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<document id="28261DD37E16DB40B79985E9DC2ECEC5" ID-CLB-Dataset="31728" ID-DOI="10.11646/zootaxa.4338.3.7" ID-GBIF-Dataset="a3f92703-00b4-4ef4-81e9-136c538017fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1037042" ID-ZooBank="5FC9941F-D455-48DD-8EDC-BD2278E571F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509002640332" checkinUser="plazi" docAuthor="Cruz-López, Jesús A." docDate="2017" docId="03E687F2947AC337A4CDFE69FB01FE86" docLanguage="en" docName="zootaxa.4338.3.7.pdf" docOrigin="Zootaxa 4338 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Martensolasma catrina Cruz-López 2017, sp. n." docType="treatment" docVersion="10" masterDocId="FFDFFF8A947BC333A45AFFAAFFD4FFC6" masterDocTitle="A second species of the genus Martensolasma (Opiliones, Dyspnoi, Nemastomatidae) from Mexico" updateTime="1702001758674" updateUser="plazi">
<mods:mods id="F6E6A720D560D175AF6E380C38AA990E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F8D247186B70971AAE82A57C71BC97AD">
<mods:title id="7F0CF7EB76A779D2D2D338F525344C36">A second species of the genus Martensolasma (Opiliones, Dyspnoi, Nemastomatidae) from Mexico</mods:title>
</mods:titleInfo>
<mods:name id="12B3210A7B36EBE1EA7EEE8D25BD36F3" type="personal">
<mods:role id="749BDA8D806EE04654521423CA55FD08">
<mods:roleTerm id="810519B74A3BF0BB56E36CC7E5DF7945">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AEA2E64C019741BC9EFFDFDC7457A9BF">Cruz-López, Jesús A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CD583CA885FCD6BA178E650E8E39CEAA">text</mods:typeOfResource>
<mods:relatedItem id="FE810CCD19244F12B6492E6DDE1478F7" type="host">
<mods:titleInfo id="9D9CDACF359AD27829DA2A47A772B9B0">
<mods:title id="F5ACE16127EB49E88F39D67A69771D2C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2CC23B6A7183E052757007744AE89F78">
<mods:date id="C46A85D113782C8E800CB513C87F519D">2017</mods:date>
<mods:detail id="6AED9EFD9ED6544EE9E7180A91C1F9A8" type="pubDate">
<mods:number id="9EB6A5B04F2EEE1C031C62941BAB918C">2017-10-26</mods:number>
</mods:detail>
<mods:detail id="D23B1D3B8378669E8639278E33F82B47" type="volume">
<mods:number id="5436292EA08F33A64A1E133FCADB6AEC">4338</mods:number>
</mods:detail>
<mods:detail id="3D232705C7D52DCB6708F1723C587973" type="issue">
<mods:number id="08398878D3C1FF985EB81C9E9DAC0D6D">3</mods:number>
</mods:detail>
</mods:part>
</mods:relatedItem>
<mods:classification id="4B60B564BB68B78338C25A40F66CE905">journal volume</mods:classification>
<mods:identifier id="36D5E88CD546303BCD66D8AED83FF6EB" type="CLB-Dataset">31728</mods:identifier>
<mods:identifier id="227A6E8670DF310CAA8A7AFE7B67020E" type="DOI">10.11646/zootaxa.4338.3.7</mods:identifier>
<mods:identifier id="74146AA6E5E3DE5417B2DFC7617BDE44" type="GBIF-Dataset">a3f92703-00b4-4ef4-81e9-136c538017fc</mods:identifier>
<mods:identifier id="96DC49C387085057BF0497DB82FDBA43" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9237571ADDBFECEC6AC8D24737C03D92" type="Zenodo-Dep">1037042</mods:identifier>
<mods:identifier id="0A7C7BA733676EC017E85B225D300747" type="ZooBank">5FC9941F-D455-48DD-8EDC-BD2278E571F6</mods:identifier>
</mods:mods>
<treatment id="03E687F2947AC337A4CDFE69FB01FE86" ID-DOI="http://doi.org/10.5281/zenodo.6004762" ID-GBIF-Taxon="135476338" ID-Zenodo-Dep="6004762" LSID="urn:lsid:plazi:treatment:03E687F2947AC337A4CDFE69FB01FE86" httpUri="http://treatment.plazi.org/id/03E687F2947AC337A4CDFE69FB01FE86" lastPageId="4" pageId="1">
<subSubSection id="C355656F947AC332A4CDFE69FED3FE39" pageId="1" type="nomenclature">
<paragraph id="8BF036E4947AC332A4CDFE69FE2DFE18" blockId="1.[151,505,451,511]" box="[151,505,451,478]" pageId="1">
<heading id="D0B88188947AC332A4CDFE69FE2DFE18" bold="true" box="[151,505,451,478]" fontSize="11" level="1" pageId="1" reason="1">
<emphasis id="B93BEAF6947AC332A4CDFE69FE2DFE18" bold="true" box="[151,505,451,478]" pageId="1">
<taxonomicName id="4C4F4D67947AC332A4CDFE69FE7BFE1B" ID-CoL="7X64D" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[151,431,451,477]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF6947AC332A4CDFE69FE7BFE1B" bold="true" box="[151,431,451,477]" italics="true" pageId="1">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D947AC332A5EDFE6EFE2DFE18" box="[439,505,452,478]" pageId="1" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF036E4947AC332A4CDFE4CFED3FE39" blockId="1.[151,505,451,511]" box="[151,263,486,511]" pageId="1">
<figureCitation id="13742A61947AC332A4CDFE4CFED3FE39" box="[151,263,486,511]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5, 6" captionStart-2="FIGURES 7 – 9" captionStart-3="FIGURES 10 – 15" captionStart-4="FIGURES 16, 17" captionStart-5="FIGURES 18 – 24" captionStartId-0="1.[151,264,1981,2004]" captionStartId-1="2.[151,264,776,799]" captionStartId-2="2.[151,264,1961,1984]" captionStartId-3="4.[151,264,1410,1433]" captionStartId-4="4.[151,264,1990,2013]" captionStartId-5="5.[151,264,1962,1985]" captionTargetBox-0="[340,1247,986,1975]" captionTargetBox-1="[340,1247,193,756]" captionTargetBox-2="[340,1247,1455,1939]" captionTargetBox-3="[340,1247,388,1357]" captionTargetBox-4="[340,1247,1615,1956]" captionTargetBox-5="[340,1247,215,1906]" captionTargetId-0="figure@1.[340,1247,986,1975]" captionTargetId-1="figure@2.[340,1247,193,756]" captionTargetId-2="figure@2.[340,1247,1455,1939]" captionTargetId-3="figure@4.[340,1247,388,1357]" captionTargetId-4="figure@4.[340,1247,1615,1956]" captionTargetId-5="figure@5.[340,1247,215,1906]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="FIGURES 1 – 4. Martensolasma catrina sp. n., habitus and prosoma. 1: male paratype, 2: female paratype, 3: male paratype, 4: female paratype. Scale bar 1 and 2: 250 mm, 3 and 4: 150 mm." captionText-1="FIGURES 5, 6. Martensolasma catrina sp. n., dorsal ornamentation. 5: male paratype, middle of scutum, 6: female paratype, detail of ornamentation. Scale bar 5: 150 mm, 6: 250 mm." captionText-2="FIGURES 7 – 9. Martensolasma catrina sp. n., male chelicera. 