401 lines
47 KiB
XML
401 lines
47 KiB
XML
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<mods:title id="26AF9819C7D075762C1726962950D067">Revision of the dwarf cannibal snails (Nata s. l.) of southern Africa — Nata s. s. and Natella (Mollusca: Gastropoda: Rhytididae), with description of three new species</mods:title>
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<mods:title id="F48A763140C5D74D0D8E41AC5E7705C3">Zootaxa</mods:title>
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<paragraph id="8BF036EAE03CD776FF1A131609DD9ABE" blockId="49.[151,727,635,695]" box="[151,727,635,662]" pageId="49" pageNumber="50">
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<heading id="D0B88186E03CD776FF1A131609DD9ABE" bold="true" box="[151,727,635,662]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
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<taxonomicName id="4C4F4D69E03CD776FF1A131609DD9ABE" ID-CoL="6S5QP" authority="Melvill & Ponsonby, 1891" authorityName="Melvill & Ponsonby" authorityYear="1891" box="[151,727,635,662]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens">
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<emphasis id="B93BEAF8E03CD776FF1A131609DD9ABE" bold="true" box="[151,727,635,662]" pageId="49" pageNumber="50">
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<emphasis id="B93BEAF8E03CD776FF1A13160A719ABD" bold="true" box="[151,379,635,661]" italics="true" pageId="49" pageNumber="50">Natella viridescens</emphasis>
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(
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<bibRefCitation id="EFDE4B1BE03CD776FE06131109C59ABE" author="Melvill" box="[395,719,636,662]" pageId="49" pageNumber="57" refString="Melvill, J. C. & Ponsonby, J. H. (1891) Descriptions of nine new terrestrial and fluviatile mollusks from South Africa. Annals and Magazine of Natural History, Series 6, 8, 237 - 240. http: // dx. doi. org / 10.1080 / 00222939109460425" type="journal article" year="1891">Melvill & Ponsonby, 1891</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BF036EAE03CD776FF1A13F3095A9A9F" blockId="49.[151,727,635,695]" box="[151,592,670,695]" pageId="49" pageNumber="50">
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<figureCitation id="13742A6FE03CD776FF1A13F30A1C9A9F" box="[151,278,670,695]" captionStart="FIGURE 2. A" captionStartId="6.[151,250,1952,1974]" captionTargetBox="[154,1433,1444,1921]" captionTargetId="figure@6.[151,1436,1439,1923]" captionTargetPageId="6" captionText="FIGURE 2. A, Natella viridescens (Melvill & Ponsonby, 1891), drowning individual with proboscis everted, Ithala Game Res., KwaZulu-Natal, shell diameter 3.7 mm (NMSA W 7263). B, Nata vernicosa (Krauss, 1848), mating pair, Qora River mouth, E. Cape, respective shell diameter 20.5 mm and 14 mm (NMSA W 4015)." httpUri="https://zenodo.org/record/266546/files/figure.png" pageId="49" pageNumber="50">Figures 2A</figureCitation>
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,
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<figureCitation id="13742A6FE03CD776FEAF13F20A259A9F" box="[290,303,671,695]" captionStart="FIGURE 3" captionStartId="8.[151,250,1532,1554]" captionTargetBox="[157,1429,235,1508]" captionTargetId="figure@8.[151,1436,229,1511]" captionTargetPageId="8" captionText="FIGURE 3. Protoconchs of Nata and Natella. A, Nata dumeticola, Kommetjie, W. Cape (NMSA 2767). B, Nata tarachodes, Spes Bona bush, Cape Peninsula (NMSA V 9848). C, Nata vernicosa, Pietermaritzburg, KwaZulu-Natal (NMSA A 7114). D, Natella viridescens, Ithala Game Res., KwaZulu-Natal (NMSA V 2530). Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/266547/files/figure.png" pageId="49" pageNumber="50">3</figureCitation>
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D, 4H, 5C, 6F, 9C, 37–40
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3556561E03CD776FF1A138E0F6F9B99" pageId="49" pageNumber="50" type="reference_group">
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<paragraph id="8BF036EAE03CD776FF1A138E0E5F9AD1" blockId="49.[151,1437,739,945]" box="[151,1365,739,761]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="AB5F11C4E03CD776FF1A138E0E5F9AD1" box="[151,1365,739,761]" pageId="49" pageNumber="50">
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<taxonomicName id="4C4F4D69E03CD776FF1A138E09E39AD1" authority="Melvill & Ponsonby, 1891: 238" authorityName="Melvill & Ponsonby" authorityPageNumber="238" authorityYear="1891" box="[151,745,739,761]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens" subGenus="Patula">
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<emphasis id="B93BEAF8E03CD776FF1A138E0BC49AD1" box="[151,206,739,761]" italics="true" pageId="49" pageNumber="50">Helix</emphasis>
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(
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<emphasis id="B93BEAF8E03CD776FF51138E0A2A9AD1" box="[220,288,739,761]" italics="true" pageId="49" pageNumber="50">Patula</emphasis>
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||
)
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<emphasis id="B93BEAF8E03CD776FEA2138E0AAB9AD1" box="[303,417,739,761]" italics="true" pageId="49" pageNumber="50">viridescens</emphasis>
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<treatmentCitation id="0AEE10FBE03CD776FE2B138E09E39AD1" author="Melvill" box="[422,745,739,761]" page="238" pageId="49" pageNumber="50" year="1891">
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<bibRefCitation id="EFDE4B1BE03CD776FE2B138E09BC9AD1" author="Melvill" box="[422,694,739,761]" pageId="49" pageNumber="57" refString="Melvill, J. C. & Ponsonby, J. H. (1891) Descriptions of nine new terrestrial and fluviatile mollusks from South Africa. Annals and Magazine of Natural History, Series 6, 8, 237 - 240. http: // dx. doi. org / 10.1080 / 00222939109460425" type="journal article" year="1891">Melvill & Ponsonby, 1891</bibRefCitation>
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: 238
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</treatmentCitation>
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</taxonomicName>
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||
.
