298 lines
37 KiB
XML
298 lines
37 KiB
XML
<document id="6A4B257B463D06D20EBEDA0ADFBAAC5D" ID-DOI="10.11646/zootaxa.3709.1.2" ID-GBIF-Dataset="64d5b399-d948-4189-8af8-75f5ffc6b62d" ID-ISSN="1175-5326" ID-Zenodo-Dep="222914" ID-ZooBank="A610056E-AE10-483A-B3E6-7D0C2AA168A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460673260853" checkinUser="plazi" docAuthor="Guéorguiev, Borislav" docDate="2013" docId="03E187FBFFEAFF97FF34FF149E97FF02" docLanguage="en" docName="zt03709p070.pdf" docOrigin="Zootaxa 3709 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrotus indicus Guéorguiev, 2013, sp. n." docType="treatment" docUuid="1A486DF2-0384-4FBC-815C-C8E0CFFB9914" docUuidSource="ZooBank" docVersion="10" masterDocId="FFD8FF83FFE0FF9BFFA3FF839C02FFD7" masterDocTitle="Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae)" updateTime="1698348865835" updateUser="plazi">
|
||
<mods:mods id="4F19C435006B378C5D7656AB46AA08E7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6CDEAC0DA881B8E383870E3703E81C1C">
|
||
<mods:title id="8406D737A21AC0BC7ECCD192EB1B5557">Taxonomic, nomenclatural, and faunistic records for species in tribes Melaenini, Moriomorphini, Pterostichini, Licinini, and Sphodrini (Coleoptera: Carabidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="5ED21BEE9733445BFDD56E4C69F1BAD0" type="personal">
|
||
<mods:role id="114B73231B9A7E22AA7168C05CC9685D">
|
||
<mods:roleTerm id="ECFF164B8AB6366069F226F4939DF4AA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3A6A8462F003C737922D52F05984DC97">Guéorguiev, Borislav</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="5646B562D0B2FD112291A6607B8591BD">text</mods:typeOfResource>
|
||
<mods:relatedItem id="A52DA2C83B390624E8EAE4C27D82AE50" type="host">
|
||
<mods:titleInfo id="EC4AD2DE97CE9CEF1991847DE6771DD6">
|
||
<mods:title id="01FEC3E953187AC34B376A8ACF5CBF79">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="75B0EB6FA28FE8ADF68027A32A5023DA">
|
||
<mods:date id="C2457D31504DF66D9213591889F88DB6">2013</mods:date>
|
||
<mods:detail id="DADC5497B25E784123EB2AB48B2A49C1" type="volume">
|
||
<mods:number id="65D12AA3AFC26DA3A8F87A6D654AC781">3709</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C639B9A5FBAEDC5D0469C9AC8FF99A1F" type="issue">
|
||
<mods:number id="283A8F527DC47C82402BFB70DEFFAD60">1</mods:number>
|
||
</mods:detail>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="210514E9344DCB2AC870D9AAEF868849">journal volume</mods:classification>
|
||
<mods:identifier id="1750C491BBB63B10F4537F8544126C2C" type="DOI">10.11646/zootaxa.3709.1.2</mods:identifier>
|
||
<mods:identifier id="95999A1A0CCB43619DE9116EF6D37C03" type="GBIF-Dataset">64d5b399-d948-4189-8af8-75f5ffc6b62d</mods:identifier>
|
||
<mods:identifier id="DB898CCDF17AFE61C44E9F2B7C37DD72" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="804146F02618E2B8328AA74489DBF1CB" type="Zenodo-Dep">222914</mods:identifier>
|
||