7: frontal view, 8: detail of mesal and mesoproximal teeth, 9: mesal view. Pink arrows indicate mesal tooth, red arrows indicate mesoproximal tooth. Scale bars 7, 9: 100 mm, 8: 50 mm." captionText-3="FIGURES 10 – 15. Martensolasma catrina sp. n., chelicera and palp. 10: male, dorsal chelicera, 11: female, dorsal chelicera, 12: male, right pedipalp, mesal view, 13: female, right pedipalp, mesal view, 14: male, glandular pores on the swollen area of pedipalpal patella, 15: female, ventral pedipalpal patella. Red circles indicate glandular pore areas, pink arrow indicates mesal tooth, red arrow indicates mesoproximal tooth, red triangles indicate some glandular pores. Scale bar 10 and 11: 50 mm, 12 and 13: 200 mm, 14: 200 mm, 15: 50 mm." captionText-4="FIGURES 16, 17. Martensolasma catrina sp. n., metatarsus II. 16: male, 17: female. Scale bar 16 and 17: 100 mm." captionText-5="FIGURES 18 – 24. Martensolasma catrina sp. n., penis. 18: entire penis, 19 – 23: rotation from dorsal to ventral view of apex, 24: stylus. Red arrow indicates a spermatic pore. Scale bars 18: 200 mm, 19 – 23: 25 mm, 24: 10 mm." httpUri-0="https://zenodo.org/record/1037044/files/figure.png" httpUri-1="https://zenodo.org/record/1037046/files/figure.png" httpUri-2="https://zenodo.org/record/1037048/files/figure.png" httpUri-3="https://zenodo.org/record/1037050/files/figure.png" httpUri-4="https://zenodo.org/record/1037052/files/figure.png" httpUri-5="https://zenodo.org/record/1037054/files/figure.png" pageId="1">Figs 1–24</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C355656F947AC332A4CDFD84FF2DFD3D" pageId="1" type="materials_examined">
<paragraph id="8BF036E4947AC332A4CDFD84FF2DFD3D" blockId="1.[151,1436,558,946]" pageId="1">
<materialsCitation id="3B273CB9947AC332A4CDFD84FE5BFD75" ID-GBIF-Occurrence="1675728551" collectingDate="2017-02-04" collectionCode="CNAN-T" collectorName="J. Cruz &amp; G. Contreras &amp; D. Barrales &amp; L. Olguin &amp; Two &amp; Other &amp; One" country="Mexico" county="Meztitlan" elevation="2" latitude="20.675226" location="Material" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" pageId="1" specimenCode="CNAN-T1109" specimenCount="2" stateProvince="Hidalgo" typeStatus="holotype">
<emphasis id="B93BEAF6947AC332A4CDFD84FE5FFD81" bold="true" box="[151,395,558,583]" pageId="1">
<location id="8E90603F947AC332A4CDFD84FF2BFD81" LSID="urn:lsid:plazi:treatment:03E687F2947AC337A4CDFE69FB01FE86:8E90603F947AC332A4CDFD84FF2BFD81" box="[151,255,558,583]" country="Mexico" county="Meztitlan" latitude="20.675226" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" name="Material" pageId="1" stateProvince="Hidalgo">Material</location>
examined
</emphasis>
.
<location id="8E90603F947AC332A5F6FD84FE32FD81" LSID="urn:lsid:plazi:treatment:03E687F2947AC337A4CDFE69FB01FE86:8E90603F947AC332A5F6FD84FE32FD81" box="[428,486,558,583]" country="Mexico" county="Meztitlan" latitude="20.675226" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" name="Male" pageId="1" stateProvince="Hidalgo">Male</location>
<typeStatus id="54F48846947AC332A5A5FD84FDB5FD81" box="[511,609,558,583]" pageId="1" type="holotype">holotype</typeStatus>
<specimenCode id="DBE99E9F947AC332A620FD85FCCAFD81" box="[634,798,559,583]" collectionCode="CNAN-T" pageId="1">CNAN-T1109</specimenCode>
,
<collectingMunicipality id="6B94AC9E947AC332A761FD84FBCFFD81" box="[827,1051,558,583]" pageId="1">Rancho Manzanal</collectingMunicipality>
,
<collectingCounty id="62914E68947AC332A063FD84FB73FD81" box="[1081,1191,558,583]" pageId="1">Meztitlán</collectingCounty>
,
<collectingRegion id="498BF806947AC332A09FFD84FAF6FD81" box="[1221,1314,558,583]" country="Mexico" name="Hidalgo" pageId="1">Hidalgo</collectingRegion>
,
<collectingCountry id="F3587674947AC332A11AFD84FA4CFD81" box="[1344,1432,558,583]" name="Mexico" pageId="1">Mexico</collectingCountry>
,
<geoCoordinate id="EE7B5023947AC332A4CDFDF9FE8AFDAC" box="[151,350,595,619]" direction="north" orientation="latitude" pageId="1" precision="1" value="20.675226">20º40’30.814’’ N</geoCoordinate>
,
<geoCoordinate id="EE7B5023947AC332A531FDF8FDF9FDAD" box="[363,557,594,619]" direction="west" orientation="longitude" pageId="1" precision="1" value="-98.71013">98º42’36.468’’W</geoCoordinate>
,
<elevation id="0062D1D7947AC332A660FDF8FD4EFDAC" box="[570,666,594,619]" metricMagnitude="0" metricUnit="m" metricValue="2.164" pageId="1" unit="m" value="2.164">
<quantity id="4CB79B01947AC332A660FDF8FD4EFDAC" box="[570,666,594,619]" metricMagnitude="0" metricUnit="m" metricValue="2.164" pageId="1" unit="m" value="2.164">2,164 m</quantity>
</elevation>
,
<collectingDate id="EFB5E9CC947AC332A6FDFDF8FCC8FDAD" box="[679,796,594,619]" pageId="1" value="2017-02-04">
<date id="FFF11024947AC332A6FDFDF8FCC8FDAD" box="[679,796,594,619]" pageId="1" value="2017-02-04">04.ii.2017</date>
</collectingDate>
,
<collectorName id="26BA5332947AC332A773FDF9FCA8FDAD" box="[809,892,595,619]" pageId="1">J. Cruz</collectorName>
,
<collectorName id="26BA5332947AC332A7D3FDF9FBCEFDAD" box="[905,1050,595,619]" pageId="1">G. Contreras</collectorName>
,
<collectorName id="26BA5332947AC332A07DFDF9FB7FFDAD" box="[1063,1195,594,619]" pageId="1">D. Barrales</collectorName>
, and
<collectorName id="26BA5332947AC332A0B3FDF9FA8CFDAD" box="[1257,1368,594,619]" pageId="1">L. Olguín</collectorName>
colls.
<collectorName id="26BA5332947AC332A4CDFDDDFF1CFD48" box="[151,200,631,654]" pageId="1">Two</collectorName>
male and two female
<typeStatus id="54F48846947AC332A590FDDDFDE1FD49" box="[458,565,631,655]" pageId="1" type="paratype">paratypes</typeStatus>
CNAN- T1111, same data as holotype.
<emphasis id="B93BEAF6947AC332A05BFDDCFACAFD49" box="[1025,1310,630,655]" italics="true" pageId="1">
<collectorName id="26BA5332947AC332A05BFDDCFB96FD49" box="[1025,1090,630,655]" pageId="1">Other</collectorName>
material examined
</emphasis>
.
<collectorName id="26BA5332947AC332A174FDDDFA88FD49" box="[1326,1372,631,655]" pageId="1">One</collectorName>
male CNAN-SEM-Op0231
</materialsCitation>
,
<materialsCitation id="3B273CB9947AC332A5C7FD31FE36FD11" ID-GBIF-Occurrence="1675728561" collectingDate="2016-01-20" collectorName="J. Mendoza &amp; Olguin &amp; One" country="Mexico" county="Meztitlan" elevation="2" latitude="20.675226" location="Material" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" pageId="1" specimenCount="1" stateProvince="Hidalgo" typeStatus="holotype">
same locality data,
<collectingDate id="EFB5E9CC947AC332A621FD30FD33FD75" box="[635,743,666,691]" pageId="1" value="2016-01-20">
<date id="FFF11024947AC332A621FD30FD33FD75" box="[635,743,666,691]" pageId="1" value="2016-01-20">20.i.2016</date>
</collectingDate>
, J. Cruz, G. Contreras,
<collectorName id="26BA5332947AC332A7A9FD31FBA2FD75" box="[1011,1142,666,691]" pageId="1">J. Mendoza</collectorName>
and L.
<collectorName id="26BA5332947AC332A088FD30FAF4FD75" box="[1234,1312,666,691]" pageId="1">Olguín</collectorName>
colls.