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<bibRefCitation id="EFDE4B1BE03CD776FD7E138E0F079AD1" author="Melvill" box="[755,1037,739,761]" pageId="49" pageNumber="57" refString="Melvill, J. C. & Ponsonby, J. H. (1892 a) Descriptions of seven new species of terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 6, 9, 84 - 87. http: // dx. doi. org / 10.1080 / 00222939208677277" type="journal article" year="1892" yearSuffix="a">Melvill & Ponsonby, 1892a</bibRefCitation>
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||
, pl. 4, fig. 1.
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||
<typeStatus id="54F48848E03CD776FB1E13890FCF9AD1" box="[1171,1221,740,761]" pageId="49" pageNumber="50">Type</typeStatus>
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loc.: Pretoria.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF036EAE03CD776FF1A126F084C9B30" blockId="49.[151,1437,739,945]" box="[151,838,770,792]" pageId="49" pageNumber="50">
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<taxonomicName id="4C4F4D69E03CD776FF1A126F0A599B30" box="[151,339,770,792]" class="Insecta" family="Urodidae" genus="Patula" kingdom="Animalia" order="Lepidoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="viridescens">
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<emphasis id="B93BEAF8E03CD776FF1A126F0A599B30" box="[151,339,770,792]" italics="true" pageId="49" pageNumber="50">Patula viridescens</emphasis>
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</taxonomicName>
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—Pilsbry,
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||
<quantity id="4CB79B0FE03CD776FE33126F09019B30" box="[446,523,770,792]" metricMagnitude="1" metricUnit="m" metricValue="4.80568" pageId="49" pageNumber="50" unit="in" value="1892.0">1892 in</quantity>
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<date id="FFF1102AE03CD776FD9F126F098D9B30" box="[530,647,770,792]" pageId="49" pageNumber="50" value="1892" valueMax="1893">1892–1893</date>
|
||
: 78, pl. 30, fig. 32.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF1A124C08A99B1F" blockId="49.[151,1437,739,945]" box="[151,931,801,823]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="AB5F11C4E03CD776FF1A124C08A99B1F" box="[151,931,801,823]" pageId="49" pageNumber="50">
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<taxonomicName id="4C4F4D69E03CD776FF1A124C0A629B1F" box="[151,360,801,823]" class="Gastropoda" family="Rhytididae" genus="Natalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03CD776FF1A124C0A629B1F" box="[151,360,801,823]" italics="true" pageId="49" pageNumber="50">Natalina viridescens</emphasis>
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</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0AEE10FBE03CD776FE0F124C09DA9B1F" author="Melvill" box="[386,720,801,823]" page="171" pageId="49" pageNumber="50" year="1898">
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||
<bibRefCitation id="EFDE4B1BE03CD776FE0F124C09949B1F" author="Melvill" box="[386,670,801,823]" pageId="49" pageNumber="58" refString="Melvill, J. C. & Ponsonby, J. H. (1898 b) A contribution towards a check-list of the non-marine molluscan fauna of South Africa. Proceedings of the Malacological Society of London, 3, 166 - 184." type="journal article" year="1898" yearSuffix="b">Melvill & Ponsonby, 1898b</bibRefCitation>
|
||
: 171
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||
</treatmentCitation>
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||
.
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<treatmentCitation id="0AEE10FBE03CD776FD57124C08959B1F" author="Connolly" box="[730,927,801,823]" page="98" pageId="49" pageNumber="50" year="1912">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FD57124C08739B1F" author="Connolly" box="[730,889,801,823]" pageId="49" pageNumber="57" refString="Connolly, M. (1912) Revised reference list of South African non-marine Mollusca; with descriptions of new species in the South African Museum. Annals of the South African Museum, 11, 9 - 306. http: // dx. doi. org / 10.5962 / bhl. title. 11477" type="journal article" year="1912">Connolly, 1912</bibRefCitation>
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||
: 98
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF1A12520E6B9B7D" blockId="49.[151,1437,739,945]" box="[151,1377,831,853]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="AB5F11C4E03CD776FF1A12520E6B9B7D" box="[151,1377,831,853]" pageId="49" pageNumber="50">
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<taxonomicName id="4C4F4D69E03CD776FF1A1252091C9B7D" box="[151,534,831,853]" class="Gastropoda" family="Rhytididae" genus="Rhytida" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens" subGenus="Macrocycloides">
|
||
<emphasis id="B93BEAF8E03CD776FF1A12520BEC9B7D" box="[151,230,831,853]" italics="true" pageId="49" pageNumber="50">Rhytida</emphasis>
|
||
(
|
||
<emphasis id="B93BEAF8E03CD776FF7912520A9F9B7D" box="[244,405,831,853]" italics="true" pageId="49" pageNumber="50">Macrocycloides</emphasis>
|
||
)
|
||
<emphasis id="B93BEAF8E03CD776FE291252091C9B7D" box="[420,534,831,853]" italics="true" pageId="49" pageNumber="50">viridescens</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EFDE4B1BE03CD776FDA21252085B9B7D" author="Mollendorff" box="[559,849,831,853]" pageId="49" pageNumber="58" refString="Mollendorff, O. von & Kobelt, W. (1903 - 1905) Die Raublungenschnecken (Agnatha). Abtheilung 1. Rhytididae & Enneidae. In: Kuster, H. C. Martini, F. H. W. & Chemnitz, J. H. (Eds.), Systematisches Conchylien-Cabinet. Vol. 1. 2 nd Edition. Part 12 b. Bauer & Raspe, Nurnburg, pp. 1 - 362, pls. 1 - 41. [Pp. 1 - 72, pls 1 - 12 = 1903; pp. 73 - 296, pls 13 - 35 = 1904; pp. 297 - 362, pls 36 - 41 = 1905, dates fide Bouchet & Rocroi, 2005.]" type="journal article" year="1903">Möllendorff & Kobelt, 1903</bibRefCitation>
|
||
, in
|
||
<date id="FFF1102AE03CD776FCF8125208E19B7D" box="[885,1003,831,853]" pageId="49" pageNumber="50" value="1903" valueMax="1905">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FCF8125208A39B7D" author="Mollendorff" box="[885,937,831,853]" pageId="49" pageNumber="58" refString="Mollendorff, O. von & Kobelt, W. (1903 - 1905) Die Raublungenschnecken (Agnatha). Abtheilung 1. Rhytididae & Enneidae. In: Kuster, H. C. Martini, F. H. W. & Chemnitz, J. H. (Eds.), Systematisches Conchylien-Cabinet. Vol. 1. 2 nd Edition. Part 12 b. Bauer & Raspe, Nurnburg, pp. 1 - 362, pls. 1 - 41. [Pp. 1 - 72, pls 1 - 12 = 1903; pp. 73 - 296, pls 13 - 35 = 1904; pp. 297 - 362, pls 36 - 41 = 1905, dates fide Bouchet & Rocroi, 2005.]" type="journal article" year="1903">1903</bibRefCitation>
|
||
–1905
|
||
</date>
|
||
: 61, pl. 10, fig. 26.
