosoma 1.42 long, 1.01 wide; tibia I: 2.34; tibia I
<abbrev id="ABBRID0EMMAG" xlink_title="length/diameter">L / d</abbrev>
: 39. Eye interdistances and diameters:
<abbrev id="ABBRID0EQMAG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EUMAG" xlink_title="posterior median eye">PME</abbrev>
0.11,
<abbrev id="ABBRID0EYMAG" xlink_title="posterior median eye">PME</abbrev>
0.04,
<abbrev id="ABBRID0E3MAG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EANAG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EENAG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.50 / 0.44. Epigyne simple and flat, with pair of posterior pockets 0.20 apart (
<abbrev id="ABBRID0EINAG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="BF6EA90F431F26357283A8D249C0B9D1" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 A</figureCitation>
,
<figureCitation id="385C9B8FEA56E8463997C5FAADD662B0" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 C</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0EUNAG" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="C7C1A4DC3650524C8F99E1761B530484" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 B</figureCitation>
,
<figureCitation id="EC4B0718BFE5158D2CCC67E476CC06E6" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 D</figureCitation>
) and pair of anteriorly narrow and posteriorly wide pore plates (
<abbrev id="ABBRID0EAOAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="F41513BB6A8D2E937511DA9567693F67" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Belisana lingui sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 mm (A – D); 0.30 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure5" httpUri="https://binary.pensoft.net/fig/1113529">5 B</figureCitation>
,
<figureCitation id="099B10A35B4B353653A3F916EB4615CF" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EMOAG" type="variation">
<paragraph id="94DAB6C4F2CA4E7EC67514A9320B6638">
<heading id="486DD75FB35263D933F9C296DB05033C" reason="title">Variation.</heading>
</paragraph>
<paragraph id="C266E151CC5DBAC674BECCEA27353775">
Tibia I in the other
<specimenCount id="35DE9DBB6C9814365A76F6410E3D85EA" count="2" type="female">two female</specimenCount>
<typeStatus id="20A5BD2395AA1339837FC16E8478CF26" type="paratype">paratypes</typeStatus>
(
<named-content id="04FA8BA4AEDDA6094F71716A1020B437" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="4C26BBB57BD16AFD13B6310A7F3563FE" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="51B8D5B77CF7319774D55F596D811407">Ar 44995</specimenCode>
–96): 2.34, 2.41.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWOAG" type="habitat">
<paragraph id="BECE931D07FC76F2B5BDFF0FFB0C5449">
<heading id="45B87C09795FC6124531043CDFF34537" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="40D81F5E631C6F9B32DA51AFE54A8E6F">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0E2OAG" type="distribution">
<paragraph id="92D95284C9F16B4846D99CDCD8272C16">
<heading id="2EC18CB32F09FAFAA570E33E2863DDA5" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="2C389FD2C1B31FA5436ECAC4E380B63B">
<collectingCountry id="65398D4789BAFA94EEF65C9158BC6F97" name="China">China</collectingCountry>
(
<collectingRegion id="FE499541F1B8AE9634139394C417F5D0" country="China" name="Guangxi">Guangxi</collectingRegion>
, type locality; Fig.
<figureCitation id="32EA4BE753B05520CDBD4A19BB474807" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guangxi, China 1 Belisana colubrina Zhang &amp; Peng, 2011 2 B. guilin Yao &amp; Li, 2020 3 B. langping sp. nov. 4 B. lingui sp. nov. 5 B. naling Yao &amp; Li, 2020 6 B. parallelica Zhang &amp; Peng, 2011 7 B. tianlinensis Zhang &amp; Peng, 2011 8 B. tianyang sp. nov. 9 B. tongle Zhang, Chen &amp; Zhu, 2008 10 B. xuanguan Yao &amp; Li, 2020 11 B. zhangi Tong &amp; Li, 2007." figureDoi="10.3897/zookeys.1209.127951.figure1" httpUri="https://binary.pensoft.net/fig/1113525">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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