<collectorName id="26BA5332947AC332A137FD31FA48FD75" box="[1389,1436,667,691]" pageId="1">One</collectorName>
female CNAN-SEM-Op0235
</materialsCitation>
,
<materialsCitation id="3B273CB9947AC332A5B7FD15FBB7FD11" ID-GBIF-Occurrence="1675728567" box="[493,1123,702,727]" collectingDate="2016-01-20" collectorName="J. Mendoza &amp; Olguin &amp; One" country="Mexico" county="Meztitlan" elevation="2" latitude="20.675226" location="Material" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" pageId="1" specimenCount="1" stateProvince="Hidalgo" typeStatus="holotype">same data as holotype. One female CNAN-DNA-Op328</materialsCitation>
,
<materialsCitation id="3B273CB9947AC332A034FD15FF2DFD3D" ID-GBIF-Occurrence="1675728569" collectingDate="2016-01-20" collectorName="J. Mendoza &amp; Olguin &amp; One" country="Mexico" county="Meztitlan" elevation="2" latitude="20.675226" location="Material" longLatPrecision="1" longitude="-98.71013" municipality="Rancho Manzanal" pageId="1" specimenCount="1" stateProvince="Hidalgo" typeStatus="holotype">same data as CNAN-SEM- Op0231.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C355656F947AC332A49DFCACFD1CFC74" pageId="1" type="etymology">
<paragraph id="8BF036E4947AC332A49DFCACFD1CFC74" blockId="1.[151,1436,558,946]" pageId="1">
<emphasis id="B93BEAF6947AC332A49DFCACFE92FCD9" bold="true" box="[199,326,774,799]" pageId="1">Etymology</emphasis>
. “Catrinas” are one of the most emblematic cultural symbols in
<collectingRegion id="498BF806947AC332A04EFCACFBB9FCD9" box="[1044,1133,774,799]" country="Mexico" name="Mexico" pageId="1">
<collectingCountry id="F3587674947AC332A04EFCACFBB9FCD9" box="[1044,1133,774,799]" name="Mexico" pageId="1">Mexico</collectingCountry>
</collectingRegion>
. They represent pictures of elegant women’s skeletons wearing elaborate (upper class) clothing and were originally intended to criticize and protest against political and economic inequality in
<collectingRegion id="498BF806947AC332A6B2FCFBFCEAFCAC" box="[744,830,849,874]" country="Mexico" name="Mexico" pageId="1">
<collectingCountry id="F3587674947AC332A6B2FCFBFCEAFCAC" box="[744,830,849,874]" name="Mexico" pageId="1">Mexico</collectingCountry>
</collectingRegion>
in latter half of the 19th century. Nowadays, they are important characters of one of the most representative holidays in
<collectingRegion id="498BF806947AC332A7C0FCDFFC24FC48" box="[922,1008,885,910]" country="Mexico" name="Mexico" pageId="1">
<collectingCountry id="F3587674947AC332A7C0FCDFFC24FC48" box="[922,1008,885,910]" name="Mexico" pageId="1">Mexico</collectingCountry>
</collectingRegion>
called “Día de Muertos” (day of the Dead). The name is a singular noun in opposition.
</paragraph>
</subSubSection>
<caption id="DF30666C947AC332A4CDF817FCC0F834" httpUri="https://zenodo.org/record/1037044/files/figure.png" pageId="1" targetBox="[340,1247,986,1975]" targetPageId="1">
<paragraph id="8BF036E4947AC332A4CDF817FCC0F834" blockId="1.[151,1436,1981,2034]" pageId="1">
<emphasis id="B93BEAF6947AC332A4CDF817FEEEF815" bold="true" box="[151,314,1981,2004]" pageId="1">FIGURES 1–4.</emphasis>
<taxonomicName id="4C4F4D67947AC332A51AF814FDF9F812" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[320,557,1982,2004]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF6947AC332A51AF814FDF9F812" box="[320,557,1982,2004]" italics="true" pageId="1">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D947AC332A669F814FDB8F815" box="[563,620,1982,2003]" pageId="1" rank="species">
<emphasis id="B93BEAF6947AC332A669F814FDB8F815" bold="true" box="[563,620,1982,2003]" pageId="1">sp. n.</emphasis>
</taxonomicNameLabel>
, habitus and prosoma. 1: male paratype, 2: female paratype, 3: male paratype, 4: female paratype. Scale bar 1 and 2: 250 mm, 3 and 4: 150 mm.
</paragraph>
</caption>
<caption id="DF30666C9479C331A4CDFCA2FD37FCFB" httpUri="https://zenodo.org/record/1037046/files/figure.png" pageId="2" targetBox="[340,1247,193,756]" targetPageId="2">
<paragraph id="8BF036E49479C331A4CDFCA2FD37FCFB" blockId="2.[151,1436,776,829]" pageId="2">
<emphasis id="B93BEAF69479C331A4CDFCA2FEEFFCD8" bold="true" box="[151,315,776,799]" pageId="2">FIGURES 5, 6.</emphasis>
<taxonomicName id="4C4F4D679479C331A518FCA3FDFBFCD9" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[322,559,777,799]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF69479C331A518FCA3FDFBFCD9" box="[322,559,777,799]" italics="true" pageId="2">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D9479C331A66CFCA3FDA4FCD8" box="[566,624,777,798]" pageId="2" rank="species">
<emphasis id="B93BEAF69479C331A66CFCA3FDA4FCD8" bold="true" box="[566,624,777,798]" pageId="2">sp. n.</emphasis>
</taxonomicNameLabel>
, dorsal ornamentation. 5: male paratype, middle of scutum, 6: female paratype, detail of ornamentation. Scale bar 5: 150 mm, 6: 250 mm.
</paragraph>
</caption>
<subSubSection id="C355656F9479C331A49DFCC0FADEFBF1" pageId="2" type="diagnosis">
<paragraph id="8BF036E49479C331A49DFCC0FADEFBF1" blockId="2.[151,1437,874,1403]" pageId="2">
<emphasis id="B93BEAF69479C331A49DFCC0FEEEFC45" bold="true" box="[199,314,874,899]" pageId="2">Diagnosis</emphasis>
.