|
||
<treatmentCitation id="0AEE10FBE03CD776FB2212520E579B7D" author="Kobelt" box="[1199,1373,831,853]" page="53" pageId="49" pageNumber="50" year="1909">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FB2212520E3D9B7D" author="Kobelt" box="[1199,1335,831,853]" pageId="49" pageNumber="57" refString="Kobelt, W. (1909) Die Molluskenausbeute der erlangerschen Reise in Nordost-Afrika. Zweite Abteilung. Verzeichnis der aus Afrika bekannten Binnenconchylien. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 32, 53 - 97." type="journal article" year="1909">Kobelt, 1909</bibRefCitation>
|
||
: 53
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF1A1232090B9BBB" blockId="49.[151,1437,739,945]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="AB5F11C4E03CD776FF1A1232090B9BBB" pageId="49" pageNumber="50">
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<taxonomicName id="4C4F4D69E03CD776FF1A12320AA39B5C" box="[151,425,862,884]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens" subGenus="Natella">
|
||
<emphasis id="B93BEAF8E03CD776FF1A12320BC29B5C" box="[151,200,863,884]" italics="true" pageId="49" pageNumber="50">Nata</emphasis>
|
||
(
|
||
<emphasis id="B93BEAF8E03CD776FF5712330A2F9B5C" box="[218,293,862,884]" italics="true" pageId="49" pageNumber="50">Natella</emphasis>
|
||
)
|
||
<emphasis id="B93BEAF8E03CD776FEBA12330AA39B5C" box="[311,425,862,884]" italics="true" pageId="49" pageNumber="50">viridescens</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0AEE10FBE03CD776FE4E123209819B5C" author="Watson" box="[451,651,862,884]" page="160" pageId="49" pageNumber="50" year="1934">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FE4E1232095F9B5C" author="Watson" box="[451,597,862,884]" pageId="49" pageNumber="59" refString="Watson, H. (1934) Natalina and other South African snails. Proceedings of the Malacological Society of London, 21, 150 - 193." type="journal article" year="1934">Watson, 1934</bibRefCitation>
|
||
: 160
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="0AEE10FBE03CD776FD151233087A9B5C" author="Connolly" box="[664,880,862,884]" page="103" pageId="49" pageNumber="50" year="1939">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FD15123308309B5C" author="Connolly" box="[664,826,862,884]" pageId="49" pageNumber="57" refString="Connolly, M. (1939) A monographic survey of South African non-marine Mollusca. Annals of the South African Museum, 33, 1 - 660." type="journal article" year="1939">Connolly, 1939</bibRefCitation>
|
||
: 103
|
||
</treatmentCitation>
|
||
, text fig. 7 (radula).
|
||
<bibRefCitation id="EFDE4B1BE03CD776FBDC12330FC19B5C" author="Zilch" box="[1105,1227,862,884]" pageId="49" pageNumber="59" refString="Zilch, A. (1959 - 1960) Euthyneura. In: Wenz, W. (Ed.), Gastropoda. In: Schindewolf, O. H. (Ed.), Handbuch der Palaozoologie. Band 6. Teil 2. Gebruder Borntraeger, Berlin, pp. 1 - 835." type="book chapter" year="1959">
|
||
Zilch,
|
||
<date id="FFF1102AE03CD776FB1A12330FC19B5C" box="[1175,1227,862,884]" pageId="49" pageNumber="50" value="1959" valueMax="1960">1959</date>
|
||
</bibRefCitation>
|
||
–60: 554, fig. 1937.
|
||
<treatmentCitation id="0AEE10FBE03CD776FF4A12100AA89BBB" author="Schileyko" box="[199,418,893,915]" page="748" pageId="49" pageNumber="50" year="2000">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FF4A12100A7A9BBB" author="Schileyko" box="[199,368,893,915]" pageId="49" pageNumber="58" refString="Schileyko, A. A. (2000) Treatise on Recent terrestrial pulmonate molluscs. Part 6. Rhytididae, Chlamydephoridae, Systrophiidae, Haplotrematidae, Streptaxidae, Spiraxidae, Oleacinidae, Testacellidae. Ruthenica, 2 (Supplement), 729 - 880." type="journal article" year="2000">Schileyko, 2000</bibRefCitation>
|
||
: 748
|
||
</treatmentCitation>
|
||
, fig. 974.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF1A12F10F6F9B99" blockId="49.[151,1437,739,945]" box="[151,1125,923,945]" pageId="49" pageNumber="50">
|
||
<treatmentCitationGroup id="AB5F11C4E03CD776FF1A12F10F6F9B99" box="[151,1125,923,945]" pageId="49" pageNumber="50">
|
||
<taxonomicName id="4C4F4D69E03CD776FF1A12F10A4A9B99" box="[151,320,923,945]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03CD776FF1A12F10A4A9B99" box="[151,320,923,945]" italics="true" pageId="49" pageNumber="50">Nata viridescens</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<treatmentCitation id="0AEE10FBE03CD776FED612F109129B99" author="Bruggen" box="[347,536,923,945]" page="28" pageId="49" pageNumber="50" year="1967">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FED612F10AF99B99" author="Bruggen" box="[347,499,923,945]" pageId="49" pageNumber="56" refString="Bruggen, A. C. van (1967) Miscellaneous notes on southern African Gastropoda Euthyneura (Mollusca). Zoologische Verhandelingen, 91, 1 - 34." type="journal article" year="1967">Bruggen, 1967</bibRefCitation>
|
||
: 28
|
||
</treatmentCitation>
|
||
, 29.