<mods:identifier id="BB745F281B75E388EE517682D8395AE4" type="ZooBank">A610056E-AE10-483A-B3E6-7D0C2AA168A1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E187FBFFEAFF97FF34FF149E97FF02" ID-DOI="http://doi.org/10.5281/zenodo.6150466" ID-GBIF-Taxon="119577703" ID-Zenodo-Dep="6150466" LSID="urn:lsid:zoobank.org:act:1A486DF2-0384-4FBC-815C-C8E0CFFB9914" httpUri="http://treatment.plazi.org/id/03E187FBFFEAFF97FF34FF149E97FF02" lastPageId="12" pageId="10">
|
||
<subSubSection id="C3526566FFEAFF91FF34FF149F3FFF07" pageId="10" type="nomenclature">
|
||
<paragraph id="8BF736EDFFEAFF91FF34FF149D9BFF65" blockId="10.[151,829,151,208]" box="[151,409,151,178]" pageId="10">
|
||
<heading id="D0BF8181FFEAFF91FF34FF149D9BFF65" bold="true" box="[151,409,151,178]" fontSize="11" level="1" pageId="10" reason="1">
|
||
<emphasis id="B93CEAFFFFEAFF91FF34FF149D9BFF65" bold="true" box="[151,409,151,178]" pageId="10">
|
||
<taxonomicName id="4C484D6EFFEAFF91FF34FF149D52FF66" ID-CoL="JM94" box="[151,336,151,177]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B93CEAFFFFEAFF91FF34FF149D52FF66" bold="true" box="[151,336,151,177]" italics="true" pageId="10">Atrotus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5784FFEAFF91FEF4FF1B9D9BFF65" box="[343,409,152,178]" pageId="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF736EDFFEAFF91FF34FF399F3FFF07" blockId="10.[151,829,151,208]" box="[151,829,186,208]" pageId="10">
|
||
Habitus:
|
||
<figureCitation id="13732A68FFEAFF91FF50FF399D3FFF07" box="[243,317,186,208]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="10">Fig. 16</figureCitation>
|
||
; mouth parts:
|
||
<figureCitation id="13732A68FFEAFF91FE6EFF399E1AFF07" box="[461,536,186,208]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="10">Fig. 17</figureCitation>
|
||
; male genitalia:
|
||
<figureCitation id="13732A68FFEAFF91FD1DFF399F38FF07" box="[702,826,186,208]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="10">Figs. 18–20</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526566FFEAFF91FF34FF7E9EC8FE56" pageId="10" type="materials_examined">
|
||
<paragraph id="8BF736EDFFEAFF91FF34FF7E9EC8FE56" blockId="10.[151,1436,253,530]" pageId="10">
|
||
<emphasis id="B93CEAFFFFEAFF91FF34FF7E9D3FFEC1" bold="true" box="[151,317,253,278]" pageId="10">
|
||
<typeStatus id="54F3884FFFEAFF91FF34FF7E9CD0FEC1" box="[151,210,253,278]" pageId="10">Type</typeStatus>
|
||
material
|
||
</emphasis>
|
||
.
|
||
<typeStatus id="54F3884FFFEAFF91FEE8FF7E9DB1FEC1" box="[331,435,253,278]" pageId="10" type="holotype">Holotype</typeStatus>
|
||
♂, left fore leg missing, mounted on card, genitalia dissected and deposited in euparal on separate triangular card pinned below the first card, “
|
||
<collectingCountry id="F35F767DFFEAFF91FCB3FEA29F57FEEE" box="[784,853,289,313]" name="India" pageId="10">INDIA</collectingCountry>
|
||
: Tamil Nadu Nilgiri Hills Coonoor,
|
||
<quantity id="4CB09B08FFEAFF91FAAFFEA2996AFEEE" box="[1292,1384,289,313]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="10" unit="m" value="1700.0">1700 m</quantity>
|
||
, 3.–
|
||
<date id="FFF6102DFFEAFF91FF34FEC69D05FE89" box="[151,263,325,350]" pageId="10" value="1991-05-10">5.10.1991</date>
|
||
leg. R.
