treatments-xml/data/52/3B/87/523B87B7FF95270606D6224BFF6AD13E.xml

<document id="AEEB3FB8BE3169343A9DC7DDEBAFC04F" ID-CLB-Dataset="45505" ID-DOI="10.11646/zootaxa.3815.1.6" ID-GBIF-Dataset="963ff3a4-4ab4-495b-95eb-a744279bc6a4" ID-ISSN="1175-5326" ID-Zenodo-Dep="226599" ID-ZooBank="51A13505-C327-49F7-8FCC-0C407927C76C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460436036422" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2014" docId="523B87B7FF95270606D6224BFF6AD13E" docLanguage="en" docName="zt03815p102.pdf" docOrigin="Zootaxa 3815 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cebudonus Ng, 2014, n. gen." docType="treatment" docVersion="9" lastPageNumber="96" masterDocId="AE02FFCFFF9427040641232CFFAFD712" masterDocTitle="Cebudonus poppeorum, a new genus and new species of eumedonine crab (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Philippines" masterLastPageNumber="102" masterPageNumber="94" pageNumber="95" updateTime="1698332252527" updateUser="ExternalLinkService">
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<mods:title id="C82AF596EAE77EC2C8EA49FC6AC114B9">Cebudonus poppeorum, a new genus and new species of eumedonine crab (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Philippines</mods:title>
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<mods:namePart id="0EFC6E6D29F3B7366153791C9E63455A">Ng, Peter K. L.</mods:namePart>
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<treatment id="523B87B7FF95270606D6224BFF6AD13E" ID-DOI="http://doi.org/10.5281/zenodo.6131210" ID-GBIF-Taxon="119431858" ID-Zenodo-Dep="6131210" LSID="urn:lsid:plazi:treatment:523B87B7FF95270606D6224BFF6AD13E" httpUri="http://treatment.plazi.org/id/523B87B7FF95270606D6224BFF6AD13E" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95">
<subSubSection id="9288652AFF95270506D6224BFED6D690" box="[151,377,359,386]" pageId="1" pageNumber="95" type="nomenclature">
<paragraph id="DA2D36A1FF95270506D6224BFED6D690" blockId="1.[151,377,359,386]" box="[151,377,359,386]" pageId="1" pageNumber="95">
<heading id="816581CDFF95270506D6224BFED6D690" bold="true" box="[151,377,359,386]" fontSize="11" level="1" pageId="1" pageNumber="95" reason="1">
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<taxonomicName id="1D924D22FF95270506D6224BFE8FD693" ID-CoL="7NWMK" box="[151,288,359,385]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270506D6224BFE8FD693" bold="true" box="[151,288,359,385]" italics="true" pageId="1" pageNumber="95">Cebudonus</emphasis>
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<taxonomicNameLabel id="F3D557C8FF95270507672244FED6D690" box="[294,377,360,386]" pageId="1" pageNumber="95" rank="genus">n. gen.</taxonomicNameLabel>
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<typeStatus id="05298803FF95270506D62282FF7FD6D5" box="[151,208,430,455]" pageId="1" pageNumber="95">Type</typeStatus>
species
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. 
<taxonomicName id="1D924D22FF95270507782282FD92D6D4" box="[313,573,430,455]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="poppeorum" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270507782282FD92D6D4" box="[313,573,430,455]" italics="true" pageId="1" pageNumber="95">Cebudonus poppeorum</emphasis>
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, n. sp.
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, by present designation.
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<subSubSection id="9288652AFF952705068622FEFF4AD315" pageId="1" pageNumber="95" type="diagnosis">
<paragraph id="DA2D36A1FF952705068622FEFF4AD315" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="95">
<emphasis id="E8E6EAB3FF952705068622FEFE95D6F9" bold="true" box="[199,314,466,491]" pageId="1" pageNumber="95">Diagnosis</emphasis>
. Carapace pentagonal; dorsal surface gently convex, covered with numerous small pits (
<figureCitation id="42A92A24FF952705036022FEFAC0D6F9" box="[1313,1391,466,491]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF952705033C22FFFA22D6F9" box="[1405,1421,467,491]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
); front with 2 long pseudorostral spines, subparallel, ca. 0.4 times total carapace length, separated by broad U-shaped cleft (
<figureCitation id="42A92A24FF952705069D2136FE81D521" box="[220,302,538,563]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270507012137FEFFD521" box="[320,336,539,563]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF95270507212137FEC1D521" box="[352,366,539,563]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">3</figureCitation>
C); anterolateral margin entire, with large, broadly triangular, sharp lateral tooth, projecting laterally, dorsal surface gently convex (
<figureCitation id="42A92A24FF952705041C2112FD02D545" box="[605,685,574,599]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270504FF2113FD61D545" box="[702,718,575,599]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF952705049C2113FD45D545" box="[733,746,575,599]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">3</figureCitation>
A, 4A); antennae with subrectangular basal article, twice as long as wide (
<figureCitation id="42A92A24FF9527050779214EFED3D569" box="[312,380,610,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Fig. 3</figureCitation>
B); antennules folding at ca. 45° (
<figureCitation id="42A92A24FF9527050541214EFCEBD569" box="[768,836,610,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Fig. 3</figureCitation>
B); epistome relatively broad with depressed median part (
<figureCitation id="42A92A24FF952705069021AAFEBDD58D" box="[209,274,646,671]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Fig. 3</figureCitation>
A–C); third maxilliped with ischium subrectangular, with deep submedian longitudinal sulcus (
<figureCitation id="42A92A24FF952705037721AAFA2CD58D" box="[1334,1411,646,671]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
C, 4A), merus subquadrate, anteroexternal angle subauriculiform (
<figureCitation id="42A92A24FF952705051E2186FC02D5D1" box="[863,941,682,707]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
C, 4A); ambulatory legs long, slender (