|
||
<treatmentCitation id="0AEE10FBE03CD776FDC412F6082F9B99" author="Richardson" box="[585,805,923,945]" page="46" pageId="49" pageNumber="50" year="1989">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FDC412F609F59B99" author="Richardson" box="[585,767,923,945]" pageId="49" pageNumber="58" refString="Richardson, C. L. (1989) Streptaxacea: a catalog of species. Part II; Ammonitellidae, Chlamydephoridae, Haplotrematidae, Rhytididae, Systrophiidae. Tryonia, 18, 1 - 154." type="journal article" year="1989">Richardson, 1989</bibRefCitation>
|
||
: 46
|
||
</treatmentCitation>
|
||
.
|
||
<treatmentCitation id="0AEE10FBE03CD776FCA212F60F6B9B99" author="Herbert" box="[815,1121,923,945]" page="219" pageId="49" pageNumber="50" year="2004">
|
||
<bibRefCitation id="EFDE4B1BE03CD776FCA212F60F259B99" author="Herbert" box="[815,1071,923,945]" pageId="49" pageNumber="57" refString="Herbert, D. & Kilburn, D. (2004) Field guide to the land snails and slugs of eastern South Africa. Natal Museum, Pietermaritzburg, 336 pp." type="book" year="2004">Herbert & Kilburn, 2004</bibRefCitation>
|
||
: 219
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03CD776FF1A12B60AEA9C3F" pageId="49" pageNumber="50" type="etymology">
|
||
<paragraph id="8BF036EAE03CD776FF1A12B60AEA9C3F" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF1A12B60A149BDC" bold="true" box="[151,286,987,1012]" pageId="49" pageNumber="50">Etymology.</emphasis>
|
||
From
|
||
<taxonomicName id="4C4F4D69E03CD776FEE512B60AEC9BDC" box="[360,486,987,1012]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03CD776FEE512B60AEC9BDC" box="[360,486,987,1012]" italics="true" pageId="49" pageNumber="50">viridescens</emphasis>
|
||
</taxonomicName>
|
||
(L.)–greenish; presumably in reference to the ‘olivaceous epidermis’ (periostracum) noted by the original authors.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03CD776FF4A154F09329C76" pageId="49" pageNumber="50" type="materials_examined">
|
||
<paragraph id="8BF036EAE03CD776FF4A154F09329C76" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A154F0A7E9C13" bold="true" box="[199,372,1058,1083]" pageId="49" pageNumber="50">
|
||
<typeStatus id="54F48848E03CD776FF4A154F0A089C13" box="[199,258,1058,1083]" pageId="49" pageNumber="50">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54F48848E03CD776FEF6154F0AE99C13" box="[379,483,1058,1083]" pageId="49" pageNumber="50" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4F4D69E03CD776FD85154F0F569C12" authority="Melvill & Ponsonby, 1891" authorityName="Melvill & Ponsonby" authorityYear="1891" box="[520,1116,1058,1083]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="49" pageNumber="50" phylum="Mollusca" rank="species" species="viridescens" subGenus="Patula">
|
||
<emphasis id="B93BEAF8E03CD776FD85154F094F9C13" box="[520,581,1058,1083]" italics="true" pageId="49" pageNumber="50">Helix</emphasis>
|
||
(
|
||
<emphasis id="B93BEAF8E03CD776FDD9154F09959C13" box="[596,671,1058,1083]" italics="true" pageId="49" pageNumber="50">Patula</emphasis>
|
||
)
|
||
<emphasis id="B93BEAF8E03CD776FD22154F08279C13" box="[687,813,1058,1083]" italics="true" pageId="49" pageNumber="50">viridescens</emphasis>
|
||
<bibRefCitation id="EFDE4B1BE03CD776FCB9154F0F569C12" author="Melvill" box="[820,1116,1058,1083]" pageId="49" pageNumber="57" refString="Melvill, J. C. & Ponsonby, J. H. (1891) Descriptions of nine new terrestrial and fluviatile mollusks from South Africa. Annals and Magazine of Natural History, Series 6, 8, 237 - 240. http: // dx. doi. org / 10.1080 / 00222939109460425" type="journal article" year="1891">
|
||
Melvill & Ponsonby,
|
||
<quantity id="4CB79B0FE03CD776FBA9154F0F569C12" box="[1060,1116,1058,1082]" metricMagnitude="1" metricUnit="m" metricValue="4.80314" pageId="49" pageNumber="50" unit="in" value="1891.0">1891</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in
|
||
<collectionCode id="ED5EAE2FE03CD776FBF2154E0FE39C12" LSID="urn:lsid:biocol.org:col:34665" box="[1151,1257,1059,1082]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="49" pageNumber="50">NHMUK</collectionCode>
|
||
(1905.1.26.24), diameter
|
||
<quantity id="4CB79B0FE03CD776FE8D15280A5C9C75" box="[256,342,1093,1118]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="49" pageNumber="50" unit="mm" value="4.5">4.5 mm</quantity>
|
||
(in poor condition).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03CD776FF4A15050F099CA9" box="[199,1027,1128,1153]" pageId="49" pageNumber="50" type="materials_examined">
|
||
<paragraph id="8BF036EAE03CD776FF4A15050F099CA9" blockId="49.[151,1437,987,2033]" box="[199,1027,1128,1153]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A150508489CA9" bold="true" box="[199,834,1128,1153]" pageId="49" pageNumber="50">Additional material examined and literature records:</emphasis>
|
||
See Appendix 6.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03CD776FF4A15E6085A9CEF" pageId="49" pageNumber="50" type="diagnosis">