<taxonomicName id="4C4F4D679479C331A514FCC0FD8FFC44" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[334,603,874,899]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF69479C331A514FCC0FD8FFC44" box="[334,603,874,899]" italics="true" pageId="2">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D9479C331A63DFCC0FD79FC45" box="[615,685,874,899]" pageId="2" rank="species">
<emphasis id="B93BEAF69479C331A63DFCC0FD79FC45" bold="true" box="[615,685,874,899]" pageId="2">sp. n.</emphasis>
</taxonomicNameLabel>
can be differentiated easily from
<taxonomicName id="4C4F4D679479C331A011FCC1FB1DFC45" authorityName="Shear" authorityYear="2006" box="[1099,1225,874,899]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="jocheni">
<emphasis id="B93BEAF69479C331A011FCC1FB1DFC45" box="[1099,1225,874,899]" italics="true" pageId="2">M. jocheni</emphasis>
</taxonomicName>
by the following combination of characters: scutal ornamentation complex—second to fifth transverse rows with five distinctly marked cells, the central three forming a forward-pointing arrowhead, cells generally closed and arranged in a rhombic pattern, whereas in
<taxonomicName id="4C4F4D679479C331A581FC7DFD81FC29" authorityName="Shear" authorityYear="2006" box="[475,597,982,1007]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="jocheni">
<emphasis id="B93BEAF69479C331A581FC7DFD81FC29" box="[475,597,982,1007]" italics="true" pageId="2">M. jocheni</emphasis>
</taxonomicName>
the cells are organized in parallel rows, with most not closed (
<figureCitation id="13742A619479C331A145FC7CFA59FC29" box="[1311,1421,982,1007]" captionStart="FIGURES 5, 6" captionStartId="2.[151,264,776,799]" captionTargetBox="[340,1247,193,756]" captionTargetId="figure@2.[340,1247,193,756]" captionTargetPageId="2" captionText="FIGURES 5, 6. Martensolasma catrina sp. n., dorsal ornamentation. 5: male paratype, middle of scutum, 6: female paratype, detail of ornamentation. Scale bar 5: 150 mm, 6: 250 mm." httpUri="https://zenodo.org/record/1037046/files/figure.png" pageId="2">Figs. 5, 6</figureCitation>
); basal segment of chelicera with small curved mesal tooth, sometimes absent (
<figureCitation id="13742A619479C331A06AFC50FB64FBD5" box="[1072,1200,1018,1043]" captionStart-0="FIGURES 7 – 9" captionStart-1="FIGURES 10 – 15" captionStartId-0="2.[151,264,1961,1984]" captionStartId-1="4.[151,264,1410,1433]" captionTargetBox-0="[340,1247,1455,1939]" captionTargetBox-1="[340,1247,388,1357]" captionTargetId-0="figure@2.[340,1247,1455,1939]" captionTargetId-1="figure@4.[340,1247,388,1357]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 7 – 9. Martensolasma catrina sp. n., male chelicera. 7: frontal view, 8: detail of mesal and mesoproximal teeth, 9: mesal view. Pink arrows indicate mesal tooth, red arrows indicate mesoproximal tooth. Scale bars 7, 9: 100 mm, 8: 50 mm." captionText-1="FIGURES 10 – 15. Martensolasma catrina sp. n., chelicera and palp. 10: male, dorsal chelicera, 11: female, dorsal chelicera, 12: male, right pedipalp, mesal view, 13: female, right pedipalp, mesal view, 14: male, glandular pores on the swollen area of pedipalpal patella, 15: female, ventral pedipalpal patella. Red circles indicate glandular pore areas, pink arrow indicates mesal tooth, red arrow indicates mesoproximal tooth, red triangles indicate some glandular pores. Scale bar 10 and 11: 50 mm, 12 and 13: 200 mm, 14: 200 mm, 15: 50 mm." httpUri-0="https://zenodo.org/record/1037048/files/figure.png" httpUri-1="https://zenodo.org/record/1037050/files/figure.png" pageId="2">Figs. 7–10</figureCitation>
); metatarsus II with subapical pseudoarticulation; three articles in tarsus II; penial stylus hook-shaped at tip (
<figureCitation id="13742A619479C331A02FFBB4FB2BFBF1" box="[1141,1279,1054,1079]" captionStart="FIGURES 18 – 24" captionStartId="5.[151,264,1962,1985]" captionTargetBox="[340,1247,215,1906]" captionTargetId="figure@5.[340,1247,215,1906]" captionTargetPageId="5" captionText="FIGURES 18 – 24. Martensolasma catrina sp. n., penis. 18: entire penis, 19 – 23: rotation from dorsal to ventral view of apex, 24: stylus. Red arrow indicates a spermatic pore. Scale bars 18: 200 mm, 19 – 23: 25 mm, 24: 10 mm." httpUri="https://zenodo.org/record/1037054/files/figure.png" pageId="2">Figs. 19–24</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C355656F9479C330A49DFBE8FE2CFAAA" lastPageId="3" pageId="2" type="description">
<paragraph id="8BF036E49479C331A49DFBE8FB9FFABD" blockId="2.[151,1437,874,1403]" pageId="2">
<emphasis id="B93BEAF69479C331A49DFBE8FE85FB9D" bold="true" box="[199,337,1090,1115]" pageId="2">Description</emphasis>
.
<emphasis id="B93BEAF69479C331A53BFBE8FDD0FB9D" box="[353,516,1090,1115]" italics="true" pageId="2">
Male
<typeStatus id="54F488469479C331A5F9FBE8FDD0FB9D" box="[419,516,1090,1115]" pageId="2" type="holotype">holotYpe</typeStatus>
</emphasis>
: Scute length = 1.80 mm. Anterolateral processes of scutum short, with rounded apices. Dorsal cells defined as follows: Two large cells lateral to eye tubercle, cell posterior to eye tubercles incomplete, continuous with the median cell of the first transverse row. First row with five cells, the two lateroexternal pairs small, with median longitudinal keel not well defined; central cell larger. Second row with five cells, latero-external cells large, rectangular; three median cells subequal in size, with longitudinal keels incomplete, three median cells arranged in a W-shaped array. Third to fifth rows with five cells, each row with latero-external pairs slightly larger than the median three; the median three cells forming an arrowhead, with central cell pentagonal, displaced anteriorly. Sixth row with five subequal cells, central cell triangular, with acute anterior apex; a small cell is between the central and the first lateral-right cells (
<figureCitation id="13742A619479C331A7ECFAC8FBF5FABD" box="[950,1057,1378,1403]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,264,1981,2004]" captionTargetBox="[340,1247,986,1975]" captionTargetId="figure@1.[340,1247,986,1975]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Martensolasma catrina sp. n., habitus and prosoma. 1: male paratype, 2: female paratype, 3: male paratype, 4: female paratype. Scale bar 1 and 2: 250 mm, 3 and 4: 150 mm." httpUri="https://zenodo.org/record/1037044/files/figure.png" pageId="2">Figs. 1, 3</figureCitation>
,
<figureCitation id="13742A619479C331A077FAC8FBE9FABD" box="[1069,1085,1378,1403]" captionStart="FIGURES 5, 6" captionStartId="2.[151,264,776,799]" captionTargetBox="[340,1247,193,756]" captionTargetId="figure@2.[340,1247,193,756]" captionTargetPageId="2" captionText="FIGURES 5, 6. Martensolasma catrina sp. n., dorsal ornamentation. 5: male paratype, middle of scutum, 6: female paratype, detail of ornamentation. Scale bar 5: 150 mm, 6: 250 mm." httpUri="https://zenodo.org/record/1037046/files/figure.png" pageId="2">5</figureCitation>
).
</paragraph>
<caption id="DF30666C9479C331A4CDF803FAA2F818" httpUri="https://zenodo.org/record/1037048/files/figure.png" pageId="2" targetBox="[340,1247,1455,1939]" targetPageId="2">
<paragraph id="8BF036E49479C331A4CDF803FAA2F818" blockId="2.[151,1436,1961,2014]" pageId="2">
<emphasis id="B93BEAF69479C331A4CDF803FEE8F878" bold="true" box="[151,316,1961,1984]" pageId="2">FIGURES 7–9.</emphasis>
<taxonomicName id="4C4F4D679479C331A519F803FDE5F879" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[323,561,1961,1983]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF69479C331A519F803FDE5F879" box="[323,561,1961,1983]" italics="true" pageId="2">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D9479C331A663F800FDA0F879" box="[569,628,1962,1983]" pageId="2" rank="species">
<emphasis id="B93BEAF69479C331A663F800FDA0F879" bold="true" box="[569,628,1962,1983]" pageId="2">sp. n.</emphasis>
</taxonomicNameLabel>
, male chelicera. 7: frontal view, 8: detail of mesal and mesoproximal teeth, 9: mesal view. Pink arrows indicate mesal tooth, red arrows indicate mesoproximal tooth. Scale bars 7, 9: 100 mm, 8: 50 mm.
</paragraph>
</caption>
<paragraph id="8BF036E49478C330A49DFF3DFB7DFF12" blockId="3.[151,1436,151,608]" pageId="3">
Chelicera small, with basal segment smooth, humped dorsally, with a small curved mesal tooth. Second segment with small, curved mesoproximal tooth, without evidence of pore field (
<figureCitation id="13742A619478C330A078FF16FB49FF12" box="[1058,1181,188,213]" captionStart-0="FIGURES 7 – 9" captionStart-1="FIGURES 10 – 15" captionStartId-0="2.[151,264,1961,1984]" captionStartId-1="4.[151,264,1410,1433]" captionTargetBox-0="[340,1247,1455,1939]" captionTargetBox-1="[340,1247,388,1357]" captionTargetId-0="figure@2.[340,1247,1455,1939]" captionTargetId-1="figure@4.[340,1247,388,1357]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 7 – 9. Martensolasma catrina sp. n., male chelicera. 7: frontal view, 8: detail of mesal and mesoproximal teeth, 9: mesal view. Pink arrows indicate mesal tooth, red arrows indicate mesoproximal tooth. Scale bars 7, 9: 100 mm, 8: 50 mm." captionText-1="FIGURES 10 – 15. Martensolasma catrina sp. n., chelicera and palp. 10: male, dorsal chelicera, 11: female, dorsal chelicera, 12: male, right pedipalp, mesal view, 13: female, right pedipalp, mesal view, 14: male, glandular pores on the swollen area of pedipalpal patella, 15: female, ventral pedipalpal patella. Red circles indicate glandular pore areas, pink arrow indicates mesal tooth, red arrow indicates mesoproximal tooth, red triangles indicate some glandular pores. Scale bar 10 and 11: 50 mm, 12 and 13: 200 mm, 14: 200 mm, 15: 50 mm." httpUri-0="https://zenodo.org/record/1037048/files/figure.png" httpUri-1="https://zenodo.org/record/1037050/files/figure.png" pageId="3">Figs. 7–10</figureCitation>
).