<figureCitation id="42A92A24FF95270503252186FF0AD5F5" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270506F521E3FF6BD5F5" box="[180,196,719,743]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF952705069121E3FF70D5F5" box="[208,223,719,743]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">4</figureCitation>
E); ambulatory merus elongated, slender, surface with margins rounded, not cristate, not distinctly granular, dorsal distal tooth distinct but low (
<figureCitation id="42A92A24FF952705046621DEFDDAD419" box="[551,629,754,779]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270504C521DFFD3BD419" box="[644,660,755,779]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF95270504E121DFFD00D419" box="[672,687,755,779]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">4</figureCitation>
E); ambulatory propodus, dactylus with ventral margins lined with dense stiff setae, with distinct dactylo-propodal lock, not forming subchelate structure (
<figureCitation id="42A92A24FF952705022A203AFB16D43D" box="[1131,1209,790,815]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF9527050286203BFB78D43D" box="[1223,1239,791,815]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF95270502A2203BFB5DD43D" box="[1251,1266,791,815]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">4</figureCitation>
D, E); cheliped merus rounded in cross-section; surface covered with numerous small pits (
<figureCitation id="42A92A24FF95270505A42016FB9DD441" box="[997,1074,826,851]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270502012017FBFFD441" box="[1088,1104,827,851]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF952705021D2017FBC5D441" box="[1116,1130,827,851]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">3</figureCitation>
C); cheliped carpus surface with long, gently curved spine on distal inner angle (
<figureCitation id="42A92A24FF95270504A82072FC98D465" box="[745,823,862,887]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270505072073FCF9D465" box="[838,854,863,887]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF95270505232073FCDFD465" box="[866,880,863,887]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">3</figureCitation>
F); chela unarmed, margins not cristate (
<figureCitation id="42A92A24FF95270503782072FAD4D465" box="[1337,1403,862,887]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Fig. 3</figureCitation>
F); thoracic sternum relatively narrow transversely, surfaces pitted, sternites 3, 4 completely fused with median depression (
<figureCitation id="42A92A24FF952705075F208AFEC3D4AD" box="[286,364,934,959]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
C, 4B); male abdominal locking mechanism present as knob-like tubercle on distal one-third of thoracic sternite 5; male abdomen broadly triangular (
<figureCitation id="42A92A24FF95270504AF20E6FC95D4F1" box="[750,826,970,995]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
D, 4C); G1 slender, sinuous, distal part elongated (
<figureCitation id="42A92A24FF952705032E20E6FF0AD315" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">Fig. 4</figureCitation>
F, G).
</paragraph>
</subSubSection>
<subSubSection id="9288652AFF9527050686273EFC2AD35D" pageId="1" pageNumber="95" type="etymology">
<paragraph id="DA2D36A1FF9527050686273EFC2AD35D" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="95">
<emphasis id="E8E6EAB3FF9527050686273EFEE9D339" bold="true" box="[199,326,1042,1067]" pageId="1" pageNumber="95">Etymology</emphasis>
. The new genus is named after Cebu City in the Visayas, 
<collectingCountry id="A2857631FF9527050589273EFBE5D339" box="[968,1098,1042,1067]" name="Philippines" pageId="1" pageNumber="95">Philippines</collectingCountry>
; in arbitrary combination with the genus name 
<taxonomicName id="1D924D22FF952705070D271AFE60D35D" box="[332,463,1078,1103]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF952705070D271AFE60D35D" box="[332,463,1078,1103]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
. The gender of the genus is masculine.
</paragraph>
</subSubSection>
<subSubSection id="9288652AFF95270606862776FD3ED5BA" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95" type="discussion">
<paragraph id="DA2D36A1FF95270506862776FB31D1A1" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="95">
<emphasis id="E8E6EAB3FF95270506862776FE9BD361" bold="true" box="[199,308,1114,1139]" pageId="1" pageNumber="95">Remarks</emphasis>
. The unusual combination of carapace, cheliped, ambulatory leg, thoracic sternal and abdominal characters in 
<taxonomicName id="1D924D22FF952705076F2752FD9BD384" box="[302,564,1150,1175]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="poppeorum" status="gen. nov.">
<emphasis id="E8E6EAB3FF952705076F2752FD9BD384" box="[302,564,1150,1175]" italics="true" pageId="1" pageNumber="95">Cebudonus poppeorum</emphasis>
</taxonomicName>
<emphasis id="E8E6EAB3FF952705047D2752FD74D385" bold="true" box="[572,731,1150,1175]" pageId="1" pageNumber="95">
<taxonomicNameLabel id="F3D557C8FF952705047D2752FD22D385" box="[572,653,1150,1175]" pageId="1" pageNumber="95" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
require the establishment of a new genus. While the carapace shape of 
<taxonomicName id="1D924D22FF95270506BD278EFED5D3A9" box="[252,378,1186,1211]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270506BD278EFED5D3A9" box="[252,378,1186,1211]" italics="true" pageId="1" pageNumber="95">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF95270507C0278EFE60D3A9" box="[385,463,1186,1211]" pageId="1" pageNumber="95" rank="genus">
<emphasis id="E8E6EAB3FF95270507C0278EFE60D3A9" bold="true" box="[385,463,1186,1211]" pageId="1" pageNumber="95">n. gen.</emphasis>
</taxonomicNameLabel>
superficially resembles species associated with echinoids like 
<taxonomicName id="1D924D22FF95270502CF278EFE51D3CD" authority="Chia &amp; Ng, 2000" authorityName="Chia &amp; Ng" authorityYear="2000" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="E8E6EAB3FF95270502CF278EFA33D3A9" box="[1166,1436,1186,1211]" italics="true" pageId="1" pageNumber="95">Eumedonus intermedius</emphasis>
<bibRefCitation id="BE034B50FF95270506D627EAFEF6D3CD" author="Chia" box="[151,345,1222,1247]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng, 2000</bibRefCitation>
(
<collectingCountry id="A2857631FF952705072927EAFE5AD3CD" box="[360,501,1222,1247]" name="Madagascar" pageId="1" pageNumber="95">Madagascar</collectingCountry>