|
||
<paragraph id="8BF036EAE03CD776FF4A15E6085A9CEF" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A15E60A7A9C8C" bold="true" box="[199,368,1163,1188]" pageId="49" pageNumber="50">Identification.</emphasis>
|
||
Shell characterised by its small size, low-discoidal profile, weak sculpture and wide umbilicus. Distinctive anatomical features are given in generic diagnosis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03CD774FF4A15BF0A7899F8" lastPageId="51" lastPageNumber="52" pageId="49" pageNumber="50" type="description">
|
||
<paragraph id="8BF036EAE03CD776FF4A15BF08959DC9" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A15BF0A779CC3" bold="true" box="[199,381,1234,1259]" pageId="49" pageNumber="50">
|
||
Shell (
|
||
<figureCitation id="13742A6FE03CD776FE9915BF0A649CC3" box="[276,366,1234,1259]" captionStart="FIGURE 37" captionStartId="50.[151,250,1179,1201]" captionTargetBox="[172,1412,198,1151]" captionTargetId="figure@50.[170,1414,193,1165]" captionTargetPageId="50" captionText="FIGURE 37. Natella viridescens (Melvill & Ponsonby, 1891). A – C, Weenen, KwaZulu-Natal, diameter 4.7 mm (NMSA A 7126). D – F, Ithala Game Reserve, KwaZulu-Natal, diameter 5.0 mm (NMSA V 3177). G – I, Pretoria, Gauteng, diameter 3.6 mm (NMSA A 7122)." httpUri="https://zenodo.org/record/266581/files/figure.png" pageId="49" pageNumber="50">Fig. 37</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Discoidal, spire low to almost flat; largest specimens with approx. 2.0 teleoconch whorls; suture above periphery, not noticeably descendant prior to aperture; periphery usually slightly below mid-whorl; surface glossy throughout. Protoconch very small, diameter 0.7–1.0 mm, comprising ±1.0 whorl, smooth and glossy; junction with teleoconch usually weakly defined. Teleoconch initially sculptured only by faint growthlines, but last adult whorl with traces of weak pliculae below suture and with occasional stronger, irregularly spaced growth-lines; sculpture weaker on base. Aperture roundly ovate; peristome interrupted by bulging parietal region; outer lip thin with membranous periostracal fringe when undamaged; base and umbilical margin evenly rounded; umbilicus broad, patent to apex; columella lip not reflected.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF4A14860A3C9E0F" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">Translucent, periostracum more or less uniformly pale golden-yellow to pale olive-yellow; little or no darker axial banding.</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF4A175F0AB89E46" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
Dimensions: Largest specimen, diameter
|
||
<quantity id="4CB79B0FE03CD776FD11175F09FF9E62" box="[668,757,1586,1611]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="49" pageNumber="50" unit="mm" value="5.3">5.3 mm</quantity>
|
||
, height
|
||
<quantity id="4CB79B0FE03CD776FCDC175F08A29E62" box="[849,936,1586,1610]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="49" pageNumber="50" unit="mm" value="2.4">2.4 mm</quantity>
|
||
(Ithala Game Res., NMSA
|
||
<date id="FFF1102AE03CD776FB6A175F0E329E63" box="[1255,1336,1586,1611]" pageId="49" pageNumber="50" value="2530-05">V2530</date>
|
||
); H:D of adults 0.42–0.47 (N=21).
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF4A171508F89F36" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A17150AF99EB9" bold="true" box="[199,499,1656,1681]" pageId="49" pageNumber="50">
|
||
Living animal (
|
||
<figureCitation id="13742A6FE03CD776FE0E17150AC19EB9" box="[387,459,1656,1681]" captionStart="FIGURE 4" captionStartId="9.[151,250,1791,1813]" captionTargetBox="[184,1395,199,1756]" captionTargetId="figure@9.[184,1396,193,1762]" captionTargetPageId="9" captionText="FIGURE 4. Living animals of Nata and Natella. A, Nata aequiplicata sp. nov., holotype, Stinkhoutkloof, E. Cape, shell diameter 6.5 mm (NMSA W 9248 / T 3066). B, Nata dumeticola, West Coast National Park, W. Cape, shell diameter 10.8 mm (NMSA W 3224). C, Nata erugata sp. nov., Tullach-Mohr, Nelspruit area, Mpumalanga, shell diameter 20 mm (MV ex NMSA W 3301). D, Nata tarachodes, Grootbos Private Nat. Res., Gansbaai, W. Cape, shell diameter 7.7 mm (NMSA W 5658). E, Nata watsoni sp. nov., Hankey, E. Cape, shell diameter 7.5 mm (paratype NMSA W 6524 / T 3921). F, Nata watsoni sp. nov. Camdeboo National Park, Graaff-Reinet, E. Cape, shell diameter 13.0 mm (paratype, NMSA W 6091 / T 2303). G, Nata vernicosa, Vernon Crookes Nat. Res., KwaZulu-Natal, shell diameter 14.0 mm (NMSA W 8020). H, Natella viridescens, Ithala Game Res., KwaZulu-Natal, shell diameter 3.7 mm (NMSA W 7263)." httpUri="https://zenodo.org/record/266548/files/figure.png" pageId="49" pageNumber="50">Fig. 4</figureCitation>
|
||
H):
|
||
</emphasis>
|
||