</paragraph>
<paragraph id="8BF036E49478C330A49DFF75FEE1FE4E" blockId="3.[151,1436,151,608]" pageId="3">
Pedipalpal segments thin. Femur with a few scattered glandular setae, distributed along entire segment. Patella swollen meso-apically, covered with few scattered glandular setae, with dense aggregation of small glandular pores on swollen surface. Tibia covered with many glandular setae, swollen meso-basally; small glandular pores on translucent swollen surface. Tarsus rounded apically, covered with many glandular setae and a few ordinary setae (
<figureCitation id="13742A619478C330A4C5FEC5FEF3FE4E" box="[159,295,367,392]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1410,1433]" captionTargetBox="[340,1247,388,1357]" captionTargetId="figure@4.[340,1247,388,1357]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Martensolasma catrina sp. n., chelicera and palp. 10: male, dorsal chelicera, 11: female, dorsal chelicera, 12: male, right pedipalp, mesal view, 13: female, right pedipalp, mesal view, 14: male, glandular pores on the swollen area of pedipalpal patella, 15: female, ventral pedipalpal patella. Red circles indicate glandular pore areas, pink arrow indicates mesal tooth, red arrow indicates mesoproximal tooth, red triangles indicate some glandular pores. Scale bar 10 and 11: 50 mm, 12 and 13: 200 mm, 14: 200 mm, 15: 50 mm." httpUri="https://zenodo.org/record/1037050/files/figure.png" pageId="3">Figs. 12, 14</figureCitation>
).
</paragraph>
<paragraph id="8BF036E49478C330A49DFE3FFD24FDA6" blockId="3.[151,1436,151,608]" pageId="3">
Leg formula: 2, 4, 3, 1 (measurements in table 1). Femora cylindrical in cross section; middle of femora I and III slightly swollen, femur IV with basal pseudoarticulation. Patellae globular. Tibiae shorter than femora, slightly swollen. Metatarsi long and slender, metatarsus II with subapical pseudoarticulation. Metatarsi covered densely with short trichomes and longer fine setae, more prominent in metatarsus II (
<figureCitation id="13742A619478C330A04DFE55FBB9FDDE" box="[1047,1133,511,536]" captionStart="FIGURES 16, 17" captionStartId="4.[151,264,1990,2013]" captionTargetBox="[340,1247,1615,1956]" captionTargetId="figure@4.[340,1247,1615,1956]" captionTargetPageId="4" captionText="FIGURES 16, 17. Martensolasma catrina sp. n., metatarsus II. 16: male, 17: female. Scale bar 16 and 17: 100 mm." httpUri="https://zenodo.org/record/1037052/files/figure.png" pageId="3">Fig. 16</figureCitation>
). Tarsi I and II with three articles, the third with a basal pseudoarticulation, giving appearance of four articles. Tarsi III and IV with four articles, the third and fourth with pseudoarticulations.
</paragraph>
<caption id="DF30666C9478C330A4CDFD3AFABCFD61" box="[151,1384,655,679]" pageId="3">
<paragraph id="8BF036E49478C330A4CDFD3AFABCFD61" blockId="3.[151,1384,655,679]" box="[151,1384,655,679]" pageId="3">
<emphasis id="B93BEAF69478C330A4CDFD3AFEDEFD61" bold="true" box="[151,266,656,679]" pageId="3">TABLE 1.</emphasis>
Pedipalp and legs measurements of holotype of
<taxonomicName id="4C4F4D679478C330A74AFD25FBDDFD61" authority="Cruz-López, 2017" authorityName="Cruz-López" authorityYear="2017" box="[784,1033,655,679]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="catrina" status="sp. nov.">
<emphasis id="B93BEAF69478C330A74AFD25FBDDFD61" box="[784,1033,655,679]" italics="true" pageId="3">Martensolasma catrina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208578D9478C330A04AFD3AFB98FD61" box="[1040,1100,656,679]" pageId="3" rank="species">
<emphasis id="B93BEAF69478C330A04AFD3AFB98FD61" bold="true" box="[1040,1100,656,679]" pageId="3">sp. n.</emphasis>
</taxonomicNameLabel>
Measurements are in mm.
</paragraph>
</caption>
<paragraph id="8BF036E49478C330A5EAFD69FADEFC61" pageId="3">
<table id="F94FC44494783CCCA4C5FD68FAF4FC61" box="[159,1312,706,935]" gridcols="6" gridrows="6" pageId="3">
<tr id="357F34A694783CCCA4C5FD68FAF4FD1E" box="[159,1312,706,728]" gridrow="0" pageId="3" rowspan-0="1">
<th id="76AE5DDA94783CCCA5EAFD68FE26FD1E" box="[432,498,706,728]" gridcol="1" gridrow="0" pageId="3">Femur</th>
<th id="76AE5DDA94783CCCA623FD68FD6AFD1E" box="[633,702,706,728]" gridcol="2" gridrow="0" pageId="3">Patella</th>
<th id="76AE5DDA94783CCCA719FD68FCA3FD1E" box="[835,887,706,728]" gridcol="3" gridrow="0" pageId="3">Tibia</th>
<th id="76AE5DDA94783CCCA051FD68FBAEFD1E" box="[1035,1146,706,728]" gridcol="4" gridrow="0" pageId="3">Metatarsus</th>
<th id="76AE5DDA94783CCCA084FD68FAF4FD1E" box="[1246,1312,706,728]" gridcol="5" gridrow="0" pageId="3">Tarsus</th>
</tr>
<tr id="357F34A694783CCCA4C5FD46FAF4FCC4" box="[159,1312,748,770]" gridrow="1" pageId="3">
<th id="76AE5DDA94783CCCA4C5FD46FF2CFCC4" box="[159,248,748,770]" gridcol="0" gridrow="1" pageId="3">Pedipalp</th>
<td id="76AE5DDA94783CCCA5EAFD46FE26FCC4" box="[432,498,748,770]" gridcol="1" gridrow="1" pageId="3">0.66</td>
<td id="76AE5DDA94783CCCA623FD46FD6AFCC4" box="[633,702,748,770]" gridcol="2" gridrow="1" pageId="3">0.60</td>
<td id="76AE5DDA94783CCCA719FD46FCA3FCC4" box="[835,887,748,770]" gridcol="3" gridrow="1" pageId="3">0.43</td>
<td id="76AE5DDA94783CCCA051FD46FBAEFCC4" box="[1035,1146,748,770]" gridcol="4" gridrow="1" pageId="3">-</td>
<td id="76AE5DDA94783CCCA084FD46FAF4FCC4" box="[1246,1312,748,770]" gridcol="5" gridrow="1" pageId="3">0.26</td>
</tr>
<tr id="357F34A694783CCCA4C5FCBFFAF4FCED" box="[159,1312,789,811]" gridrow="2" pageId="3">
<th id="76AE5DDA94783CCCA4C5FCBFFF2CFCED" box="[159,248,789,811]" gridcol="0" gridrow="2" pageId="3">LI</th>
<td id="76AE5DDA94783CCCA5EAFCBFFE26FCED" box="[432,498,789,811]" gridcol="1" gridrow="2" pageId="3">0.66</td>
<td id="76AE5DDA94783CCCA623FCBFFD6AFCED" box="[633,702,789,811]" gridcol="2" gridrow="2" pageId="3">0.33</td>
<td id="76AE5DDA94783CCCA719FCBFFCA3FCED" box="[835,887,789,811]" gridcol="3" gridrow="2" pageId="3">0.53</td>
<td id="76AE5DDA94783CCCA051FCBFFBAEFCED" box="[1035,1146,789,811]" gridcol="4" gridrow="2" pageId="3">0.90</td>
<td id="76AE5DDA94783CCCA084FCBFFAF4FCED" box="[1246,1312,789,811]" gridcol="5" gridrow="2" pageId="3">0.73</td>
</tr>
<tr id="357F34A694783CCCA4C5FC94FAF4FC92" box="[159,1312,830,852]" gridrow="3" pageId="3">
<th id="76AE5DDA94783CCCA4C5FC94FF2CFC92" box="[159,248,830,852]" gridcol="0" gridrow="3" pageId="3">LII</th>