)
</taxonomicName>
, 
<taxonomicName id="1D924D22FF952705044827EBFC07D3CD" authority="Rathbun, 1918" authorityName="Rathbun" authorityYear="1918" box="[521,936,1222,1247]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="vicinus">
<emphasis id="E8E6EAB3FF952705044827EBFDD8D3CC" box="[521,631,1223,1246]" italics="true" pageId="1" pageNumber="95">E. vicinus</emphasis>
Rathbun, 1918 (
<collectingCountry id="A2857631FF952705057527EAFC0FD3CD" box="[820,928,1222,1247]" name="Australia" pageId="1" pageNumber="95">Australia</collectingCountry>
)
</taxonomicName>
, and 
<taxonomicName id="1D924D22FF95270505A527EBFB76D3CD" authority="Alcock 1895" authorityName="Alcock" authorityYear="1895" box="[996,1241,1222,1247]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="species" species="zebra">
<emphasis id="E8E6EAB3FF95270505A527EBFBEDD3CD" box="[996,1090,1222,1247]" italics="true" pageId="1" pageNumber="95">E. zebra</emphasis>
Alcock 1895
</taxonomicName>
(Red Sea to East 
<collectingCountry id="A2857631FF95270506D627C6FF75D211" box="[151,218,1258,1283]" name="China" pageId="1" pageNumber="95">China</collectingCountry>
Sea), especially with regard to the shape of the laterally directed spines; its front is completely different, with two long pseudorostral spines (
<figureCitation id="42A92A24FF95270507BA2622FDE5D235" box="[507,586,1294,1319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF952705041B2623FDC6D235" box="[602,617,1295,1319]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
) in contrast to 
<taxonomicName id="1D924D22FF95270505582622FC33D235" box="[793,924,1294,1319]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270505582622FC33D235" box="[793,924,1294,1319]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
, which has a lobiform pseudorostrum that is bifurcated distally to form two teeth (cf. 
<bibRefCitation id="BE034B50FF952705041F261EFCB3D259" author="Chia" box="[606,796,1330,1355]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 8, 9A, D, 10, 11, 12A, D, N, 13, 14, 15A, D, Q, R). The lateral carapace tooth and the dorsal surface adjacent to it is gently convex dorsally in 
<taxonomicName id="1D924D22FF95270502CD267AFAA5D27D" box="[1164,1290,1366,1391]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270502CD267AFAA5D27D" box="[1164,1290,1366,1391]" italics="true" pageId="1" pageNumber="95">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF9527050350267AFAF0D27D" box="[1297,1375,1366,1391]" pageId="1" pageNumber="95" rank="genus">
<emphasis id="E8E6EAB3FF9527050350267AFAF0D27D" bold="true" box="[1297,1375,1366,1391]" pageId="1" pageNumber="95">n. gen.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="42A92A24FF952705032F267AFF0BD281" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Fig. 3</figureCitation>
A) (the tooth is stout with the dorsal surface distinctly convex in 
<taxonomicName id="1D924D22FF95270505CE2656FBBDD281" box="[911,1042,1402,1427]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270505CE2656FBBDD281" box="[911,1042,1402,1427]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF95270502C42650FB10D281" box="[1157,1215,1402,1427]" italics="true" pageId="1" pageNumber="95">et al.</emphasis>
1995: fig. 3A); the cheliped merus is relatively long and unarmed (
<figureCitation id="42A92A24FF952705048C26B2FCB0D2A5" box="[717,799,1438,1463]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF952705057326B3FCEDD2A5" box="[818,834,1439,1463]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF952705051226B3FCCFD2A5" box="[851,864,1439,1463]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">3</figureCitation>
C) (relatively short with numerous tubercles and granules in 
<taxonomicName id="1D924D22FF952705075A26EEFE31D2C9" box="[283,414,1474,1499]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF952705075A26EEFE31D2C9" box="[283,414,1474,1499]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF952705079226EEFD3CD2C9" author="Chia" box="[467,659,1474,1499]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 8, 9B, 10, 11, 12B, L, 14, 15B, O); the meri of the ambulatory legs are long, subcylindrical and smooth, without any marginal cristae, and the dorsal distal tooth is relatively low (
<figureCitation id="42A92A24FF95270506DE2526FF5CD131" box="[159,243,1546,1571]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270507482527FEB6D131" box="[265,281,1547,1571]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF952705076D2527FE95D131" box="[300,314,1547,1571]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">4</figureCitation>
E) (short, laterally flattened with distinct marginal cristae and a strong dorsal distal tooth in 
<taxonomicName id="1D924D22FF95270506D62502FEB5D155" box="[151,282,1582,1607]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270506D62502FEB5D155" box="[151,282,1582,1607]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF952705070D2502FDA6D155" author="Chia" box="[332,521,1582,1607]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 8, 9E, G, 10, 11, 12E, K, 13, 14, 15E, K); the anterior male thoracic sternum (sternites 1–4) are proportionately narrow transversely (
<figureCitation id="42A92A24FF952705056E257EFC2DD179" box="[815,898,1618,1643]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
C, 4B) (proportionately wider transversely in 
<taxonomicName id="1D924D22FF95270506D6255AFEB5D19D" box="[151,282,1654,1679]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270506D6255AFEB5D19D" box="[151,282,1654,1679]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF952705070F255AFDBED19D" author="Chia" box="[334,529,1654,1679]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 9C, 11B, 12C, 14B, 15N); and the male abdomen is relatively wide (
<figureCitation id="42A92A24FF9527050325255AFF0BD1A1" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="1" pageNumber="95">Figs. 3</figureCitation>
D, 4C) (proportionately narrower in 
<taxonomicName id="1D924D22FF952705047F25B6FD6ED1A1" box="[574,705,1690,1715]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF952705047F25B6FD6ED1A1" box="[574,705,1690,1715]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF95270504B225B6FC1ED1A1" author="Chia" box="[755,945,1690,1715]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 12J, 14J, 15N).