Few specimens examined alive; head and neck grey to dark grey, with scattered microscopic orange pigment granules, tentacles and retractor muscles often somewhat darker; foot paler ventrally, particularly the tail and beneath shell; ventro-lateral margins of foot with more numerous orange pigment granules; skin texture relatively smooth; mantle edge pale, often densely speckled with orange pigment granules; lining of mantle cavity lacking dark makings, whitish kidney sometimes conspicuous.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03CD776FF4A16450E229FD9" blockId="49.[151,1437,987,2033]" pageId="49" pageNumber="50">
|
||
<emphasis id="B93BEAF8E03CD776FF4A16450ACB9F69" bold="true" box="[199,449,1832,1857]" pageId="49" pageNumber="50">
|
||
Radula (
|
||
<figureCitation id="13742A6FE03CD776FEA016450A7E9F69" box="[301,372,1832,1857]" captionStart="FIGURE 6" captionStartId="12.[151,250,1492,1514]" captionTargetBox="[154,1432,339,1464]" captionTargetId="figure@12.[151,1436,338,1471]" captionTargetPageId="12" captionText="FIGURE 6. Radulae of Nata and Natella, automontage light micrographs of half-row portions of stained radula mounts. A, Nata aequiplicata sp. nov., type locality (paratype, NMSA W 9249 / T 3067). B, Nata dumeticola, Jacobsbaai, W. Cape (NMSA V 7760). C, Nata erugata sp. nov., Golden Gate Highlands National Park, Free State (paratype, NMSA W 4243 / T 3913). D, Nata tarachodes, Oubos, W. Cape (NMSA W 5671). E, Nata watsoni sp. nov., Graaff-Reinet, E. Cape (paratype, NMSA W 6091 / T 2303). F, Natella viridescens, Ithala Game Res., KwaZulu-Natal (NMSA V 2530). Scale bars: A = 100 µm; B = 200 µm; C – E = 250 µm; F = 50 µm." httpUri="https://zenodo.org/record/266550/files/figure.png" pageId="49" pageNumber="50">Figs 6</figureCitation>
|
||
F, 38):
|
||
</emphasis>
|
||
Formula 1+7 +(3–4); length up to
|
||
<quantity id="4CB79B0FE03CD776FCC7164408A89F68" box="[842,930,1833,1857]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="49" pageNumber="50" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, with 27–32 broadly V-shaped rows of teeth; rachidian present, narrow with a short, curved cusp; no clear distinction between lateral and marginal series, largest tooth taken to be outermost lateral; inner lateral teeth slender, the cusp down-curved; teeth progressively increasing in length from first to sixth; seventh tooth considerably larger (fang tooth), and much broader and blade-like; 3–4 teeth (marginals) beyond largest lateral, the first also flattened, but somewhat smaller and rather more narrowly acuminate, remaining marginals decreasing rapidly in size, a minute fourth marginal sometimes evident.
|
||
</paragraph>
|
||
<caption id="DF306662E03FD775FF1A15F60A7B9CC6" httpUri="https://zenodo.org/record/266581/files/figure.png" pageId="50" pageNumber="51" targetBox="[172,1412,198,1151]" targetPageId="50">
|
||
<paragraph id="8BF036EAE03FD775FF1A15F60A7B9CC6" blockId="50.[151,1436,1179,1262]" pageId="50" pageNumber="51">
|
||
<emphasis id="B93BEAF8E03FD775FF1A15F60A299C99" bold="true" box="[151,291,1179,1201]" pageId="50" pageNumber="51">FIGURE 37.</emphasis>
|
||
<taxonomicName id="4C4F4D69E03FD775FEA015F608219C99" authority="Melvill & Ponsonby, 1891" authorityName="Melvill & Ponsonby" authorityYear="1891" box="[301,811,1179,1201]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03FD775FEA015F60AFF9C99" box="[301,501,1179,1201]" italics="true" pageId="50" pageNumber="51">Natella viridescens</emphasis>
|
||
(Melvill & Ponsonby, 1891)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93BEAF8E03FD775FCB415F6087A9C99" bold="true" box="[825,880,1179,1201]" pageId="50" pageNumber="51">A–C,</emphasis>
|
||
Weenen, KwaZulu-Natal, diameter 4.7 mm (NMSA A7126).
|
||
<emphasis id="B93BEAF8E03FD775FF7F15D70A2E9CF8" bold="true" box="[242,292,1210,1232]" pageId="50" pageNumber="51">D–F,</emphasis>
|
||
Ithala Game Reserve, KwaZulu-Natal, diameter 5.0 mm (NMSA V3177). G–I, Pretoria, Gauteng, diameter 3.6 mm (NMSA A7122).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF306662E03FD775FF1A16F008149FD9" httpUri="https://zenodo.org/record/266582/files/figure.png" pageId="50" pageNumber="51" targetBox="[190,1414,1314,1927]" targetPageId="50">
|
||
<paragraph id="8BF036EAE03FD775FF1A16F008149FD9" blockId="50.[151,1436,1949,2033]" pageId="50" pageNumber="51">
|
||
<emphasis id="B93BEAF8E03FD775FF1A16F00A299F9B" bold="true" box="[151,291,1949,1971]" pageId="50" pageNumber="51">FIGURE 38.</emphasis>
|
||
<taxonomicName id="4C4F4D69E03FD775FEA116F0082C9F9B" authority="Melvill & Ponsonby, 1891" authorityName="Melvill & Ponsonby" authorityYear="1891" box="[300,806,1949,1971]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="50" pageNumber="51" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03FD775FEA116F00AF89F9B" box="[300,498,1949,1971]" italics="true" pageId="50" pageNumber="51">Natella viridescens</emphasis>
|
||
(Melvill & Ponsonby, 1891)
|
||
</taxonomicName>
|
||
, radula.
|
||
<emphasis id="B93BEAF8E03FD775FC0E16F008919F9B" bold="true" box="[899,923,1949,1971]" pageId="50" pageNumber="51">A,</emphasis>
|
||
whole width of radula, showing small rachidian tooth, stained with Shirlastain A, Ithala Game Reserve, KwaZulu-Natal (NMSA V2530).
|
||
<emphasis id="B93BEAF8E03FD775FBBF16D10F429FFA" bold="true" box="[1074,1096,1980,2002]" pageId="50" pageNumber="51">B,</emphasis>
|
||
SEM of half row, Soutpansberg, Limpopo (NMSA V5664). Scale bars: A = 100 µm; B = 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036EAE03ED774FF4A11FA0A7899F8" blockId="51.[151,1437,151,464]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF4A11FA0AF09898" bold="true" box="[199,506,151,176]" pageId="51" pageNumber="52">
|
||
Distal genitalia (
|
||
<figureCitation id="13742A6FE03ED774FE1C11FA0AE19898" box="[401,491,151,176]" captionStart="FIGURE 39" captionStartId="51.[151,250,1169,1191]" captionTargetBox="[268,1301,560,1138]" captionTargetId="figure@51.[264,1323,547,1147]" captionTargetPageId="51" captionText="FIGURE 39. Natella viridescens (Melvill & Ponsonby, 1891), genitalia (NMSA V 3177). A, middle and lower portion of genital tract. B, internal structure of penis. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursa copulatrix duct; fod, free oviduct; ga, genital atrium; m, muscle fibres; p, penis; pf, folds inside penis base; pr, penial retractor muscle; pv, penial verge; sod, spermoviduct; v, vagina; vd, vas deferens." httpUri="https://zenodo.org/record/266583/files/figure.png" pageId="51" pageNumber="52">Fig. 39</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Genital pore below pneumostome. Penis short and stout (length ±
|
||
<quantity id="4CB79B0FE03ED774FA8911F50E689898" box="[1284,1378,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="51" pageNumber="52" unit="mm" value="0.7">0.7 mm</quantity>
|
||
), not obviously narrowing basally, bluntly rounded at its apex, from which the vas deferens and penial retractor muscle arise. Interior of penis with a broad verge extending for half its length from apex, basal section lined with low longitudinal folds, lacking papillation. No evidence of a swollen epiphallic region prior to insertion of vas deferens; the latter not reflexed and adnate to side of penis, but runs directly backward to its origin at base of spermoviduct. Genital atrium simple and vagina short with thickened wall. Bursa copulatrix and its duct about one-third length of spermoviduct, slender basally, but apical portion slightly swollen; the whole adnate to free oviduct and lower spermoviduct. A concentration of muscle fibres arises close to base of bursa copulatrix duct, connecting with adjacent body wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF306662E03ED774FF1A15FC09BC9D2B" httpUri="https://zenodo.org/record/266583/files/figure.png" pageId="51" pageNumber="52" targetBox="[268,1301,560,1138]" targetPageId="51">
|
||
<paragraph id="8BF036EAE03ED774FF1A15FC09BC9D2B" blockId="51.[151,1436,1169,1283]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF1A15FC0A299C8F" bold="true" box="[151,291,1169,1191]" pageId="51" pageNumber="52">FIGURE 39.</emphasis>
|
||
<taxonomicName id="4C4F4D69E03ED774FEA115FC082D9C8F" authority="Melvill & Ponsonby, 1891" authorityName="Melvill & Ponsonby" authorityYear="1891" box="[300,807,1169,1191]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03ED774FEA115FC0AF99C8F" box="[300,499,1169,1191]" italics="true" pageId="51" pageNumber="52">Natella viridescens</emphasis>
|
||
(Melvill & Ponsonby, 1891)
|
||
</taxonomicName>
|
||
, genitalia (NMSA V3177).
|
||
<emphasis id="B93BEAF8E03ED774FBC215FC0F6D9C8F" bold="true" box="[1103,1127,1169,1191]" pageId="51" pageNumber="52">A,</emphasis>
|
||
middle and lower portion of genital tract.
|
||
<emphasis id="B93BEAF8E03ED774FE9315C20A3F9CED" bold="true" box="[286,309,1199,1221]" pageId="51" pageNumber="52">B,</emphasis>
|
||
internal structure of penis. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursa copulatrix duct; fod, free oviduct; ga, genital atrium; m, muscle fibres; p, penis; pf, folds inside penis base; pr, penial retractor muscle; pv, penial verge; sod, spermoviduct; v, vagina; vd, vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3556561E03ED774FF4A142409E89D81" pageId="51" pageNumber="52" type="distribution">
|
||
<paragraph id="8BF036EAE03ED774FF4A142409E89D81" blockId="51.[151,1437,1353,1882]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF4A14240ADD9D4A" bold="true" box="[199,471,1353,1378]" pageId="51" pageNumber="52">
|
||
Distribution (
|
||
<figureCitation id="13742A6FE03ED774FEE014240AC39D4A" box="[365,457,1353,1378]" captionStart="FIGURE 40" captionStartId="52.[151,250,1199,1221]" captionTargetBox="[180,1412,198,1182]" captionTargetId="figure@52.[179,1413,193,1186]" captionTargetPageId="52" captionText="FIGURE 40. Distribution of Natella viridescens (Melvill & Ponsonby, 1891). Triangles with a white centre indicate localities for specimens used in DNA analyses (Moussalli & Herbert 2016); contour at 1000 m." httpUri="https://zenodo.org/record/266584/files/figure.png" pageId="51" pageNumber="52">Fig. 40</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Endemic to north-eastern
|
||
<collectingCountry id="F358767AE03ED774FC98142408A79D4A" box="[789,941,1353,1378]" name="South Africa" pageId="51" pageNumber="52">South Africa</collectingCountry>
|
||
; from the Soutpansberg (Limpopo), south through N.W. Province, Gauteng and Mpumalanga, to the KwaZulu-Natal Midlands (Howick–Dargle area); mostly at altitudes over