|
||
<collectionCode id="ED59AE28FFEAFF91FEFEFEC59DAEFE89" box="[349,428,326,350]" pageId="10">SCHUH</collectionCode>
|
||
” [black print on white label] / “H O L O T Y P E
|
||
<taxonomicName id="4C484D6EFFEAFF91FC7AFEC59880FE8A" box="[985,1154,325,349]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B93CEAFFFFEAFF91FC7AFEC59880FE8A" box="[985,1154,325,349]" italics="true" pageId="10">Atrotus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5784FFEAFF91FB2AFEC598DDFE8A" box="[1161,1247,326,349]" pageId="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Guéorguiev des. 2012” [black print on red label] [
|
||
<figureCitation id="13732A68FFEAFF91FDA4FEEA9E58FE55" box="[519,602,361,386]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="10">Fig. 16</figureCitation>
|
||
] (
|
||
<collectionCode id="ED59AE28FFEAFF91FDD3FEEA9EBCFE56" box="[624,702,361,385]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="10">NMW</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526566FFEAFF91FF64FE0E9F2EFDC6" pageId="10" type="diagnosis">
|
||
<paragraph id="8BF736EDFFEAFF91FF64FE0E9F2EFDC6" blockId="10.[151,1436,253,530]" pageId="10">
|
||
<emphasis id="B93CEAFFFFEAFF91FF64FE0E9D38FE71" bold="true" box="[199,314,397,422]" pageId="10">Diagnosis</emphasis>
|
||
. This species is easily diagnosed from all congeners by the following set of characters: 1/ size of body
|
||
<quantity id="4CB09B08FFEAFF91FF7BFE329D31FE1E" box="[216,307,433,458]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="10" unit="mm" value="7.5">7.5 mm</quantity>
|
||
; 2/ microsculpture distinct on the whole dorsal surface; 3/ terminal segment of the maxillary palpi attenuate; 4/ pronotum widest around the middle, with plane lateral margins; 5/ elytral striae impunctate; 6/ parascutellar stria absent; 7/ median lobe with disc at apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF376665FFEAFF91FF34F9959EDDF9BD" httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="10" targetBox="[173,1405,589,1527]" targetPageId="10">
|
||
<paragraph id="8BF736EDFFEAFF91FF34F9959EDDF9BD" blockId="10.[151,1436,1558,1642]" pageId="10">
|
||
<emphasis id="B93CEAFFFFEAFF91FF34F9959D4DF9FB" bold="true" box="[151,335,1558,1580]" pageId="10">FIGURES 16–20</emphasis>
|
||
. Holotype of
|
||
<taxonomicName id="4C484D6EFFEAFF91FE43F9949E78F9FB" box="[480,634,1558,1580]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B93CEAFFFFEAFF91FE43F9949E78F9FB" box="[480,634,1558,1580]" italics="true" pageId="10">Atrotus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5784FFEAFF91FD21F9949EBEF9FB" box="[642,700,1559,1580]" pageId="10" rank="species">sp. n.</taxonomicNameLabel>
|
||
: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18
|
||
<emphasis id="B93CEAFFFFEAFF91FD09F9D79EB4F9BD" bold="true" box="[682,694,1620,1642]" pageId="10">–</emphasis>
|
||
20).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3526566FFEAFF90FF64F9159F61F843" lastPageId="11" pageId="10" type="description">
|
||
<paragraph id="8BF736EDFFEAFF90FF64F9159DA7FBAB" blockId="10.[151,1436,1686,2036]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" pageId="10">
|
||
<emphasis id="B93CEAFFFFEAFF91FF64F9159D53F978" bold="true" box="[199,337,1686,1711]" pageId="10">Description</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEAFF91FEC3F9149DBBF978" box="[352,441,1687,1711]" italics="true" pageId="10">Habitus</emphasis>
|
||
. Moderately large-sized species of
|
||
<taxonomicName id="4C484D6EFFEAFF91FCEEF9149F9CF979" box="[845,926,1687,1710]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CEAFFFFEAFF91FCEEF9149F9CF979" box="[845,926,1687,1710]" italics="true" pageId="10">Atrotus</emphasis>
|
||
</taxonomicName>
|
||
, with elongate and flat body.
|
||
<emphasis id="B93CEAFFFFEAFF91FB50F91B9997F978" box="[1267,1429,1688,1711]" italics="true" pageId="10">Measurements</emphasis>
|
||
. BL:
|
||
<quantity id="4CB09B08FFEAFF91FF68F9399D27F905" box="[203,293,1722,1747]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="10" unit="mm" value="7.5">7.5 mm</quantity>
|
||
; BW:
|
||
<quantity id="4CB09B08FFEAFF91FECEF9399DD4F905" box="[365,470,1722,1747]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" pageId="10" unit="mm" value="2.95">2.95 mm</quantity>
|
||
.