<td id="76AE5DDA94783CCCA5EAFC94FE26FC92" box="[432,498,830,852]" gridcol="1" gridrow="3" pageId="3">1.06</td>
<td id="76AE5DDA94783CCCA623FC94FD6AFC92" box="[633,702,830,852]" gridcol="2" gridrow="3" pageId="3">0.36</td>
<td id="76AE5DDA94783CCCA719FC94FCA3FC92" box="[835,887,830,852]" gridcol="3" gridrow="3" pageId="3">0.66</td>
<td id="76AE5DDA94783CCCA051FC94FBAEFC92" box="[1035,1146,830,852]" gridcol="4" gridrow="3" pageId="3">2.06</td>
<td id="76AE5DDA94783CCCA084FC94FAF4FC92" box="[1246,1312,830,852]" gridcol="5" gridrow="3" pageId="3">1.03</td>
</tr>
<tr id="357F34A694783CCCA4C5FCC2FAF4FCB8" box="[159,1312,872,894]" gridrow="4" pageId="3">
<th id="76AE5DDA94783CCCA4C5FCC2FF2CFCB8" box="[159,248,872,894]" gridcol="0" gridrow="4" pageId="3">LIII</th>
<td id="76AE5DDA94783CCCA5EAFCC2FE26FCB8" box="[432,498,872,894]" gridcol="1" gridrow="4" pageId="3">0.80</td>
<td id="76AE5DDA94783CCCA623FCC2FD6AFCB8" box="[633,702,872,894]" gridcol="2" gridrow="4" pageId="3">0.33</td>
<td id="76AE5DDA94783CCCA719FCC2FCA3FCB8" box="[835,887,872,894]" gridcol="3" gridrow="4" pageId="3">0.53</td>
<td id="76AE5DDA94783CCCA051FCC2FBAEFCB8" box="[1035,1146,872,894]" gridcol="4" gridrow="4" pageId="3">0.86</td>
<td id="76AE5DDA94783CCCA084FCC2FAF4FCB8" box="[1246,1312,872,894]" gridcol="5" gridrow="4" pageId="3">0.83</td>
</tr>
<tr id="357F34A694783CCCA4C5FC3BFAF4FC61" box="[159,1312,913,935]" gridrow="5" pageId="3">
<th id="76AE5DDA94783CCCA4C5FC3BFF2CFC61" box="[159,248,913,935]" gridcol="0" gridrow="5" pageId="3">LIV</th>
<td id="76AE5DDA94783CCCA5EAFC3BFE26FC61" box="[432,498,913,935]" gridcol="1" gridrow="5" pageId="3">1.10</td>
<td id="76AE5DDA94783CCCA623FC3BFD6AFC61" box="[633,702,913,935]" gridcol="2" gridrow="5" pageId="3">0.36</td>
<td id="76AE5DDA94783CCCA719FC3BFCA3FC61" box="[835,887,913,935]" gridcol="3" gridrow="5" pageId="3">0.70</td>
<td id="76AE5DDA94783CCCA051FC3BFBAEFC61" box="[1035,1146,913,935]" gridcol="4" gridrow="5" pageId="3">1.10</td>
<td id="76AE5DDA94783CCCA084FC3BFAF4FC61" box="[1246,1312,913,935]" gridcol="5" gridrow="5" pageId="3">1.06</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF036E49478C330A49DFC46FDE2FBEE" blockId="3.[151,1437,1004,2037]" pageId="3">
Penis long and thin. Stylus narrow, tip noticeably hook-shaped, apex rounded. Monomorphic setation, no more than seven small setae (
<figureCitation id="13742A619478C330A5FAFBA5FDFEFBEE" box="[416,554,1039,1064]" captionStart="FIGURES 18 – 24" captionStartId="5.[151,264,1962,1985]" captionTargetBox="[340,1247,215,1906]" captionTargetId="figure@5.[340,1247,215,1906]" captionTargetPageId="5" captionText="FIGURES 18 – 24. Martensolasma catrina sp. n., penis. 18: entire penis, 19 – 23: rotation from dorsal to ventral view of apex, 24: stylus. Red arrow indicates a spermatic pore. Scale bars 18: 200 mm, 19 – 23: 25 mm, 24: 10 mm." httpUri="https://zenodo.org/record/1037054/files/figure.png" pageId="3">Figs. 18–24</figureCitation>
).
</paragraph>
<paragraph id="8BF036E49478C330A49DFB9EFCA9FB7E" blockId="3.[151,1437,1004,2037]" pageId="3">
<emphasis id="B93BEAF69478C330A49DFB9EFE5EFB8A" box="[199,394,1076,1101]" italics="true" pageId="3">
Female
<typeStatus id="54F488469478C330A57DFB9FFE5EFB8A" box="[295,394,1077,1100]" pageId="3" type="paratype">paratYpe</typeStatus>
</emphasis>
: Scute length = 1.96. Similar to the male, but with the following differences: Sulci on mesotergum slightly developed, tending toward
<emphasis id="B93BEAF69478C330A688FBF3FC50FBB6" box="[722,900,1113,1136]" italics="true" pageId="3">scutum parvum</emphasis>
; median scutal cells of second to fifth rows complex, irregular in shape; second segment of chelicera without mesoproximal tooth; pedipalpal patella and tibia not swollen, without glandular pores (
<figureCitation id="13742A619478C330A61AFB35FD7FFB7E" box="[576,683,1183,1208]" captionStart="FIGURES 1 – 4" captionStartId="1.[151,264,1981,2004]" captionTargetBox="[340,1247,986,1975]" captionTargetId="figure@1.[340,1247,986,1975]" captionTargetPageId="1" captionText="FIGURES 1 – 4. Martensolasma catrina sp. n., habitus and prosoma. 1: male paratype, 2: female paratype, 3: male paratype, 4: female paratype. Scale bar 1 and 2: 250 mm, 3 and 4: 150 mm." httpUri="https://zenodo.org/record/1037044/files/figure.png" pageId="3">Figs. 2, 4</figureCitation>
,
<figureCitation id="13742A619478C330A6EDFB35FD13FB7E" box="[695,711,1183,1208]" captionStart="FIGURES 5, 6" captionStartId="2.[151,264,776,799]" captionTargetBox="[340,1247,193,756]" captionTargetId="figure@2.[340,1247,193,756]" captionTargetPageId="2" captionText="FIGURES 5, 6. Martensolasma catrina sp. n., dorsal ornamentation. 5: male paratype, middle of scutum, 6: female paratype, detail of ornamentation. Scale bar 5: 150 mm, 6: 250 mm." httpUri="https://zenodo.org/record/1037046/files/figure.png" pageId="3">6</figureCitation>
,
<figureCitation id="13742A619478C330A689FB0AFC90FB7E" box="[723,836,1183,1208]" captionStart="FIGURES 10 – 15" captionStartId="4.[151,264,1410,1433]" captionTargetBox="[340,1247,388,1357]" captionTargetId="figure@4.[340,1247,388,1357]" captionTargetPageId="4" captionText="FIGURES 10 – 15. Martensolasma catrina sp. n., chelicera and palp. 10: male, dorsal chelicera, 11: female, dorsal chelicera, 12: male, right pedipalp, mesal view, 13: female, right pedipalp, mesal view, 14: male, glandular pores on the swollen area of pedipalpal patella, 15: female, ventral pedipalpal patella. Red circles indicate glandular pore areas, pink arrow indicates mesal tooth, red arrow indicates mesoproximal tooth, red triangles indicate some glandular pores. Scale bar 10 and 11: 50 mm, 12 and 13: 200 mm, 14: 200 mm, 15: 50 mm." httpUri="https://zenodo.org/record/1037050/files/figure.png" pageId="3">11, 13, 15</figureCitation>
,
<figureCitation id="13742A619478C330A70AFB0AFCBBFB7E" box="[848,879,1184,1208]" captionStart="FIGURES 16, 17" captionStartId="4.[151,264,1990,2013]" captionTargetBox="[340,1247,1615,1956]" captionTargetId="figure@4.[340,1247,1615,1956]" captionTargetPageId="4" captionText="FIGURES 16, 17. Martensolasma catrina sp. n., metatarsus II. 16: male, 17: female. Scale bar 16 and 17: 100 mm." httpUri="https://zenodo.org/record/1037052/files/figure.png" pageId="3">17</figureCitation>
).
</paragraph>
<paragraph id="8BF036E49478C330A49DFB6EFE2CFAAA" blockId="3.[151,1437,1004,2037]" pageId="3">
<emphasis id="B93BEAF69478C330A49DFB6EFEA9FB1B" bold="true" box="[199,381,1220,1245]" pageId="3">Natural history</emphasis>
. All specimens were found under small rocks in the boundary area between a cattle ranch and a pine forest. The specimens were found in sympatry with an undescribed species of
<taxonomicName id="4C4F4D679478C330A0C9FB4DFE81FAE2" authority="Goodnight &amp; Goodnight, 1942" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A0C9FB4DFB23FAC6" box="[1171,1271,1255,1280]" italics="true" pageId="3">Trilasma</emphasis>
Goodnight &amp; Goodnight, 1942
</taxonomicName>
and the stygnopsid
<taxonomicName id="4C4F4D679478C330A662FAA6FB1EFAE3" authority="Goodnight &amp; Goodnight, 1946" authorityName="Goodnight &amp; Goodnight" authorityYear="1946" box="[568,1226,1292,1317]" class="Arachnida" family="Stygnopsidae" genus="Chapulobunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="unispinosus">
<emphasis id="B93BEAF69478C330A662FAA6FCB3FAE2" box="[568,871,1292,1317]" italics="true" pageId="3">Chapulobunus unispinosus</emphasis>
Goodnight &amp; Goodnight, 1946
</taxonomicName>
and undetermined species of the genera
<taxonomicName id="4C4F4D679478C330A5CAFA9AFC80FA8E" authority="Cruz-Lopez &amp; Francke, 2015" authorityName="Cruz-Lopez &amp; Francke" authorityYear="2015" box="[400,852,1327,1352]" class="Arachnida" family="Stygnopsidae" genus="Crettaros" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A5CAFA9AFE2DFA81" box="[400,505,1328,1351]" italics="true" pageId="3">Crettaros</emphasis>
Cruz-López &amp; Francke, 2015
</taxonomicName>
,
<taxonomicName id="4C4F4D679478C330A73BFA9BFAC4FA8E" authority="Goodnight &amp; Goodnight, 1944" authorityName="Goodnight &amp; Goodnight" authorityYear="1944" box="[865,1296,1327,1352]" class="Magnoliopsida" family="Apiaceae" genus="Karos" higherTaxonomySource="CoL" kingdom="Plantae" order="Apiales" pageId="3" phylum="Tracheophyta" rank="genus">
<emphasis id="B93BEAF69478C330A73BFA9BFC71FA8E" box="[865,933,1329,1352]" italics="true" pageId="3">Karos</emphasis>
Goodnight &amp; Goodnight, 1944
</taxonomicName>
, and
<taxonomicName id="4C4F4D679478C330A114FA9BFE27FAAA" authority="Goodnight &amp; Goodnight, 1947" authorityName="Goodnight &amp; Goodnight" authorityYear="1947" class="Insecta" family="Pyralidae" genus="Potosa" kingdom="Animalia" order="Lepidoptera" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A114FA9BFA48FA8E" box="[1358,1436,1329,1352]" italics="true" pageId="3">Potosa</emphasis>
Goodnight &amp; Goodnight, 1947
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C355656F9478C337A49DFADDFB01FE86" lastPageId="4" pageId="3" type="discussion">
<paragraph id="8BF036E49478C330A49DFADDFB1BF86B" blockId="3.[151,1437,1004,2037]" pageId="3">
<emphasis id="B93BEAF69478C330A49DFADDFE91FA56" bold="true" box="[199,325,1399,1424]" pageId="3">Discussion</emphasis>
.
<bibRefCitation id="EFDE4B159478C330A507FADDFE2EFA56" author="Shear" box="[349,506,1399,1424]" pageId="3" refString="Shear, W. A. (2006) Martensolasma jocheni, a new genus and species of harvestman from Mexico (Opiliones: Nemastomatidae: Ortholasmatinae). Zootaxa, 1325, 191 - 198." type="journal article" year="2006">Shear (2006)</bibRefCitation>
discussed the phylogenetic affinities of
<taxonomicName id="4C4F4D679478C330A7A6FADDFB7EFA56" authorityName="Shear" authorityYear="2006" box="[1020,1194,1399,1424]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A7A6FADDFB7EFA56" box="[1020,1194,1399,1424]" italics="true" pageId="3">Martensolasma</emphasis>
</taxonomicName>
and the remaining
<taxonomicName id="4C4F4D679478C330A4CDFA36FE87FA73" authorityName="Shear &amp; Gruber" authorityYear="1983" box="[151,339,1436,1461]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
, and suggested that the unusual absence of the modified eye process, the dorsal ornamentation without complex cells, and the paucity or absence of pseudoarticulations on the femora and metatarsi could represent plesiomorphic characters within the subfamily.
<bibRefCitation id="EFDE4B159478C330A745FA4EFC7FFA3B" author="Shear" box="[799,939,1508,1533]" pageId="3" refString="Shear, W. A. (2006) Martensolasma jocheni, a new genus and species of harvestman from Mexico (Opiliones: Nemastomatidae: Ortholasmatinae). Zootaxa, 1325, 191 - 198." type="journal article" year="2006">Shear (2006</bibRefCitation>
,
<bibRefCitation id="EFDE4B159478C330A7E2FA4EFC26FA3A" author="Shear" box="[952,1010,1508,1532]" pageId="3" refString="Shear, W. A. (2010) New species and records of ortholasmatine harvestmen from Mexico, Honduras, and the western United States (Opiliones, Nemastomatidae, Ortholasmatinae). Zookeys, 52, 9 - 45. https: // doi. org / 10.3897 / zookeys. 52.471" type="journal article" year="2010">2010</bibRefCitation>
) considered
<taxonomicName id="4C4F4D679478C330A0D9FA4EFAE5FA3B" authorityName="Shear" authorityYear="2006" box="[1155,1329,1508,1533]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A0D9FA4EFAE5FA3B" box="[1155,1329,1508,1533]" italics="true" pageId="3">Martensolasma</emphasis>
</taxonomicName>
to be the most divergent member of
<taxonomicName id="4C4F4D679478C330A5B3F9ADFD72F9E6" authorityName="Shear &amp; Gruber" authorityYear="1983" box="[489,678,1543,1568]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
. With the recent discovery of
<taxonomicName id="4C4F4D679478C330A073F9A3FB7CF9E6" authorityName="Cruz-López" authorityYear="2017" box="[1065,1192,1545,1568]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="catrina">
<emphasis id="B93BEAF69478C330A073F9A3FB7CF9E6" box="[1065,1192,1545,1568]" italics="true" pageId="3">M. catrina</emphasis>
</taxonomicName>
described here, the morphological features of the genus are more heterogeneous, with those of the new species being shared with other members of the subfamily. According to the morphological-phylogenetic hypothesis of
<bibRefCitation id="EFDE4B159478C330A033F9E5FABAF9AE" author="Shear" box="[1129,1390,1615,1640]" pageId="3" refString="Shear, W. A. &amp; Gruber, J. (1983) The opilionid subfamily Ortholasmatinae (Opiliones, Troguloidea, Nemastomatidae). American Museum Novitates, 2757, 1 - 65." type="journal article" year="1983">Shear &amp; Gruber (1983)</bibRefCitation>
, the dorsal ornamentation with well-marked cells and hooked stylus are synapomorphies for the clade
<taxonomicName id="4C4F4D679478C330A0A7F9DEFA57F94B" authorityName="Banks" authorityYear="1894" box="[1277,1411,1652,1677]" class="Arachnida" family="Nemastomatidae" genus="Ortholasma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A0A7F9DEFA57F94B" box="[1277,1411,1652,1677]" italics="true" pageId="3">Ortholasma</emphasis>
</taxonomicName>
+
<taxonomicName id="4C4F4D679478C330A4CDF93DFF2EF976" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[151,250,1687,1712]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A4CDF93DFF2EF976" box="[151,250,1687,1712]" italics="true" pageId="3">Trilasma</emphasis>
</taxonomicName>
. However, these characters are also present in
<emphasis id="B93BEAF69478C330A772F933FC7DF976" box="[808,937,1689,1712]" italics="true" pageId="3">
<taxonomicName id="4C4F4D679478C330A772F933FC72F976" authorityName="Cruz-López" authorityYear="2017" box="[808,934,1689,1712]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="catrina">M. catrina</taxonomicName>
,
</emphasis>
although with the dorsal cells not as well defined as in
<taxonomicName id="4C4F4D679478C330A576F916FE5BF913" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[300,399,1724,1749]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A576F916FE5BF913" box="[300,399,1724,1749]" italics="true" pageId="3">Trilasma</emphasis>
</taxonomicName>
. Thus,
<taxonomicName id="4C4F4D679478C330A5BEF917FD8EF912" authorityName="Cruz-López" authorityYear="2017" box="[484,602,1725,1748]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="catrina">
<emphasis id="B93BEAF69478C330A5BEF917FD8EF912" box="[484,602,1725,1748]" italics="true" pageId="3">M. catrina</emphasis>
</taxonomicName>
presents an intermediate condition between
<taxonomicName id="4C4F4D679478C330A017F916FB2FF913" authorityName="Shear" authorityYear="2006" box="[1101,1275,1724,1749]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A017F916FB2FF913" box="[1101,1275,1724,1749]" italics="true" pageId="3">Martensolasma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D679478C330A16BF916FA47F913" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[1329,1427,1724,1749]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A16BF916FA47F913" box="[1329,1427,1724,1749]" italics="true" pageId="3">Trilasma</emphasis>
</taxonomicName>
. This observation opens the question of whether
<taxonomicName id="4C4F4D679478C330A6E0F975FCBCF93E" authorityName="Shear" authorityYear="2006" box="[698,872,1759,1784]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A6E0F975FCBCF93E" box="[698,872,1759,1784]" italics="true" pageId="3">Martensolasma</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C4F4D679478C330A014F975FB00F93E" authorityName="Banks" authorityYear="1894" box="[1102,1236,1759,1784]" class="Arachnida" family="Nemastomatidae" genus="Ortholasma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A014F975FB00F93E" box="[1102,1236,1759,1784]" italics="true" pageId="3">Ortholasma</emphasis>
</taxonomicName>
+
<taxonomicName id="4C4F4D679478C330A0AFF975FA8DF93E" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[1269,1369,1759,1784]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A0AFF975FA8DF93E" box="[1269,1369,1759,1784]" italics="true" pageId="3">Trilasma</emphasis>
</taxonomicName>
clade based on genital synapomorphies (and with a reversal in
<taxonomicName id="4C4F4D679478C330A767F8AFFC6DF8DB" authorityName="Shear" authorityYear="2006" box="[829,953,1796,1821]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="jocheni">
<emphasis id="B93BEAF69478C330A767F8AFFC6DF8DB" box="[829,953,1796,1821]" italics="true" pageId="3">M. jocheni</emphasis>
</taxonomicName>
), or whether the hooked stylus appeared independently in
<taxonomicName id="4C4F4D679478C330A53EF88DFE3EF886" authorityName="Banks" authorityYear="1894" box="[356,490,1831,1856]" class="Arachnida" family="Nemastomatidae" genus="Ortholasma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A53EF88DFE3EF886" box="[356,490,1831,1856]" italics="true" pageId="3">Ortholasma</emphasis>
</taxonomicName>
+
<taxonomicName id="4C4F4D679478C330A649F88DFDA3F886" authorityName="Goodnight &amp; Goodnight" authorityYear="1942" box="[531,631,1831,1856]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A649F88DFDA3F886" box="[531,631,1831,1856]" italics="true" pageId="3">Trilasma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4F4D679478C330A6E3F883FCE1F886" authorityName="Cruz-López" authorityYear="2017" box="[697,821,1833,1856]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="catrina">
<emphasis id="B93BEAF69478C330A6E3F883FCE1F886" box="[697,821,1833,1856]" italics="true" pageId="3">M. catrina</emphasis>
</taxonomicName>
. However, it is possible that the stylus of the
<typeStatus id="54F488469478C330A136F883FA48F887" box="[1388,1436,1833,1857]" pageId="3">type</typeStatus>
specimen of
<taxonomicName id="4C4F4D679478C330A57FF8E7FE48F8A3" authorityName="Shear" authorityYear="2006" box="[293,412,1868,1893]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="jocheni">
<emphasis id="B93BEAF69478C330A57FF8E7FE48F8A3" box="[293,412,1868,1893]" italics="true" pageId="3">M. jocheni</emphasis>
</taxonomicName>
depicted by
<bibRefCitation id="EFDE4B159478C330A670F8E6FCC6F8A2" author="Shear" box="[554,786,1868,1893]" pageId="3" refString="Shear, W. A. (2006) Martensolasma jocheni, a new genus and species of harvestman from Mexico (Opiliones: Nemastomatidae: Ortholasmatinae). Zootaxa, 1325, 191 - 198." type="journal article" year="2006">Shear (2006: fig. 14)</bibRefCitation>
was broken and that the hooked terminal portion was lost. If so, this would indicate that the hooked stylus is ancestral for
<taxonomicName id="4C4F4D679478C330A73BF8C5FBDBF84E" authorityName="Shear" authorityYear="2006" box="[865,1039,1903,1928]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF69478C330A73BF8C5FBDBF84E" box="[865,1039,1903,1928]" italics="true" pageId="3">Martensolasma</emphasis>
</taxonomicName>
and that the “straight stylus” of
<taxonomicName id="4C4F4D679478C330A127F8DBFF3EF86B" authorityName="Shear" authorityYear="2006" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" phylum="Arthropoda" rank="species" species="jocheni">
<emphasis id="B93BEAF69478C330A127F8DBFF3EF86B" italics="true" pageId="3">M. jocheni</emphasis>
</taxonomicName>
is an artifact. Indeed, the specimens of this species available to me all have hooked styli.
</paragraph>
<paragraph id="8BF036E49478C337A48BF812FB01FE86" blockId="3.[151,1437,1004,2037]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" pageId="3">
Morphological variation among ortholasmatines is noted only in the number of pseudoarticulations on the metatarsi and number of tarsomeres of all legs. Dorsal ornamentation, shape and ornamentation of the ocular process, and the number of lateral processes are constant among species. On the other hand, male genital morphology is conserved, being very similar and apparently without variation among the genera (
<bibRefCitation id="EFDE4B15947FC337A0BCFF16FF06FF3E" author="Shear" pageId="4" refString="Shear, W. A. &amp; Gruber, J. (1983) The opilionid subfamily Ortholasmatinae (Opiliones, Troguloidea, Nemastomatidae). American Museum Novitates, 2757, 1 - 65." type="journal article" year="1983">Shear &amp; Gruber 1983</bibRefCitation>
, 2010). The presence of straight and hooked styli and large and mixed dorsal cells in the same genus, opens the possibility that morphology has evolved in a different way than has been proposed. A future phylogeny of
<taxonomicName id="4C4F4D67947FC337A4CDFE8DFE84FE86" authorityName="Shear &amp; Gruber" authorityYear="1983" box="[151,336,295,320]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Opiliones" pageId="4" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
might illuminate the evolutionary morphology of taxonomic traits in this group.
</paragraph>
</subSubSection>
</treatment>
</document>
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