</paragraph>
<paragraph id="DA2D36A1FF95270506862592FB94D0BD" blockId="1.[151,1437,430,2003]" pageId="1" pageNumber="95">
The unarmed and non-cristate ambulatory meri of 
<taxonomicName id="1D924D22FF95270504BE2592FCD2D1C5" box="[767,893,1726,1751]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270504BE2592FCD2D1C5" box="[767,893,1726,1751]" italics="true" pageId="1" pageNumber="95">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF95270505C52592FC7CD1C5" box="[900,979,1726,1751]" pageId="1" pageNumber="95" rank="genus">
<emphasis id="E8E6EAB3FF95270505C52592FC7CD1C5" bold="true" box="[900,979,1726,1751]" pageId="1" pageNumber="95">n. gen.</emphasis>
</taxonomicNameLabel>
resembles that of the echinoid symbiont 
<taxonomicName id="1D924D22FF95270506D625CFFE87D1E8" box="[151,296,1763,1786]" class="Malacostraca" family="Pilumnidae" genus="Gonatonotus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270506D625CFFE87D1E8" box="[151,296,1763,1786]" italics="true" pageId="1" pageNumber="95">Gonatonotus</emphasis>
</taxonomicName>
, but in the new taxon, it is proportionately much longer, more slender and almost smooth (
<figureCitation id="42A92A24FF952705036A25CEFAD6D1E9" box="[1323,1401,1762,1787]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="1" pageNumber="95">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF95270503C625CFFA38D1E9" box="[1415,1431,1763,1787]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="1" pageNumber="95">2</figureCitation>
, 
<figureCitation id="42A92A24FF95270506D6242BFF0AD00D" box="[151,165,1799,1823]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="1" pageNumber="95">4</figureCitation>
E) (proportionately shorter and distinctly granular in 
<taxonomicName id="1D924D22FF95270504B6242BFC27D00C" box="[759,904,1799,1822]" class="Malacostraca" family="Pilumnidae" genus="Gonatonotus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270504B6242BFC27D00C" box="[759,904,1799,1822]" italics="true" pageId="1" pageNumber="95">Gonatonotus</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF95270505FB242AFBD6D00D" author="Chia" box="[954,1145,1798,1823]" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 17, 18E, G, 20, 21E, K, 22, 23E, K). The differences previously noted in the form of the front, anterolateral region, cheliped, thoracic sternum and abdomen between 
<taxonomicName id="1D924D22FF95270504442462FD2CD075" box="[517,643,1870,1895]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF95270504442462FD2CD075" box="[517,643,1870,1895]" italics="true" pageId="1" pageNumber="95">Cebudonus</emphasis>
</taxonomicName>
<emphasis id="E8E6EAB3FF95270504CC2462FD71D075" bold="true" box="[653,734,1870,1895]" pageId="1" pageNumber="95">
<taxonomicNameLabel id="F3D557C8FF95270504CC2462FD71D075" box="[653,734,1870,1895]" pageId="1" pageNumber="95" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="1D924D22FF952705055A2462FC31D075" box="[795,926,1870,1895]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF952705055A2462FC31D075" box="[795,926,1870,1895]" italics="true" pageId="1" pageNumber="95">Eumedonus</emphasis>
</taxonomicName>
also apply for 
<taxonomicName id="1D924D22FF95270502102463FB4DD074" box="[1105,1250,1871,1894]" class="Malacostraca" family="Pilumnidae" genus="Gonatonotus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270502102463FB4DD074" box="[1105,1250,1871,1894]" italics="true" pageId="1" pageNumber="95">Gonatonotus</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="BE034B50FF952705035A2462FF7BD099" author="Chia" pageId="1" pageNumber="95" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia &amp; Ng 2000</bibRefCitation>
: figs. 17, 18A, D, 19A, 20, 21A, D, M, N, 22, 23A, D [front]; figs. 17, 18B, 20, 21B, 22, 23B [chelipeds]; figs. 17B, 19G, 20B, 21C, 22B, 23C [sternum]; figs. 19H, 21J, 22, 23J [abdomen]).
</paragraph>
<paragraph id="DA2D36A1FF95270606862496FD3ED5BA" blockId="1.[151,1437,430,2003]" lastBlockId="2.[151,1437,151,1580]" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="95">
The form of the front, with the two long pseudorostral spines is similar to that of 
<taxonomicName id="1D924D22FF95270502152496FB74D0C1" box="[1108,1243,1978,2003]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270502152496FB74D0C1" box="[1108,1243,1978,2003]" italics="true" pageId="1" pageNumber="95">Tiaramedon</emphasis>
</taxonomicName>
and 
<taxonomicName id="1D924D22FF95270503502496FAC5D0C1" box="[1297,1386,1978,2003]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="95" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF95270503502496FAC5D0C1" box="[1297,1386,1978,2003]" italics="true" pageId="1" pageNumber="95">Zebrida</emphasis>
</taxonomicName>
, but these genera differ markedly from 
<taxonomicName id="1D924D22FF962706047223BBFD1ED7A2" box="[563,689,151,176]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF962706047223BBFD1ED7A2" box="[563,689,151,176]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF96270604FD23BBFCA0D7A2" box="[700,783,151,176]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF96270604FD23BBFCA0D7A2" bold="true" box="[700,783,151,176]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
in several other characters. 
<taxonomicName id="1D924D22FF962706022123BBFB71D7A2" box="[1120,1246,151,176]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF962706022123BBFB71D7A2" box="[1120,1246,151,176]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF96270602A823BBFA93D7A2" box="[1257,1340,151,176]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF96270602A823BBFA93D7A2" bold="true" box="[1257,1340,151,176]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
, can be separated from 
<taxonomicName id="1D924D22FF96270607042390FE63D7C7" box="[325,460,188,213]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270607042390FE63D7C7" box="[325,460,188,213]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
by its smooth dorsal surface of the carapace (
<figureCitation id="42A92A24FF96270605912390FBB1D7C6" box="[976,1054,188,213]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF962706026D2390FB94D7C6" box="[1068,1083,188,212]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
) (armed with additional gastric and branchial spines in 
<taxonomicName id="1D924D22FF96270607F023F3FD97D7EA" box="[433,568,223,248]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270607F023F3FD97D7EA" box="[433,568,223,248]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf 
<bibRefCitation id="BE034B50FF962706042D23F3FC98D7EA" author="Chia" box="[620,823,223,248]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: figs. 7A, 8A); the two pseudorostral spines joining medially without any trace of a median plate (
<figureCitation id="42A92A24FF96270604F32228FCADD60E" box="[690,770,260,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270605522228FC8DD60E" box="[787,802,260,284]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
) (with a small plate present between the pseudorostral spines in 
<taxonomicName id="1D924D22FF9627060745220BFE24D652" box="[260,395,295,320]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF9627060745220BFE24D652" box="[260,395,295,320]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF9627060780220BFD2BD652" author="Chia" box="[449,644,295,320]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: fig. 8A, B); the chela is unarmed (
<figureCitation id="42A92A24FF962706025B220BFBC6D652" box="[1050,1129,295,320]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270602392204FB27D652" box="[1144,1160,296,320]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF96270602D72204FB0CD652" box="[1174,1187,296,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">3</figureCitation>
F) (with an additional spine on the dorsal distal margin in 
<taxonomicName id="1D924D22FF962706046F2260FD1AD677" box="[558,693,332,357]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706046F2260FD1AD677" box="[558,693,332,357]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF96270604AB2260FC03D676" author="Chia" box="[746,940,332,357]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: figs. 7A, B, 8K); the cheliped carpus has a single long inner spine (
<figureCitation id="42A92A24FF96270607E22243FE5FD69A" box="[419,496,367,392]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270607BF225CFDA1D69A" box="[510,526,368,392]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF9627060458225CFD89D69A" box="[537,550,368,392]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">3</figureCitation>
F) (2 large spines in 
<taxonomicName id="1D924D22FF962706054D2243FC3CD69A" box="[780,915,367,392]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706054D2243FC3CD69A" box="[780,915,367,392]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF96270605842243FB2DD69A" author="Chia" box="[965,1154,367,392]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: fig. 7A); the ambulatory merus is long and slender with a short dorsal distal tooth (
<figureCitation id="42A92A24FF962706056822B8FCD8D6BE" box="[809,887,404,429]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270605C722B8FC39D6BE" box="[902,918,404,428]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF96270605E222B8FC1ED6BE" box="[931,945,404,428]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="2" pageNumber="96">4</figureCitation>
E) (short and stout with a long distal dorsal tooth in 
<taxonomicName id="1D924D22FF96270606BA229BFE2DD6C2" box="[251,386,439,464]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270606BA229BFE2DD6C2" box="[251,386,439,464]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF96270607FD229BFD2AD6C2" author="Chia" box="[444,645,439,464]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: figs. 7, 8C); and the male anterior thoracic sternum is transversely narrow (
<figureCitation id="42A92A24FF96270606B722F0FEECD6E6" box="[246,323,476,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
C, 4B) (transversely broad in 
<taxonomicName id="1D924D22FF96270604D222F0FCB5D6E7" box="[659,794,476,501]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270604D222F0FCB5D6E7" box="[659,794,476,501]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="BE034B50FF962706050C22F0FBA2D6E6" author="Chia" box="[845,1037,476,501]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: figs. 7B, 8D). The male abdomens of 
<taxonomicName id="1D924D22FF96270606F722D3FE9BD50A" box="[182,308,511,536]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF96270606F722D3FE9BD50A" box="[182,308,511,536]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF962706077D22D3FE23D50A" box="[316,396,511,536]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF962706077D22D3FE23D50A" bold="true" box="[316,396,511,536]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="1D924D22FF962706078422D3FDE3D50A" box="[453,588,511,536]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706078422D3FDE3D50A" box="[453,588,511,536]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
are nevertheless similar in shape (
<figureCitation id="42A92A24FF962706059722D3FB8CD50A" box="[982,1059,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
D, 4C; cf. 
<bibRefCitation id="BE034B50FF96270602DF22D3FAF0D50A" author="Chia" box="[1182,1375,511,536]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (1998) A revision of Ceratocarcinus White, 1847, and Harrovia Adams &amp; White, 1849 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46 (2), 493 - 563." type="journal article" year="1998">Chia &amp; Ng 1998</bibRefCitation>
: figs. 7B, 8F). The two genera are also different in terms of host specificity, with 
<taxonomicName id="1D924D22FF96270602442108FB22D52F" box="[1029,1165,548,573]" class="Malacostraca" family="Pilumnidae" genus="Tiaramedon" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270602442108FB22D52F" box="[1029,1165,548,573]" italics="true" pageId="2" pageNumber="96">Tiaramedon</emphasis>
</taxonomicName>
always associated with crinoids (and with young crabs characteristically transversely striped like many other crinoid-associated eumedonines, see 
<bibRefCitation id="BE034B50FF96270607282140FD93D597" author="Castro" box="[361,572,620,645]" pageId="2" pageNumber="96" refString="Castro, P., Chia, D. G. B. &amp; Ng, P. K. L. (1995) On the taxonomic status of Ceratocarcinus longimanus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), a crab symbiotic with comatulid crinoids. Raffles Bulletin of Zoology, 43 (1), 239 - 250." type="journal article" year="1995">
Castro 
<emphasis id="E8E6EAB3FF96270607FD2141FE57D597" box="[444,504,620,645]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995
</bibRefCitation>
; Chia &amp; Ng 1995, 1998) whereas 
<taxonomicName id="1D924D22FF96270605902140FBE0D597" box="[977,1103,620,645]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF96270605902140FBE0D597" box="[977,1103,620,645]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF96270602182140FB05D597" box="[1113,1194,620,645]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF96270602182140FB05D597" bold="true" box="[1113,1194,620,645]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
with its longitudinal stripes is probably associated with echinoids.
</paragraph>
</subSubSection>
<subSubSection id="9288652AFF96270606862198FF6AD13E" pageId="2" pageNumber="96" type="reference_group">
<paragraph id="DA2D36A1FF96270606862198FAD5D34A" blockId="2.[151,1437,151,1580]" pageId="2" pageNumber="96">
<taxonomicName id="1D924D22FF96270606862198FEEAD5DF" ID-CoL="7NWMK" box="[199,325,692,717]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF96270606862198FEEAD5DF" box="[199,325,692,717]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF96270607132198FE08D5DF" box="[338,423,692,717]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF96270607132198FE08D5DF" bold="true" box="[338,423,692,717]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
can be separated from the echinoid symbiont 
<taxonomicName id="1D924D22FF962706059C2198FB99D5DF" ID-CoL="7Q6XB" box="[989,1078,692,717]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706059C2198FB99D5DF" box="[989,1078,692,717]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
by having two subcylindrical pseudorostral spines (
<figureCitation id="42A92A24FF96270607CA21FBFE77D5E2" box="[395,472,727,752]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270607A721F4FE5AD5E2" box="[486,501,728,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
) (distinctly lamelliform in 
<taxonomicName id="1D924D22FF962706056721FBFCD0D5E2" ID-CoL="7Q6XB" box="[806,895,727,752]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706056721FBFCD0D5E2" box="[806,895,727,752]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF96270605B821FBFBDED5E2" author="Chia" box="[1017,1137,727,752]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2, 3A, B); the lateral carapace tooth and the dorsal surface adjacent to the tooth is dorsally convex (
<figureCitation id="42A92A24FF962706027921D0FBD1D406" box="[1080,1150,764,789]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Fig. 3</figureCitation>
B) (the area is distinctly dorsally flattened in 
<taxonomicName id="1D924D22FF96270607C12033FE76D42A" ID-CoL="7Q6XB" box="[384,473,799,824]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270607C12033FE76D42A" box="[384,473,799,824]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF962706040B200DFD2CD42A" box="[586,643,799,824]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: fig. 3B); the chela and merus are unarmed and the carpus has a long spine at the inner angle (
<figureCitation id="42A92A24FF96270607A32068FD80D44E" box="[482,559,836,861]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF962706047C2068FDE2D44E" box="[573,589,836,860]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF96270604192068FDCAD44E" box="[600,613,836,860]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">3</figureCitation>
F) (the chela has a large subdistal dorsal tooth, the carpus and merus each having three large lamelliform teeth in 
<taxonomicName id="1D924D22FF962706040C204BFD09D492" ID-CoL="7Q6XB" box="[589,678,871,896]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706040C204BFD09D492" box="[589,678,871,896]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF962706055F204BFC38D492" author="Chia" box="[798,919,871,896]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2, 3J); the ambulatory merus is long and slender with a short dorsal distal tooth (
<figureCitation id="42A92A24FF962706041020A0FD31D4B6" box="[593,670,908,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270604ED20A0FD13D4B6" box="[684,700,908,932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF962706048920A0FD79D4B6" box="[712,726,908,932]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="2" pageNumber="96">4</figureCitation>
E) (short and stout with the margins cristate to dentate and with a large distal dorsal tooth in 
<taxonomicName id="1D924D22FF96270607902083FD85D4DA" ID-CoL="7Q6XB" box="[465,554,943,968]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270607902083FD85D4DA" box="[465,554,943,968]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF96270604DE2083FCB8D4DA" author="Chia" box="[671,791,943,968]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2, 3C, K, M); the ambulatory propodus and dactylus are normal (
<figureCitation id="42A92A24FF962706075E20F8FEC3D4FE" box="[287,364,980,1005]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF962706073B20F8FE25D4FE" box="[378,394,980,1004]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF96270607D720F8FE0AD4FE" box="[406,421,980,1004]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="2" pageNumber="96">4</figureCitation>
D, E) (forming a subchelate clasping structure in 
<taxonomicName id="1D924D22FF962706058D20F8FB8AD4FF" ID-CoL="7Q6XB" box="[972,1061,980,1005]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706058D20F8FB8AD4FF" box="[972,1061,980,1005]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF96270602DF20F8FAB9D4FE" author="Chia" box="[1182,1302,980,1005]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2, 3C, K, M); and the male anterior thoracic sternum is proportionately narrower transversely (
<figureCitation id="42A92A24FF96270602AE20DBFAE8D302" box="[1263,1351,1015,1040]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
C, 4B) (proportionately wider in 
<taxonomicName id="1D924D22FF96270607FA2730FDBBD327" ID-CoL="7Q6XB" box="[443,532,1052,1077]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270607FA2730FDBBD327" box="[443,532,1052,1077]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF96270604D02730FCA4D326" author="Chia" box="[657,779,1052,1077]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2B, 3D). The male abdomens of 
<taxonomicName id="1D924D22FF96270602872730FAEBD327" ID-CoL="7NWMK" box="[1222,1348,1052,1077]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF96270602872730FAEBD327" box="[1222,1348,1052,1077]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<emphasis id="E8E6EAB3FF962706030D2730FA33D327" bold="true" box="[1356,1436,1052,1077]" pageId="2" pageNumber="96">
<taxonomicNameLabel id="F3D557C8FF962706030D2730FA33D327" box="[1356,1436,1052,1077]" pageId="2" pageNumber="96" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="1D924D22FF96270606872713FEB0D34A" ID-CoL="7Q6XB" box="[198,287,1087,1112]" class="Malacostraca" family="Pilumnidae" genus="Zebrida" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270606872713FEB0D34A" box="[198,287,1087,1112]" italics="true" pageId="2" pageNumber="96">Zebrida</emphasis>
</taxonomicName>
are similar, with both relatively wide transversely (
<figureCitation id="42A92A24FF96270605212713FC02D34A" box="[864,941,1087,1112]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
D, 4C; cf. Ng &amp; 
<bibRefCitation id="BE034B50FF96270602282713FB4ED34A" author="Chia" box="[1129,1249,1087,1112]" pageId="2" pageNumber="96" refString="Chia, D. G. B. &amp; Ng, P. K. L. (2000) A revision of Eumedonus H. Milne Edwards, 1834 and Gonatonotus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), two genera of crabs symbiotic with sea urchins. Journal of Natural History, 34, 15 - 56. http: // dx. doi. org / 10.1080 / 002229300299679" type="journal article" year="2000">Chia 2000</bibRefCitation>
: figs. 2B, 3F)
</paragraph>
<paragraph id="DA2D36A1FF96270606862748FF6AD13E" blockId="2.[151,1437,151,1580]" pageId="2" pageNumber="96">
There are also similarities with 
<taxonomicName id="1D924D22FF96270604102748FD7ED36F" ID-CoL="7Q6XC" box="[593,721,1124,1149]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270604102748FD7ED36F" box="[593,721,1124,1149]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
but 
<taxonomicName id="1D924D22FF96270605532748FC3FD36F" ID-CoL="7NWMK" box="[786,912,1124,1149]" class="Malacostraca" family="Pilumnidae" genus="Cebudonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E8E6EAB3FF96270605532748FC3FD36F" box="[786,912,1124,1149]" italics="true" pageId="2" pageNumber="96">Cebudonus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F3D557C8FF96270605DE2748FC59D36F" box="[927,1014,1124,1149]" pageId="2" pageNumber="96" rank="genus">
<emphasis id="E8E6EAB3FF96270605DE2748FC59D36F" bold="true" box="[927,1014,1124,1149]" pageId="2" pageNumber="96">n. gen.</emphasis>
</taxonomicNameLabel>
can be separated by its two long pseudorostral spines (
<figureCitation id="42A92A24FF96270607DB27ABFE40D3B2" box="[410,495,1159,1184]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF962706044427A4FDBBD3B2" box="[517,532,1160,1184]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
) (with a distinctly dorsoventrally flattened lobiform pseudorostrum that is bifurcated distally to form two teeth in 
<taxonomicName id="1D924D22FF96270604122780FD79D3D7" ID-CoL="7PF77" box="[595,726,1196,1221]" class="Malacostraca" family="Pilumnidae" genus="Eumedonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270604122780FD79D3D7" box="[595,726,1196,1221]" italics="true" pageId="2" pageNumber="96">Eumedonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF96270605072781FCD0D3D7" box="[838,895,1196,1221]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: figs. 1, 2A, K); the lateral carapace tooth and the dorsal surface adjacent to it is dorsally convex (
<figureCitation id="42A92A24FF962706057A27E3FC2DD3FA" box="[827,898,1231,1256]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Fig. 3</figureCitation>
B) (the area is distinctly dorsally flattened in 
<taxonomicName id="1D924D22FF96270606D627D8FEB9D21F" ID-CoL="7Q6XC" box="[151,278,1268,1293]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF96270606D627D8FEB9D21F" box="[151,278,1268,1293]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF96270607C527D9FE13D21F" box="[388,444,1268,1293]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: figs. 1, 2A, 3C); the chela and merus are unarmed, and the carpus has a long spine on the inner angle (
<figureCitation id="42A92A24FF9627060735263BFE6DD222" box="[372,450,1303,1328]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF96270607902634FE4ED222" box="[465,481,1304,1328]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF96270607AC2634FE55D222" box="[493,506,1304,1328]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">3</figureCitation>
F) (the chela has a distinct distal dorsal tooth, the carpus 2 large lamelliform teeth and the merus several distinct teeth in 
<taxonomicName id="1D924D22FF962706040A2610FD65D247" ID-CoL="7Q6XC" box="[587,714,1340,1365]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706040A2610FD65D247" box="[587,714,1340,1365]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF962706057D2611FCDAD247" box="[828,885,1340,1365]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: figs. 1, 
<date id="AE2C1061FF962706025B2610FB85D246" box="[1050,1066,1340,1364]" pageId="2" pageNumber="96">2</date>
I); the ambulatory merus is long, subcylindrical and smooth, without any marginal cristae and the dorsal distal tooth is relatively low (
<figureCitation id="42A92A24FF96270602BC2673FAE5D26A" box="[1277,1354,1375,1400]" captionStart="FIGURE 1" captionStartId="3.[151,250,1940,1962]" captionTargetBox="[151,1436,897,1918]" captionTargetId="figure@3.[151,1436,897,1918]" captionTargetPageId="3" captionText="FIGURE 1. Cebudonus poppeorum n. gen., n. sp. Colour in life, female, Olango Channel, Cebu, Visayas, Philippines, 380 m, coll. 2012 (specimen not extant) (photograph by G. Poppe)." httpUri="https://zenodo.org/record/226600/files/figure.png" pageId="2" pageNumber="96">Figs. 1</figureCitation>
, 
<figureCitation id="42A92A24FF9627060319264CFAC7D26A" box="[1368,1384,1376,1400]" captionStart="FIGURE 2" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[193,1393,193,1930]" captionTargetId="figure@4.[193,1393,193,1932]" captionTargetPageId="4" captionText="FIGURE 2. Cebudonus poppeorum n. gen., n. sp., overall dorsal views. A, holotype male (10.8 × 11.9 mm) (NMCR); B, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines." httpUri="https://zenodo.org/record/226601/files/figure.png" pageId="2" pageNumber="96">2</figureCitation>
, 
<figureCitation id="42A92A24FF9627060335264CFA2DD26A" box="[1396,1410,1376,1400]" captionStart="FIGURE 4" captionStartId="6.[151,250,1917,1939]" captionTargetBox="[230,1347,203,1883]" captionTargetId="figure@6.[215,1362,193,1898]" captionTargetPageId="6" captionText="FIGURE 4. Cebudonus poppeorum n. gen., n. sp. A – D, F – H, holotype male (10.8 × 11.9 mm) (NMCR); E, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211). Specimens from Cebu, Philippines. A, right third maxilliped (setae not drawn); B, male anterior thoracic sternum (sternites 1 – 4) and telson; C, male abdomen; D, right third ambulatory propodus and dactylus; E, left fourth ambulatory leg; F, ventral view of left G 1; dorsal view of left G 1; H, left G 2. Scales: A – E = 1.0 mm; F – H = 0.5 mm." httpUri="https://zenodo.org/record/226603/files/figure.png" pageId="2" pageNumber="96">4</figureCitation>
E) (short, laterally flattened with distinct marginal cristae and a strong dorsal distal tooth in 
<taxonomicName id="1D924D22FF962706023626A8FB59D28F" ID-CoL="7Q6XC" box="[1143,1270,1412,1437]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706023626A8FB59D28F" box="[1143,1270,1412,1437]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF962706032526A9FA33D28F" box="[1380,1436,1412,1437]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: figs. 1, 2B, C); the anterior male thoracic sternum (sternites 1–4) are proportionately narrower transversely (
<figureCitation id="42A92A24FF96270606DE26E0FF5FD2F6" box="[159,240,1484,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
C, 4B) (proportionately wider transversely in 
<taxonomicName id="1D924D22FF962706054826E0FC27D2F7" ID-CoL="7Q6XC" box="[777,904,1484,1509]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706054826E0FC27D2F7" box="[777,904,1484,1509]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF962706024226E1FBEFD2F7" box="[1027,1088,1484,1509]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: fig. 2D); and the male abdomen is proportionately wider (
<figureCitation id="42A92A24FF962706046126C3FDC3D11A" box="[544,620,1519,1544]" captionStart="FIGURE 3" captionStartId="5.[151,250,1469,1491]" captionTargetBox="[177,1414,183,1450]" captionTargetId="figure@5.[177,1415,183,1451]" captionTargetPageId="5" captionText="FIGURE 3. Cebudonus poppeorum n. gen., n. sp. A – E, holotype male (10.8 × 11.9 mm) (NMCR); G, paratype female (10.6 × 12.5 mm) (ZRC 2014.0211); H, paratype female (10.9 × 12.4 mm) (ZRC 2014.0214). Specimens from Philippines. A, frontal view of carapace; B, front, antennae, antennules, epistome and orbits; C, anterior thoracic sternum and male abdomen; D, male anterior thoracic sternum and abdomen; E, first two male abdominal somites; F, outer view of left chela; G, female thoracic sternum and abdomen; H, female thoracic sternum showing vulvae." httpUri="https://zenodo.org/record/226602/files/figure.png" pageId="2" pageNumber="96">Figs. 3</figureCitation>
D, 4C) (proportionately narrower in 
<taxonomicName id="1D924D22FF962706024526C3FB2BD11A" ID-CoL="7Q6XC" box="[1028,1156,1519,1544]" class="Malacostraca" family="Pilumnidae" genus="Zebridonus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="E8E6EAB3FF962706024526C3FB2BD11A" box="[1028,1156,1519,1544]" italics="true" pageId="2" pageNumber="96">Zebridonus</emphasis>
</taxonomicName>
; cf. Chia 
<emphasis id="E8E6EAB3FF96270602B026DDFA86D11A" box="[1265,1321,1519,1544]" italics="true" pageId="2" pageNumber="96">et al.</emphasis>
1995: fig. 2F).
</paragraph>
</subSubSection>
</treatment>
</document>