|
||
<emphasis id="B93CEAFFFFEAFF91FE42F9389E2BF905" box="[481,553,1723,1746]" italics="true" pageId="10">Ratios</emphasis>
|
||
. PW/HW: 1.73; PW/PL: 1.29; PbW/PaW: 1.36; EW/PW: 1.31; EL/EW: 1.54.
|
||
<emphasis id="B93CEAFFFFEAFF91FF34F95C9CD5F920" box="[151,215,1759,1783]" italics="true" pageId="10">Color</emphasis>
|
||
. Head piceous, with base of clypeus, mandibles and most of terminal segments of maxillary and labial palpi testaceus, apex of clypeus, labrum, palpi, and antennae yellowish; pronotum piceous on disc and apex, sides and base narrowly testaceus; elytra dark brown to piceous on disc and base, apex and fields outside stria 8 testaceus to yellowish, scutellum and suture somewhat darker than disc; ventral side mostly dark brown to piceous, mouthparts testaceous, elytral epipleura and entire legs yellowish.
|
||
<emphasis id="B93CEAFFFFEAFF91FCA0F8EC9FACF850" box="[771,942,1903,1927]" italics="true" pageId="10">Microsculpture</emphasis>
|
||
. Distinct on entire dorsal surface, head and pronotum with isodiametric, elytra with slight transverse microsculpture.
|
||
<emphasis id="B93CEAFFFFEAFF91FC6DF8109816F87D" box="[974,1044,1939,1962]" italics="true" pageId="10">Lustre</emphasis>
|
||
. Dorsal and ventral surfaces shiny.
|
||
<emphasis id="B93CEAFFFFEAFF91FF34F8349CD6F818" box="[151,212,1975,1999]" italics="true" pageId="10">Head</emphasis>
|
||
. Narrower with respect to pronotum, frons smooth, frontal furrows superficial, short, converging behind, two supraorbital setae on each side; eyes fairly large, convex, moderately prominent, tempora short; clypeus incompletely fused with labrum, with anterior margin almost straight and a seta on each side, same incompletely fused with frons as clypeal suture is perceptible; labrum fairly bilobed, with V-shaped incision in middle, incision with two short setae medially, lobes inflated, each with a large seta at apex; mandibles long, curved downwards at apex, incisor tooth and terebral tooth sharp-pointed (angle at tip <90º) in both mandibles, as incisor teeth longer, more curved and sharper than terebral teeth (
|
||
<figureCitation id="13732A68FFEBFF90FD11FEA49F08FE97" box="[690,778,295,320]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="11">Fig. 17</figureCitation>
|
||
); apical segments of maxillary palpi attenuate, sparsely pubescent; apical segment of labial palpi somewhat fusiform, sparsely pubescent; antennae very long, basal antennomere almost as long as two succeeding segments, antennomere 7 crosses posterior margin of pronotum.
|
||
<emphasis id="B93CEAFFFFEBFF90FF34FE169D04FE7B" box="[151,262,405,428]" italics="true" pageId="11">Pronotum</emphasis>
|
||
. Wide, sub-quadrate, widest at middle, with lateral margins distinctly, narrowly bordered (bordering reduced only before posterolateral setae), flat; sides more convex apically than basally, with two pairs of setae situated inward at discal position (not at margin), lateral setae situated in pronotal apical second fifth, removed from border at a distance larger than diameter of pore, posterolateral setae situated in pronotal basal sixth before hind angles, at distance from border equal to diameter of pore; anterior margin emarginated, shorter than basal margin, fore angles round, moderately prominent; posterior margin convex, hind angles broadly rounded, without perceptible tips; basal impressions rather faint, sub-linear, slanting, convergent posteriorly; disc flat, smooth (only basal part with several faint and short longitudinal wrinkles), median line short, fine, faintly impressed.
|
||
<emphasis id="B93CEAFFFFEBFF90FAECFD139997FD7F" box="[1359,1429,656,680]" italics="true" pageId="11">Elytra</emphasis>
|
||
. Sub-elongate, widest just after middle, coalesced along suture; sides with very slight subapical sinuation; humeri well-marked, obtusely angulate; striae impunctate, moderately impressed, parascutellar striae absent, all striae disappeared close to base; intervals flat, smooth; basal margin complete, almost reaching stria 1; parascutellar setiferous puncture present, situated in stria 2; interval 3 with a small discal setiferous puncture adjoining stria 2 at basal second fifth of elytra; hind wings vestigial.
|
||
<emphasis id="B93CEAFFFFEBFF90FD61FCC79F6EFC8B" box="[706,876,836,860]" italics="true" pageId="11">Ventral surface</emphasis>
|
||
(
|
||
<emphasis id="B93CEAFFFFEBFF90FCDEFCC79864FC8B" box="[893,1126,836,860]" italics="true" pageId="11">thorax and abdomen</emphasis>
|
||
). Sternal part of thorax and abdomen smooth and shiny; metepisterna very wide, shorter than long, weakly narrowed behind, anterior margin appreciably longer than each of other three margins; last visible sternum with two minute marginal setiferous punctures.
|
||
<emphasis id="B93CEAFFFFEBFF90FEB2FC329D44FC1F" box="[273,326,945,968]" italics="true" pageId="11">Legs</emphasis>
|
||
. Long, slender; basal segments 1–3 of male protarsi dilated and asymmetrical, first one longer than wide, longer than each of two succeeding segments, segments 2–3 nearly as long as wide.
|
||
<emphasis id="B93CEAFFFFEBFF90FB30FC57993AFC3B" box="[1171,1336,980,1004]" italics="true" pageId="11">Male genitalia</emphasis>
|
||
. Median lobe of aedeagus moderately long, curved down viewed laterally (
|
||
<figureCitation id="13732A68FFEBFF90FCDEFC749FD2FBC7" box="[893,976,1015,1040]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="11">Fig. 18</figureCitation>
|
||
), basal and medial parts uniformly wide, basal orifice rather concave, apical part thinner towards hooked tip (dorsal hook larger, ventral hook minute); right paramere long and narrow, dilated at tip with a minute seta (
|
||
<figureCitation id="13732A68FFEBFF90FC99FBBC9F8EFB8F" box="[826,908,1087,1112]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="11">Fig. 19</figureCitation>
|
||
), left paramere conchoid, with a minute seta in distal position (
|
||
<figureCitation id="13732A68FFEBFF90FEE7FBE79D9AFBAB" box="[324,408,1124,1149]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="11">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF736EDFFEBFF90FF64FB049EBAFADA" blockId="11.[151,1437,151,2013]" pageId="11">
|
||
<emphasis id="B93CEAFFFFEBFF90FF64FB049D37FB77" bold="true" box="[199,309,1159,1184]" pageId="11">Affinities</emphasis>
|
||
. The morphologic features of the new species conform rather well to the extended diagnosis of
|
||
<taxonomicName id="4C484D6EFFEBFF90FF34FB2F9DC5FB12" authority="Basilewsky 1951" authorityName="Basilewsky" authorityYear="1951" box="[151,455,1196,1221]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93CEAFFFFEBFF90FF34FB2F9CEBFB14" box="[151,233,1196,1219]" italics="true" pageId="11">Atrotus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD94B1CFFEBFF90FF54FB2F9DBCFB12" author="Basilewsky" box="[247,446,1196,1221]" pageId="11" refString="Basilewsky, P. (1951) Sur le genre Atrotus Peringuey (Col. Carabidae) et quelques considerations sur le peuplement entomologique des hautes montagnes de l'Afrique. Annales de la Societe royale de Zoologie du Belgique, 82, 5 - 18." type="journal article" year="1951">Basilewsky 1951</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. However, there are two features with different states in the new taxon, compared with those in the other congeners: 1/ terminal segment of the maxillary palpi attenuate (not fusiform); 2/ complete reduction of the parascutellar striae of the elytra.
|
||
</paragraph>
|
||
<paragraph id="8BF736EDFFEBFF90FF64FA949802F9A3" blockId="11.[151,1437,151,2013]" pageId="11">
|
||
Based on
|
||
<bibRefCitation id="EFD94B1CFFEBFF90FE9BFA949DD9FAE7" author="Britton" box="[312,475,1303,1328]" pageId="11" refString="Britton, E. B. (1948) Coleoptera: Cicindelidae and Carabidae. In: Expedition to south-west Arabia 1937 - 8. British Museum (Natural History), 1, 87 - 131." type="journal article" year="1948">Britton (1948)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFD94B1CFFEBFF90FDB7FA949EE9FAE7" author="Basilewsky" box="[532,747,1303,1328]" pageId="11" refString="Basilewsky, P. (1951) Sur le genre Atrotus Peringuey (Col. Carabidae) et quelques considerations sur le peuplement entomologique des hautes montagnes de l'Afrique. Annales de la Societe royale de Zoologie du Belgique, 82, 5 - 18." type="journal article" year="1951">Basilewsky (1951)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C484D6EFFEBFF90FD5BFA9B9F68FAE7" box="[760,874,1304,1328]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B93CEAFFFFEBFF90FD5BFA9B9F0BFAF8" box="[760,777,1304,1327]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FCBAFA9B9F68FAE7" box="[793,874,1304,1328]" italics="true" pageId="11">indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5784FFEBFF90FCD1FA9B9FB2FAF8" box="[882,944,1304,1327]" pageId="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
appears close to two species, one from the Arabian Peninsula and another from
|
||
<collectingCountry id="F35F767DFFEBFF90FD9AFABF9ECCFA82" box="[569,718,1340,1365]" name="South Africa" pageId="11">South Africa</collectingCountry>
|
||
. The new species resembles
|
||
<taxonomicName id="4C484D6EFFEBFF90FBB7FABF995FFA83" authority="Britton, 1948" authorityName="Britton" authorityYear="1948" box="[1044,1373,1340,1365]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FBB7FABF98BDFA83" box="[1044,1215,1340,1364]" italics="true" pageId="11">Atrotus bicolor</emphasis>
|
||
<bibRefCitation id="EFD94B1CFFEBFF90FB64FABF995FFA83" author="Britton" box="[1223,1373,1340,1365]" pageId="11" refString="Britton, E. B. (1948) Coleoptera: Cicindelidae and Carabidae. In: Expedition to south-west Arabia 1937 - 8. British Museum (Natural History), 1, 87 - 131." type="journal article" year="1948">Britton, 1948</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F35F767DFFEBFF90FF34FAE29CE7FAAF" box="[151,229,1377,1400]" name="Yemen" pageId="11">Yemen</collectingCountry>
|
||
in the pronotum widest near to the middle, with sides rectilinear to hardly convex towards the base, and elytra with impunctate striae and smooth intervals. But the former is distinct from latter, in: 1/ larger size of body (7.4. mm in
|
||
<taxonomicName id="4C484D6EFFEBFF90FE86FA2B9D94FA17" box="[293,406,1448,1472]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="B93CEAFFFFEBFF90FE86FA2B9D34FA68" box="[293,310,1448,1471]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FEE6FA2B9D94FA17" box="[325,406,1448,1472]" italics="true" pageId="11">indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20F5784FFEBFF90FE3DFA2A9DDFFA17" box="[414,477,1449,1472]" pageId="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
, vs.
|
||
<quantity id="4CB09B08FFEBFF90FDB7FA249E69FA17" box="[532,619,1447,1472]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="11" unit="mm" value="6.5">6.5 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C484D6EFFEBFF90FD32FA2B9F01FA17" box="[657,771,1448,1472]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FD32FA2B9EA0FA68" box="[657,674,1448,1471]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FD11FA2B9F01FA17" box="[690,771,1448,1472]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
); 2/ uniformly colored pronotum and elytra (vs. pronotum and elytra bicoloured in
|
||
<taxonomicName id="4C484D6EFFEBFF90FE15FA4F9E28FA33" box="[438,554,1484,1508]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FE15FA4F9DC5FA34" box="[438,455,1484,1507]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FE7AFA4F9E28FA33" box="[473,554,1484,1508]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
); 3/ microsculpture distinct on the whole dorsal surface (vs. microsculpture absent on the pronotum and disc of elytra in
|
||
<taxonomicName id="4C484D6EFFEBFF90FD2EFA739EFFF9DF" box="[653,765,1520,1544]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FD2EFA739E9CF9D0" box="[653,670,1520,1543]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FD0FFA739EFFF9DF" box="[684,765,1520,1544]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
); 4/ pronotum with sides and base narrowly testaceus, apex and disc piceous (vs. sides, apex and base broadly testaceus, disc piceous in
|
||
<taxonomicName id="4C484D6EFFEBFF90FBBCF9979891F9FB" box="[1055,1171,1556,1580]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FBBCF9979832F9FC" box="[1055,1072,1556,1579]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FBE1F9979891F9FB" box="[1090,1171,1556,1580]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
); 5/ lateral margins of pronotum completely flat (vs. lateral margins “slightly but broadly reflexed behind” in
|
||
<taxonomicName id="4C484D6EFFEBFF90FBC0F9BB98D1F987" box="[1123,1235,1592,1616]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FBC0F9BB9876F998" box="[1123,1140,1592,1615]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FB21F9BB98D1F987" box="[1154,1235,1592,1616]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
); 6/ base and disc of elytra dark brown (vs. base of elytra testaceous, disc piceous in
|
||
<taxonomicName id="4C484D6EFFEBFF90FC23F9DF9FF2F9A3" box="[896,1008,1628,1652]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B93CEAFFFFEBFF90FC23F9DF9F93F9A4" box="[896,913,1628,1651]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FC3CF9DF9FF2F9A3" box="[927,1008,1628,1652]" italics="true" pageId="11">bicolor</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF736EDFFEBFF90FF64F9039F61F843" blockId="11.[151,1437,151,2013]" pageId="11">
|
||
On the other hand, there is similarity in the microsculpture of pronotum and elytra, as well as, shape of aedeagus of the new species (
|
||
<figureCitation id="13732A68FFEBFF90FE4CF9279E40F96B" box="[495,578,1700,1725]" captionStart="FIGURES 16 – 20" captionStartId="10.[151,264,1558,1580]" captionTargetBox="[173,1405,589,1527]" captionTargetId="figure@10.[150,1436,566,1537]" captionTargetPageId="10" captionText="FIGURES 16 – 20. Holotype of Atrotus indicus sp. n.: Fig. 16, habitus (photo by Assen Ignatov); Fig. 17, mouthparts, dorsal aspect; Fig. 18, median lobe of aedeagus, lateral aspect; Fig. 19, right paramere, internal face; Fig. 20, left paramere, internal face. Scale bars: 0.5 mm (Fig. 17); 0.2 mm (Figs. 18 – 20)." httpUri="https://zenodo.org/record/222920/files/figure.png" pageId="11">Fig. 18</figureCitation>
|
||
) to that of
|
||
<taxonomicName id="4C484D6EFFEBFF90FD6AF9279FD1F96A" authority="Peringuey" authorityName="Peringuey" box="[713,979,1700,1725]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FD6AF9279ED8F96C" box="[713,730,1700,1723]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FD48F9279F5AF96B" box="[747,856,1700,1724]" italics="true" pageId="11">fortipatus</emphasis>
|
||
Péringuey
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD94B1CFFEBFF90FC47F927991CF96B" author="Basilewsky" box="[996,1310,1700,1725]" pageId="11" refString="Basilewsky, P. (1951) Sur le genre Atrotus Peringuey (Col. Carabidae) et quelques considerations sur le peuplement entomologique des hautes montagnes de l'Afrique. Annales de la Societe royale de Zoologie du Belgique, 82, 5 - 18." type="journal article" year="1951">
|
||
Basilewsky 1951: 8,
|
||
<figureCitation id="13732A68FFEBFF90FB7AF927991CF96B" box="[1241,1310,1700,1725]" captionStart="FIGURES 2 – 6" captionStartId="3.[151,264,1018,1040]" captionTargetBox="[216,1396,181,997]" captionTargetId="figure@3.[151,1436,173,997]" captionTargetPageId="3" captionText="FIGURES 2 – 6. Holotype of Mecyclothorax baehri sp. n.: Fig. 2, habitus; Fig. 3, median lobe of aedeagus, left lateral aspect; Fig. 4, right paramere; Fig. 5, left paramere; Fig. 6, urite, dorsal view. Scale bars: 1.5 mm (Fig. 2); 0.5 mm (Figs. 3 – 6)." httpUri="https://zenodo.org/record/222916/files/figure.png" pageId="11">Fig. 2</figureCitation>