|
||
<quantity id="4CB79B0FE03ED774FEDC14FC0A929D81" box="[337,408,1425,1450]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="51" pageNumber="52" unit="m" value="500.0">500 m</quantity>
|
||
and commonly over
|
||
<quantity id="4CB79B0FE03ED774FD0B14FC09D49D81" box="[646,734,1425,1449]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03ED774FF4A14D809D89E12" pageId="51" pageNumber="52" type="biology_ecology">
|
||
<paragraph id="8BF036EAE03ED774FF4A14D809D89E12" blockId="51.[151,1437,1353,1882]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF4A14D80A209DE6" bold="true" box="[199,298,1461,1486]" pageId="51" pageNumber="52">Habitat.</emphasis>
|
||
Evidently catholic in habitat preferences. Most records from the northern part of the Great Escarpment are from forested habitats, but some of the localities in Gauteng, N.W. Province and KwaZulu-Natal indicate that the species also inhabits somewhat drier, thicket-type vegetation. Primarily in leaf-litter and under logs, but will climb walls and probably tree trunks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556561E03ED774FF4A172809099F71" pageId="51" pageNumber="52" type="discussion">
|
||
<paragraph id="8BF036EAE03ED774FF4A172809499F3A" blockId="51.[151,1437,1353,1882]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF4A17280A319E76" bold="true" box="[199,315,1605,1630]" pageId="51" pageNumber="52">Remarks.</emphasis>
|
||
<taxonomicName id="4C4F4D69E03ED774FECA172809299E76" box="[327,547,1605,1630]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03ED774FECA172809299E76" box="[327,547,1605,1630]" italics="true" pageId="51" pageNumber="52">Natella viridescens</emphasis>
|
||
</taxonomicName>
|
||
is the smallest southern African rhytidid.
|
||
<taxonomicName id="4C4F4D69E03ED774FBAD172B0FB39E75" box="[1056,1209,1606,1629]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="erugata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E03ED774FBAD172B0FB39E75" box="[1056,1209,1606,1629]" italics="true" pageId="51" pageNumber="52">Nata erugata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E03ED774FB4B17280E2F9E76" box="[1222,1317,1605,1630]" pageId="51" pageNumber="52" rank="species">
|
||
<emphasis id="B93BEAF8E03ED774FB4B17280E2F9E76" bold="true" box="[1222,1317,1605,1630]" pageId="51" pageNumber="52">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C4F4D69E03ED774FAE8172B0A0E9EA9" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="vernicosa">
|
||
<emphasis id="B93BEAF8E03ED774FAE8172B0A0E9EA9" italics="true" pageId="51" pageNumber="52">Nata vernicosa</emphasis>
|
||
</taxonomicName>
|
||
, with which it is known to co-occur, are substantially larger species and lack such a wide umbilicus. Juvenile specimens of both have deeper, fewer whorled shells with a much narrower umbilicus. The smaller
|
||
<taxonomicName id="4C4F4D69E03ED774FAE817E30E919E8D" box="[1381,1435,1678,1701]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="51" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93BEAF8E03ED774FAE817E30E919E8D" box="[1381,1435,1678,1701]" italics="true" pageId="51" pageNumber="52">Nata</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="4C4F4D69E03ED774FF7517DF0AA29EE2" box="[248,424,1713,1738]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="aequiplicata" status="sp. nov.">
|
||
<emphasis id="B93BEAF8E03ED774FF7517DF0AA29EE2" box="[248,424,1713,1738]" italics="true" pageId="51" pageNumber="52">N. aequiplicata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2085783E03ED774FE3F17DC09049EE2" box="[434,526,1713,1738]" pageId="51" pageNumber="52" rank="species">
|
||
<emphasis id="B93BEAF8E03ED774FE3F17DC09049EE2" bold="true" box="[434,526,1713,1738]" pageId="51" pageNumber="52">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C4F4D69E03ED774FDC717DF09E09EE2" box="[586,746,1713,1738]" class="Gastropoda" family="Rhytididae" genus="Nata" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pulmonata" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="tarachodes">
|
||
<emphasis id="B93BEAF8E03ED774FDC717DF09E09EE2" box="[586,746,1713,1738]" italics="true" pageId="51" pageNumber="52">N. tarachodes</emphasis>
|
||
</taxonomicName>
|
||
have stronger sculpture and occur only in the southern and western Cape respectively. Specimens of
|
||
<taxonomicName id="4C4F4D69E03ED774FDF817B808459EC6" box="[629,847,1749,1774]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03ED774FDF817B808459EC6" box="[629,847,1749,1774]" italics="true" pageId="51" pageNumber="52">Natella viridescens</emphasis>
|
||
</taxonomicName>
|
||
from northern KwaZulu-Natal are often somewhat larger than those from other localities.
|
||
</paragraph>
|
||
<paragraph id="8BF036EAE03ED774FF4A167009099F71" blockId="51.[151,1437,1353,1882]" pageId="51" pageNumber="52">
|
||
<emphasis id="B93BEAF8E03ED774FF4A16700A659F1E" bold="true" box="[199,367,1821,1846]" pageId="51" pageNumber="52">Conservation.</emphasis>
|
||
With its wide distribution and broad habitat tolerance,
|
||
<taxonomicName id="4C4F4D69E03ED774FC7F16700FC69F1E" box="[1010,1228,1821,1846]" class="Gastropoda" family="Rhytididae" genus="Natella" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="52" phylum="Mollusca" rank="species" species="viridescens">
|
||
<emphasis id="B93BEAF8E03ED774FC7F16700FC69F1E" box="[1010,1228,1821,1846]" italics="true" pageId="51" pageNumber="52">Natella viridescens</emphasis>
|
||
</taxonomicName>
|
||
is not currently a species of conservation concern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |