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<mods:title id="A7ACA88110041CACB175B63743034BF3">A revision of the Geothelphusa leIJicerIJix species group (Crustacea: Decapoda: Brachyura: Potamidae), with descriptions of three new species</mods:title>
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<mods:namePart id="26D4619D1E60703C9F8DA124D53214E7">Shokita, Shigemitsu</mods:namePart>
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<mods:namePart id="F2FF2A1E6E9F331FC289753EF0508892">Ng, Peter K. L.</mods:namePart>
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<mods:title id="34EE583CF9A15358F18BD5CA84E41B56">Journal of Natural History</mods:title>
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<mods:date id="CF80507A8429761A7FB5FFA295FA3890">2006</mods:date>
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<mods:number id="9F287927093B74BACCC8359C2E95AB2B">40</mods:number>
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<mods:number id="4324D086926CDD618320FFDBD4F4E734">13 - 14</mods:number>
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<heading id="C5147B231D65FFB2221FF9C971F1271C" box="[325,956,1553,1577]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="776" reason="2">
Key to species of
<emphasis id="AC97105D1D65FFB2214FF9C97158271C" box="[533,789,1553,1577]" italics="true" pageId="19" pageNumber="776">
<taxonomicName id="59E3B7CC1D65FFB2214FF9C970EB271C" ID-CoL="7NVV5" authorityName="Stimpson" authorityYear="1858" box="[533,678,1553,1577]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="776" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
leυicerυix
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species-group
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<keyLead id="25126F7A1D65FFB22327F98A76D9275F" box="[125,1172,1618,1642]" pageId="19" pageNumber="776">1. Anterolateral margin faintly cristate or not cristate......... 2</keyLead>
</paragraph>
<paragraph id="9E5CCC4F1D65FFB22327F9AB76D927BE" blockId="19.[125,1172,1618,1675]" box="[125,1172,1651,1675]" pageId="19" pageNumber="776">
<keyLead id="25126F7A1D65FFB22327F9AB76D927BE" box="[125,1172,1651,1675]" pageId="19" pageNumber="776"> Anterolateral margin distinctly cristate............. 4</keyLead>
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<caption id="CA9C9CC71D62FFB523D0FB0A72AE242F" ID-DOI="http://doi.org/10.5281/zenodo.4581783" ID-Zenodo-Dep="4581783" httpUri="https://zenodo.org/record/4581783/files/figure.png" pageId="20" pageNumber="777" startId="20.[138,198,1234,1253]" targetBox="[138,1187,133,1198]" targetPageId="20">
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Figure 9. Frontal view of
<taxonomicName id="59E3B7CC1D62FFB52220FB0A700525D0" ID-CoL="3FT3V" authority="Naruse &amp; Shokita &amp; Ng, 2006" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" box="[378,584,1234,1253]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="species" species="kumejima" status="sp. nov.">
<emphasis id="AC97105D1D62FFB52220FB0A700525D0" box="[378,584,1234,1253]" italics="true" pageId="20" pageNumber="777">Geothelphusa kumejima</emphasis>
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<taxonomicNameLabel id="B7A4AD261D62FFB52114FB0A70F725D0" box="[590,698,1234,1253]" pageId="20" pageNumber="777" rank="species">new species</taxonomicNameLabel>
and
<taxonomicName id="59E3B7CC1D62FFB521B2FB0A71C225D0" ID-CoL="3FT3R" authority="Naruse &amp; Shokita &amp; Ng, 2006" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" box="[744,911,1234,1253]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="species" species="iheya" status="sp. nov.">
<emphasis id="AC97105D1D62FFB521B2FB0A71C225D0" box="[744,911,1234,1253]" italics="true" pageId="20" pageNumber="777">Geothelphusa iheya</emphasis>
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. (a)
<taxonomicName id="59E3B7CC1D62FFB52777FB0A729225CA" ID-CoL="3FT3V" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="species" species="kumejima">
<emphasis id="AC97105D1D62FFB52777FB0A729225CA" italics="true" pageId="20" pageNumber="777">Geothelphusa kumejima</emphasis>
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, paratype, ZRC 2006.0077, male, CL 35.0 mm; (b)
<taxonomicName id="59E3B7CC1D62FFB521BAFB34716225CA" ID-CoL="3FT3R" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" box="[736,815,1260,1279]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="species" species="iheya">
<emphasis id="AC97105D1D62FFB521BAFB34716225CA" box="[736,815,1260,1279]" italics="true" pageId="20" pageNumber="777">G. iheya</emphasis>
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, paratype, ZRC 2006.0078, male, CL 28.7 mm.
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2. Male G1 very slender, total length to width ratio of G1 5.456.69, distal margin of subterminal segment narrower than middle part.........................
<emphasis id="AC97105D1D62FFB5218EFA7F719E248A" box="[724,979,1447,1471]" italics="true" pageId="20" pageNumber="777">
<taxonomicName id="59E3B7CC1D62FFB5218EFA7F7128248A" ID-CoL="7NVV5" authorityName="Stimpson" authorityYear="1858" box="[724,869,1447,1471]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
leυicerυix
</emphasis>
(
<bibRefCitation id="FA72B1BE1D62FFB520BEFA7F76D5248A" author="Rathbun MJ" box="[996,1176,1447,1471]" pageId="20" pageNumber="777" pagination="27 - 30" refId="ref9237" refString="Rathbun MJ. 1898. Description of three new species of fresh-water crabs of the genus Potamon. Proceedings of the Biological Society of Washington 12: 27 - 30. Plates 1, 2." type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
)
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<paragraph id="9E5CCC4F1D62FFB523D0FA1076EC2734" blockId="20.[138,1185,1382,1537]" pageId="20" pageNumber="777">
<keyLead id="25126F7A1D62FFB523D0FA1076EC2734" pageId="20" pageNumber="777"> Male G1 not very slender, distal margin of subterminal segment subequal with middle part..................... 3</keyLead>
</paragraph>
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<paragraph id="9E5CCC4F1D62FFB523D0F9C276EC2767" blockId="20.[138,1185,1562,1683]" pageId="20" pageNumber="777">
<keyLead id="25126F7A1D62FFB523D0F9C276EC2767" pageId="20" pageNumber="777">
3. Outer proximal angle of the subterminal segment of G1 with prominent knob...............
<taxonomicName id="59E3B7CC1D62FFB52008F9E2761A2767" ID-CoL="3FT3V" authority="Naruse &amp; Shokita &amp; Ng, 2006" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" box="[850,1111,1594,1618]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="species" species="kumejima" status="sp. nov.">
<emphasis id="AC97105D1D62FFB52008F9E2761A2767" box="[850,1111,1594,1618]" italics="true" pageId="20" pageNumber="777">Geothelphusa kumejima</emphasis>
</taxonomicName>
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<paragraph id="9E5CCC4F1D62FFB523D0F98376EC27A6" blockId="20.[138,1185,1562,1683]" pageId="20" pageNumber="777">
<keyLead id="25126F7A1D62FFB523D0F98376EC27A6" pageId="20" pageNumber="777">
Outer proximal angle of the subterminal segment of G1 with small knob..............
<emphasis id="AC97105D1D62FFB52060F9A3761A27A6" box="[826,1111,1659,1683]" italics="true" pageId="20" pageNumber="777">
<taxonomicName id="59E3B7CC1D62FFB52060F9A3718627A6" ID-CoL="7NVV5" authorityName="Stimpson" authorityYear="1858" box="[826,971,1659,1683]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="777" phylum="Arthropoda" rank="genus">Geothelphusa</taxonomicName>
grandioυata
</emphasis>
n. sp.
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<paragraph id="9E5CCC4F1D63FFB42327FF5176D92194" blockId="21.[125,1172,137,226]" box="[125,1172,137,161]" pageId="21" pageNumber="778">
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4. G1 stout, terminal segment strongly directed outwards.
<taxonomicName id="59E3B7CC1D63FFB4202CFF5176072194" ID-CoL="3FT3R" authority="Naruse &amp; Shokita &amp; Ng, 2006" authorityName="Naruse &amp; Shokita &amp; Ng" authorityYear="2006" box="[886,1098,137,161]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="778" phylum="Arthropoda" rank="species" species="iheya" status="sp. nov.">
<emphasis id="AC97105D1D63FFB4202CFF5176072194" box="[886,1098,137,161]" italics="true" pageId="21" pageNumber="778">Geothelphusa iheya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B7A4AD261D63FFB42709FF5176D92194" box="[1107,1172,137,161]" pageId="21" pageNumber="778" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
<paragraph id="9E5CCC4F1D63FFB42327FF7176D921D7" blockId="21.[125,1172,137,226]" pageId="21" pageNumber="778">
<keyLead id="25126F7A1D63FFB42327FF7176D921D7" pageId="21" pageNumber="778">
G1 slender, terminal segment gently directed outwards................
<taxonomicName id="59E3B7CC1D63FFB422ACFF1276D921D7" ID-CoL="6KCVN" authority="Shokita, Naruse, and Fujii, 2002" authorityName="Shokita, Naruse, and Fujii" authorityYear="2002" box="[502,1172,202,226]" class="Malacostraca" family="Potamidae" genus="Geothelphusa" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="778" phylum="Arthropoda" rank="species" species="miyakoensis">
<emphasis id="AC97105D1D63FFB422ACFF12715D21D7" box="[502,784,202,226]" italics="true" pageId="21" pageNumber="778">Geothelphusa miyakoensis</emphasis>
<bibRefCitation id="FA72B1BE1D63FFB42042FF1276D921D7" author="Shokita S &amp; Naruse T &amp; Fujii H" box="[792,1172,202,226]" pageId="21" pageNumber="778" pagination="443 - 448" refId="ref9704" refString="Shokita S, Naruse T, Fujii H. 2002. Geothelphusa miyakoensis, a new species of freshwater crab (Crustacea: Decapoda: Brachyura: Potamidae) from Miyako Island, Southern Ryukyus, Japan. Raffles Bulletin of Zoology 50: 443 - 448." type="journal article" year="2002">Shokita, Naruse, and Fujii, 2002</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
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