|
||
</bibRefCitation>
|
||
), the latter being the
|
||
<typeStatus id="54F3884FFFEBFF90FEA7F94A9D36F936" box="[260,308,1737,1761]" pageId="11">type</typeStatus>
|
||
species of the genus which inhabits the provinces of Western Cape and Eastern Cape of the Republic of South African. However, the former is distinct from the latter in: 1/ the almost symmetrical lobes of labrum (highly asymmetric in
|
||
<taxonomicName id="4C484D6EFFEBFF90FE30F8939E1DF8FF" box="[403,543,1808,1832]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FE30F8939DA6F8F0" box="[403,420,1808,1831]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FE11F8939E1DF8FF" box="[434,543,1808,1832]" italics="true" pageId="11">fortipatus</emphasis>
|
||
</taxonomicName>
|
||
); 2/ pronotum widest around the middle (vs. pronotum widest before the middle in
|
||
<taxonomicName id="4C484D6EFFEBFF90FF14F8B79D45F89B" box="[183,327,1844,1868]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FF14F8B79CCAF89C" box="[183,200,1844,1867]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FF79F8B79D45F89B" box="[218,327,1844,1868]" italics="true" pageId="11">fortipatus</emphasis>
|
||
</taxonomicName>
|
||
); 3/ sides of pronotum rectilinear to hardly convex towards the base (vs. sides slightly sinuate towards the base in
|
||
<taxonomicName id="4C484D6EFFEBFF90FED6F8DB9E03F8A7" box="[373,513,1880,1904]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FED6F8DB9D84F8B8" box="[373,390,1880,1903]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FE37F8DB9E03F8A7" box="[404,513,1880,1904]" italics="true" pageId="11">fortipatus</emphasis>
|
||
</taxonomicName>
|
||
); 4/ moderately impressed elytral striae (vs. striae deep in
|
||
<taxonomicName id="4C484D6EFFEBFF90FB30F8DB991DF8A7" box="[1171,1311,1880,1904]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FB30F8DB98A6F8B8" box="[1171,1188,1880,1903]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FB11F8DB991DF8A7" box="[1202,1311,1880,1904]" italics="true" pageId="11">fortipatus</emphasis>
|
||
</taxonomicName>
|
||
); 5/ elytral intervals flat (vs. intervals more or less convex in
|
||
<taxonomicName id="4C484D6EFFEBFF90FD64F8FF9F51F843" box="[711,851,1916,1940]" class="Insecta" family="Carabidae" genus="Atrotus" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="fortipatus">
|
||
<emphasis id="B93CEAFFFFEBFF90FD64F8FF9EDAF844" box="[711,728,1916,1939]" italics="true" pageId="11">A</emphasis>
|
||
.
|
||
<emphasis id="B93CEAFFFFEBFF90FD45F8FF9F51F843" box="[742,851,1916,1940]" italics="true" pageId="11">fortipatus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526566FFEBFF90FF64F8239D99F80A" pageId="11" type="distribution">
|
||
<paragraph id="8BF736EDFFEBFF90FF64F8239D99F80A" blockId="11.[151,1437,151,2013]" pageId="11">
|
||
<emphasis id="B93CEAFFFFEBFF90FF64F8239D5BF86E" bold="true" box="[199,345,1952,1977]" pageId="11">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F35F767DFFEBFF90FEC9F8239DA4F86E" box="[362,422,1952,1977]" name="India" pageId="11">India</collectingCountry>
|
||
, state Tamil Nadu, Nilgiri Hills Mountain, the vicinity of Coonoor Town, around 1700 meters above sea level.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3526566FFECFF97FF64FF149E97FF02" pageId="12" type="etymology">
|
||
<paragraph id="8BF736EDFFECFF97FF64FF149E97FF02" blockId="12.[151,1436,151,213]" pageId="12">
|
||
<emphasis id="B93CEAFFFFECFF97FF64FF149D44FF67" bold="true" box="[199,326,151,176]" pageId="12">Etymology</emphasis>
|
||
. The specific epithet is a noun in the nominative singular in preposition. It honors
|
||
<collectingCountry id="F35F767DFFECFF97FA8EFF14996BFF67" box="[1325,1385,151,176]" name="India" pageId="12">India</collectingCountry>
|
||
, the country where the new species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |