323 lines
43 KiB
XML
323 lines
43 KiB
XML
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<mods:title id="9D671984DA4056A0F2AED9134ED3EF08">The genus Troglophilus Krauss, 1879 (Orthoptera: Rhaphidophoridae) in the west Balkans</mods:title>
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<mods:namePart id="827A1F6553016D5C6A5D29F7CDF0F1B1">Karaman, Ivo</mods:namePart>
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<mods:namePart id="0475F5A0069BF8D76B604B5728CA0387">Hammouti, Nasera</mods:namePart>
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<mods:namePart id="3A0393DD82A0C55FF75A0A21386594DF">Pavićević, Dragan</mods:namePart>
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<mods:namePart id="FC0E3DD3FFBDC8FFDBCB2B7D77A5A2D9">Kiefer, Andreas</mods:namePart>
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<mods:namePart id="1785FEDD5E4DD9A8FDECC809F703662B">Horvatović, Mladen</mods:namePart>
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<mods:namePart id="8818FA96D3ECC05A3F96CC0E113A2EE8">Seitz, Alfred</mods:namePart>
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<mods:title id="3BFC1276CA0929527D221CBD60FEED51">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="4F052E794AC068CB51EA995483C02627">
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<mods:date id="18D427D0E7089CA145A6DC74173A9904">2011</mods:date>
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<mods:number id="E0B260666D77F12E244BE26777AAE795">2011-11-25</mods:number>
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<mods:number id="96DBFB75A3AF5928CDFB2E7C9C45B18E">163</mods:number>
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<mods:url id="A180B8C545095C466EDFC65AC4EF2CF2">http://dx.doi.org/10.1111/j.1096-3642.2011.00738.x</mods:url>
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<mods:identifier id="E9DE97F458150C273E7976B3D6E71B06" type="DOI">10.1111/j.1096-3642.2011.00738.x</mods:identifier>
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<mods:identifier id="37C73B2ABDC876AD7E0996379F32DA56" type="ISSN">0024-4082</mods:identifier>
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<subSubSection id="C344BC93FFDFFFA6FCFBF963FA83F914" box="[867,1360,1720,1741]" pageId="4" pageNumber="1039" type="nomenclature">
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<paragraph id="8BE1EF18FFDFFFA6FCFBF963FA83F914" blockId="4.[867,1360,1720,1741]" box="[867,1360,1720,1741]" pageId="4" pageNumber="1039">
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<heading id="D0A95874FFDFFFA6FCFBF963FA83F914" box="[867,1360,1720,1741]" centered="true" fontSize="8" level="2" pageId="4" pageNumber="1039" reason="8">
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<taxonomicName id="4C5E949BFFDFFFA6FCFBF963FA83F914" ID-CoL="5993L" authority=": POPOV & CHOBANOV, 2004" authorityName=": POPOV & CHOBANOV" authorityYear="2004" box="[867,1360,1720,1741]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="4" pageNumber="1039" phylum="Arthropoda" rank="species" species="neglectus">
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<emphasis id="B92A330AFFDFFFA6FCFBF963FBD7F915" box="[867,1028,1720,1741]" italics="true" pageId="4" pageNumber="1039">T. NEGLECTUS</emphasis>
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:
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<bibRefCitation id="EFCF92E9FFDFFFA6FB8DF963FA83F914" author="Popov A & Chobanov D" box="[1045,1360,1720,1741]" pageId="4" pageNumber="1039" pagination="241 - 309" refId="ref11349" refString="Popov A, Chobanov D. 2004. Blattodea, mantodea, isoptera, orthoptera and dermaptera of the Eastern Rhodopes (bulgaria and greece). In: Beron P, Popov A, eds. Biodiversity of Bulgaria 2. Biodiversity of Eastern Rhodopes (Bulgaria and Greece). Sofia: Pensoft & National Museum of Natural History, 241 - 309." type="book chapter" year="2004">POPOV & CHOBANOV, 2004</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C344BC93FFDFFFA5FCBEF905FB4CFB8A" lastPageId="7" lastPageNumber="1042" pageId="4" pageNumber="1039" type="materials_examined">
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<paragraph id="8BE1EF18FFDFFFA6FCBEF905FB65F92D" blockId="4.[806,1206,1758,1780]" box="[806,1206,1758,1780]" pageId="4" pageNumber="1039">
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<emphasis id="B92A330AFFDFFFA6FCBEF905FBD5F92A" box="[806,1030,1758,1779]" italics="true" pageId="4" pageNumber="1039">Material examined:</emphasis>
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See Appendix.
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</paragraph>
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<paragraph id="8BE1EF18FFDFFFA7FCBEF8C7FD4BF8B6" blockId="4.[806,1421,1820,1903]" lastBlockId="5.[164,779,1514,1903]" lastPageId="5" lastPageNumber="1040" pageId="4" pageNumber="1039">
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<emphasis id="B92A330AFFDFFFA6FCBEF8C7FC4EF8E8" box="[806,925,1820,1841]" italics="true" pageId="4" pageNumber="1039">Diagnosis:</emphasis>
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Small to medium sized species. Male body length ranges between
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<quantity id="4CA642FDFFDFFFA6FB06F8E1FAC5F889" box="[1182,1302,1850,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.8" metricValueMin="1.1" pageId="4" pageNumber="1039" unit="mm" value="14.5" valueMax="18.0" valueMin="11.0">11–18 mm</quantity>
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; females,
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<quantity id="4CA642FDFFDFFFA6FCBEF882FC73F8B7" box="[806,928,1881,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="4" pageNumber="1039" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 mm</quantity>
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. Body colour dark brown to light brown marbled. The male tenth tergite with wide, partially square, medial protrusion distally divided by a central depression in two triangularly shaped lobes (
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<figureCitation id="1365F39DFFDEFFA7FF34F99EFEF1F985" box="[172,290,1605,1628]" captionStart="Figure 3" captionStartId="5.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-207@5.[323,1283,196,1286]" captionTargetPageId="5" captionText="Figure 3. A–F, Troglophilus brevicauda, tenth tergite, dorsal view: A, male 15 mm (Bjeloševina, Montenegro); B, male 16.5 mm (Plačkovica, Macedonia); C, male 14 mm (Olovo, Bosnia); D, female 15 mm (Olovo, Bosnia); E, female 14 mm (Dērekare, Serbia); F, female 16 mm (Petnica, Serbia)." figureDoi="http://doi.org/10.5281/zenodo.5442329" httpUri="https://zenodo.org/record/5442329/files/figure.png" pageId="5" pageNumber="1040">Fig. 3A–C</figureCitation>
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). Posterior edge of the lobes slightly undulating. The female tenth abdominal tergite with two paramedial tooth form protrusions, variable in form from slightly noticeable (
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<figureCitation id="1365F39DFFDEFFA7FE50F97AFDCAF96E" box="[456,537,1697,1719]" captionStart="Figure 3" captionStartId="5.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-207@5.[323,1283,196,1286]" captionTargetPageId="5" captionText="Figure 3. A–F, Troglophilus brevicauda, tenth tergite, dorsal view: A, male 15 mm (Bjeloševina, Montenegro); B, male 16.5 mm (Plačkovica, Macedonia); C, male 14 mm (Olovo, Bosnia); D, female 15 mm (Olovo, Bosnia); E, female 14 mm (Dērekare, Serbia); F, female 16 mm (Petnica, Serbia)." figureDoi="http://doi.org/10.5281/zenodo.5442329" httpUri="https://zenodo.org/record/5442329/files/figure.png" pageId="5" pageNumber="1040">Fig. 3F</figureCitation>
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) to very pronounced and wide (
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<figureCitation id="1365F39DFFDEFFA7FEB3F91BFEACF90F" box="[299,383,1728,1750]" captionStart="Figure 3" captionStartId="5.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-207@5.[323,1283,196,1286]" captionTargetPageId="5" captionText="Figure 3. A–F, Troglophilus brevicauda, tenth tergite, dorsal view: A, male 15 mm (Bjeloševina, Montenegro); B, male 16.5 mm (Plačkovica, Macedonia); C, male 14 mm (Olovo, Bosnia); D, female 15 mm (Olovo, Bosnia); E, female 14 mm (Dērekare, Serbia); F, female 16 mm (Petnica, Serbia)." figureDoi="http://doi.org/10.5281/zenodo.5442329" httpUri="https://zenodo.org/record/5442329/files/figure.png" pageId="5" pageNumber="1040">Fig. 3D</figureCitation>
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).The titillator almost triangular in shape, distally widely obtuse (
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<figureCitation id="1365F39DFFDEFFA7FDB7F905FD50F92C" box="[559,643,1758,1781]" captionStart="Figure 4" captionStartId="6.[144,223,1316,1335]" captionTargetBox="[303,1263,196,1286]" captionTargetId="figure-234@6.[303,1263,196,1286]" captionTargetPageId="6" captionText="Figure 4. A–E, Troglophilus zorae sp. nov., paratypes: A, male 16 mm, tenth tergite, dorsal view (Mlečnik, Macedonia); B, male 19.5 mm, tenth tergite, dorsal view (Matka, Macedonia); C, female 15.5 mm, tenth tergite, dorsal view (Mlečnik, Macedonia); D, female 18 mm, tenth tergite, dorsal view (Matka, Macedonia); E, male 19.5 mm, phallus with titillator (arrow), dorsal view (Matka, Macedonia). F, Troglophilus brevicauda, male 15 mm, phallus with titillator (arrow), dorsal view (Olovo, Bosnia)." figureDoi="http://doi.org/10.5281/zenodo.5442331" httpUri="https://zenodo.org/record/5442331/files/figure.png" pageId="5" pageNumber="1040">Fig. 4F</figureCitation>
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). In lateral view, slender, distally slightly voluminous. The ovipositor short and wide, length ranges between 7.5–
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<quantity id="4CA642FDFFDEFFA7FF3CF8E1FF27F889" box="[164,244,1850,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="1040" unit="mm" value="10.0">10 mm</quantity>
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; the widest point is in its proximal third (
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<figureCitation id="1365F39DFFDEFFA7FF34F882FEC2F8B6" box="[172,273,1881,1903]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="5" pageNumber="1040">Fig. 12G</figureCitation>
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), ventral margin almost straight.
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</paragraph>
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<caption id="DF21BF90FFDEFFA7FF3CFAFFFD38FAA8" ID-DOI="http://doi.org/10.5281/zenodo.5442329" ID-Zenodo-Dep="5442329" httpUri="https://zenodo.org/record/5442329/files/figure.png" pageId="5" pageNumber="1040" startId="5.[164,243,1316,1335]" targetBox="[323,1283,196,1286]" targetPageId="5">
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<paragraph id="8BE1EF18FFDEFFA7FF3CFAFFFD38FAA8" blockId="5.[164,1441,1316,1394]" pageId="5" pageNumber="1040">
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<emphasis id="B92A330AFFDEFFA7FF3CFAFFFEDFFAEE" bold="true" box="[164,268,1316,1335]" pageId="5" pageNumber="1040">Figure 3.</emphasis>
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A–F,
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<taxonomicName id="4C5E949BFFDEFFA7FEC9FAFFFD81FAEE" authorityName=": HARZ" authorityYear="1969" box="[337,594,1316,1335]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="1040" phylum="Arthropoda" rank="species" species="brevicauda">
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<emphasis id="B92A330AFFDEFFA7FEC9FAFFFD81FAEE" box="[337,594,1316,1335]" italics="true" pageId="5" pageNumber="1040">Troglophilus brevicauda</emphasis>
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</taxonomicName>
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, tenth tergite, dorsal view: A, male 15 mm (Bjeloševina, Montenegro); B, male 16.5 mm (Plačkovica, Macedonia); C, male 14 mm (Olovo, Bosnia); D, female 15 mm (Olovo, Bosnia); E, female 14 mm (Dērekare, Serbia); F, female 16 mm (Petnica, Serbia).
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</paragraph>
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</caption>
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<paragraph id="8BE1EF18FFDEFFA7FCA2FA31FC08F8B7" blockId="5.[826,1441,1514,1902]" pageId="5" pageNumber="1040">
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<emphasis id="B92A330AFFDEFFA7FCA2FA31FC1FFA26" box="[826,972,1514,1535]" italics="true" pageId="5" pageNumber="1040">Distribution:</emphasis>
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This taxon appears with disjunct distribution in three regions (
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<figureCitation id="1365F39DFFDEFFA7FBC7F9D3FB72F9C7" box="[1119,1185,1544,1566]" captionStart="Figure 5" captionStartId="7.[164,243,778,797]" captionTargetBox="[387,1219,196,749]" captionTargetId="figure-561@7.[387,1219,196,749]" captionTargetPageId="7" captionText="Figure 5. The geographical distribution of Troglophilus brevicauda (circles) and Troglophilus zorae sp. nov. (triangles) (square – literature data). (SLO-Slovenia; CRO-Croatia; BIH-Bosnia and Herzegovina; SRB-Serbia; MNE-Montenegro; BUL-Bulgaria; ALB-Albania; MK-Macedonia; GRE-Greece)." figureDoi="http://doi.org/10.5281/zenodo.5442333" httpUri="https://zenodo.org/record/5442333/files/figure.png" pageId="5" pageNumber="1040">Fig. 5</figureCitation>
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). It is widely distributed in montane and submontane parts of western
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<collectingCountry id="F349AF88FFDEFFA7FCA2F99EFC55F982" box="[826,902,1605,1627]" name="Serbia" pageId="5" pageNumber="1040">Serbia</collectingCountry>
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, eastern
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<collectingCountry id="F349AF88FFDEFFA7FC6FF99EFB9BF982" box="[1015,1096,1605,1627]" name="Bosnia and Herzegovina" pageId="5" pageNumber="1040">Bosnia</collectingCountry>
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, and northern parts of
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<collectingCountry id="F349AF88FFDEFFA7FAFEF99DFC40F9A0" name="Serbia" pageId="5" pageNumber="1040">Montenegro</collectingCountry>
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. A remote local population occurs in the region of Rhodopes in
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<collectingCountry id="F349AF88FFDEFFA7FBC5F958FB17F940" box="[1117,1220,1667,1689]" name="Bulgaria" pageId="5" pageNumber="1040">Bulgaria</collectingCountry>
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. Unfortunately, it was not possible for us to carry out a morphological study of this population, so only a genetic analysis on legs kindly sent to us by our colleague D. Čobanov (
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<collectingRegion id="499A21FAFFDEFFA7FCD9F926FCA8F8CA" box="[833,891,1789,1811]" country="Bulgaria" name="Sofiya-Grad" pageId="5" pageNumber="1040">Sofia</collectingRegion>
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) was performed. An isolated population from eastern
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<collectingCountry id="F349AF88FFDEFFA7FC04F8C7FBC4F8EB" box="[924,1047,1820,1842]" name="Macedonia" pageId="5" pageNumber="1040">Macedonia</collectingCountry>
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(Plačkovica Mt.) showed a large genetic distance (
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<collectionCode id="ED4F77DDFFDEFFA7FB9DF8E1FBFFF889" box="[1029,1068,1850,1872]" pageId="5" pageNumber="1040">HT</collectionCode>
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3) from the other populations of
|
||
<taxonomicName id="4C5E949BFFDEFFA7FCA2F882FC07F8B7" authorityName=": HARZ" authorityYear="1969" box="[826,980,1881,1902]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="5" pageNumber="1040" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDEFFA7FCA2F882FC07F8B7" box="[826,980,1881,1902]" italics="true" pageId="5" pageNumber="1040">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF21BF90FFDDFFA4FF08FAFFFE68FA75" ID-DOI="http://doi.org/10.5281/zenodo.5442331" ID-Zenodo-Dep="5442331" httpUri="https://zenodo.org/record/5442331/files/figure.png" pageId="6" pageNumber="1041" startId="6.[144,223,1316,1335]" targetBox="[303,1263,196,1286]" targetPageId="6">
|
||
<paragraph id="8BE1EF18FFDDFFA4FF08FAFFFE68FA75" blockId="6.[144,1421,1315,1452]" pageId="6" pageNumber="1041">
|
||
<emphasis id="B92A330AFFDDFFA4FF08FAFFFF2BFAEE" bold="true" box="[144,248,1316,1335]" pageId="6" pageNumber="1041">Figure 4.</emphasis>
|
||
A–E,
|
||
<emphasis id="B92A330AFFDDFFA4FEDAFAF8FD56FAEE" bold="true" box="[322,645,1315,1335]" pageId="6" pageNumber="1041">
|
||
<taxonomicName id="4C5E949BFFDDFFA4FEDAFAF8FDF3FAEE" authority="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz, 2011" authorityName="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz" authorityYear="2011" box="[322,544,1315,1335]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="zorae" status="sp. nov.">
|
||
<emphasis id="B92A330AFFDDFFA4FEDAFAF8FDF3FAEE" bold="true" box="[322,544,1315,1335]" italics="true" pageId="6" pageNumber="1041">Troglophilus zorae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2198E71FFDDFFA4FDB3FAFFFD56FAEE" box="[555,645,1316,1335]" pageId="6" pageNumber="1041" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratypes: A, male 16 mm, tenth tergite, dorsal view (Mlečnik, Macedonia); B, male 19.5 mm, tenth tergite, dorsal view (Matka, Macedonia); C, female 15.5 mm, tenth tergite, dorsal view (Mlečnik, Macedonia); D, female 18 mm, tenth tergite, dorsal view (Matka, Macedonia); E, male 19.5 mm, phallus with titillator (arrow), dorsal view (Matka, Macedonia). F,
|
||
<taxonomicName id="4C5E949BFFDDFFA4FD23FAA7FC6AFA56" authorityName=": HARZ" authorityYear="1969" box="[699,953,1404,1423]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDDFFA4FD23FAA7FC6AFA56" box="[699,953,1404,1423]" italics="true" pageId="6" pageNumber="1041">Troglophilus brevicauda</emphasis>
|
||
</taxonomicName>
|
||
, male 15 mm, phallus with titillator (arrow), dorsal view (Olovo, Bosnia).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1EF18FFDDFFA4FF08FA31FBE1F8B6" blockId="6.[144,759,1514,1903]" lastBlockId="6.[806,1422,1514,1903]" pageId="6" pageNumber="1041">
|
||
<emphasis id="B92A330AFFDDFFA4FF08FA31FF28FA26" box="[144,251,1514,1535]" italics="true" pageId="6" pageNumber="1041">Remarks:</emphasis>
|
||
This species was originally described by
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FF08F9D3FE94F9C7" author="Chopard L" box="[144,327,1544,1566]" pageId="6" pageNumber="1041" pagination="137 - 139" refId="ref10416" refString="Chopard L. 1934. Diagnoses d'Orthopteres cavernicoles nouveaux. Bulletin de la Societe entomologique de France 39: 137 - 139." type="journal article" year="1934">Chopard (1934)</bibRefCitation>
|
||
, based on a single female specimen from a cave in the village of Bjeloševina (now in
|
||
<collectingCountry id="F349AF88FFDDFFA4FF08F99DFECEF982" box="[144,285,1606,1627]" name="Serbia" pageId="6" pageNumber="1041">Montenegro</collectingCountry>
|
||
). Later on,
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FE38F99EFD81F985" author="Chopard L" box="[416,594,1605,1628]" pageId="6" pageNumber="1041" pagination="509 - 514" refId="ref10439" refString="Chopard L. 1940. Orthopteres cavernicoles recueillis par M. P. Remy en Yougoslavie et en Macedoine. Comparaison entre la distribution des Orthopteres et des Coleopteres cavernicoles d'Europe. VI Congreso Internacional Entomologia, Madrid 6 - 12 Septiembre, 1935: 509 - 514." type="book chapter" year="1940">Chopard (1940)</bibRefCitation>
|
||
cited the type series of this species, with the female
|
||
<typeStatus id="54E551BAFFDDFFA4FDAEF9BFFD44F9A3" box="[566,663,1636,1658]" pageId="6" pageNumber="1041" type="holotype">holotype</typeStatus>
|
||
and
|
||
<specimenCount id="9D582491FFDDFFA4FD55F9BFFF1FF940" pageId="6" pageNumber="1041" type="adult">two adult</specimenCount>
|
||
males and
|
||
<specimenCount id="9D582491FFDDFFA4FED6F958FE0BF940" box="[334,472,1667,1689]" pageId="6" pageNumber="1041" type="juvenile">one juvenile</specimenCount>
|
||
female. It remains unexplained as to why he did not present the characteristics of the males in his description. As the description is very featureless in most parts and without any illustrations, this led some authors to interpret it differently. Thus, in papers of
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FE56F8C7FD43F8E8" author="Karaman Z" box="[462,656,1819,1841]" pageId="6" pageNumber="1041" pagination="211 - 228" refId="ref10844" refString="Karaman Z. 1958. Spiljski zrikavci Jugoslavije. Godisnji Zbornik Univerziteta Skopje 11: 211 - 228. (In Serbian)." type="journal article" year="1958">Karaman (1958)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FD4FF8C7FF0AF889" author="Us P" pageId="6" pageNumber="1041" pagination="301 - 319" refId="ref11629" refString="Us P. 1970. Prispevek k poznavanju jamskih ortopterov Jugoslavije (Orthoptera - Tettigonoidea). Acta Carsologica SAZU 5: 301 - 319." type="journal article" year="1970">Us (1970)</bibRefCitation>
|
||
, some specimens of
|
||
<taxonomicName id="4C5E949BFFDDFFA4FE5BF8E0FDE8F896" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[451,571,1850,1872]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDDFFA4FE5BF8E0FDE8F896" box="[451,571,1850,1872]" italics="true" pageId="6" pageNumber="1041">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
were treated as
|
||
<taxonomicName id="4C5E949BFFDDFFA4FF08F882FEFAF8B7" authorityName=": HARZ" authorityYear="1969" box="[144,297,1881,1902]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDDFFA4FF08F882FEFAF8B7" box="[144,297,1881,1902]" italics="true" pageId="6" pageNumber="1041">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
. The probable reason for this is the specimen of a subadult female of
|
||
<taxonomicName id="4C5E949BFFDDFFA4FB03FA31FAC0FA26" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[1179,1299,1514,1535]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDDFFA4FB03FA31FAC0FA26" box="[1179,1299,1514,1535]" italics="true" pageId="6" pageNumber="1041">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
from locus typicus (collected in the same time as the type specimen of
|
||
<taxonomicName id="4C5E949BFFDDFFA4FCE0F9FCFBC1F9E5" authorityName=": HARZ" authorityYear="1969" box="[888,1042,1575,1596]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDDFFA4FCE0F9FCFBC1F9E5" box="[888,1042,1575,1596]" italics="true" pageId="6" pageNumber="1041">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
), and which Z. Karaman received from the National Museum of Natural History in Paris as
|
||
<taxonomicName id="4C5E949BFFDDFFA4FCDEF9BEFC33F9A0" authorityName=": HARZ" authorityYear="1969" box="[838,992,1636,1658]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDDFFA4FCDEF9BEFC33F9A0" box="[838,992,1636,1658]" italics="true" pageId="6" pageNumber="1041">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
(now in the collection of I. Karaman). It can be justly supposed that this is the same specimen that
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FC01F97AFB9FF96E" author="Chopard L" box="[921,1100,1697,1719]" pageId="6" pageNumber="1041" pagination="509 - 514" refId="ref10439" refString="Chopard L. 1940. Orthopteres cavernicoles recueillis par M. P. Remy en Yougoslavie et en Macedoine. Comparaison entre la distribution des Orthopteres et des Coleopteres cavernicoles d'Europe. VI Congreso Internacional Entomologia, Madrid 6 - 12 Septiembre, 1935: 509 - 514." type="book chapter" year="1940">Chopard (1940)</bibRefCitation>
|
||
cited for the type series. Z.
|
||
<bibRefCitation id="EFCF92E9FFDDFFA4FCBEF91BFC3AF90F" author="Karaman Z" box="[806,1001,1728,1750]" pageId="6" pageNumber="1041" pagination="211 - 228" refId="ref10844" refString="Karaman Z. 1958. Spiljski zrikavci Jugoslavije. Godisnji Zbornik Univerziteta Skopje 11: 211 - 228. (In Serbian)." type="journal article" year="1958">Karaman (1958)</bibRefCitation>
|
||
treated this specimen as
|
||
<taxonomicName id="4C5E949BFFDDFFA4FAA1F91BFCA0F92A" authorityName=": HARZ" authorityYear="1969" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDDFFA4FAA1F91BFCA0F92A" italics="true" pageId="6" pageNumber="1041">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
, as well as the specimens of
|
||
<taxonomicName id="4C5E949BFFDDFFA4FB5AF904FAE9F92A" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[1218,1338,1758,1780]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="1041" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDDFFA4FB5AF904FAE9F92A" box="[1218,1338,1758,1780]" italics="true" pageId="6" pageNumber="1041">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
from a locality in western
|
||
<collectingCountry id="F349AF88FFDDFFA4FC65F926FB9AF8CA" box="[1021,1097,1789,1811]" name="Serbia" pageId="6" pageNumber="1041">Serbia</collectingCountry>
|
||
, where males had somewhat more distinct protrusions of the tenth tergite. Chopard’s undisputed authority should not be ignored in this misunderstanding.
|
||
</paragraph>
|
||
<caption id="DF21BF90FFDCFFA5FF3CFCD1FCCCFC81" ID-DOI="http://doi.org/10.5281/zenodo.5442333" ID-Zenodo-Dep="5442333" httpUri="https://zenodo.org/record/5442333/files/figure.png" pageId="7" pageNumber="1042" startId="7.[164,243,778,797]" targetBox="[387,1219,196,749]" targetPageId="7">
|
||
<paragraph id="8BE1EF18FFDCFFA5FF3CFCD1FCCCFC81" blockId="7.[164,1442,778,856]" pageId="7" pageNumber="1042">
|
||
<emphasis id="B92A330AFFDCFFA5FF3CFCD1FEDFFCC4" bold="true" box="[164,268,778,797]" pageId="7" pageNumber="1042">Figure 5.</emphasis>
|
||
The geographical distribution of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FDF6FCD1FCBFFCC4" authorityName=": HARZ" authorityYear="1969" box="[622,876,778,797]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDCFFA5FDF6FCD1FCBFFCC4" box="[622,876,778,797]" italics="true" pageId="7" pageNumber="1042">Troglophilus brevicauda</emphasis>
|
||
</taxonomicName>
|
||
(circles) and
|
||
<emphasis id="B92A330AFFDCFFA5FC60FCD1FAFCFCC4" bold="true" box="[1016,1327,778,798]" pageId="7" pageNumber="1042">
|
||
<taxonomicName id="4C5E949BFFDCFFA5FC60FCD1FB01FCC7" authority="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz, 2011" authorityName="Karaman & Hammouti & Pavićević & Kiefer & Horvatović & Seitz" authorityYear="2011" box="[1016,1234,778,798]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="zorae" status="sp. nov.">
|
||
<emphasis id="B92A330AFFDCFFA5FC60FCD1FB01FCC7" bold="true" box="[1016,1234,778,798]" italics="true" pageId="7" pageNumber="1042">Troglophilus zorae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2198E71FFDCFFA5FB40FCD1FAFCFCC4" box="[1240,1327,778,797]" pageId="7" pageNumber="1042" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(triangles) (square – literature data). (SLO-Slovenia; CRO-Croatia; BIH-Bosnia and Herzegovina; SRB-Serbia; MNE-Montenegro; BUL-Bulgaria; ALB-Albania; MK-Macedonia; GRE-Greece).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE1EF18FFDCFFA5FF24FC7FFB4CFB8A" blockId="7.[164,779,932,1903]" lastBlockId="7.[826,1441,932,1107]" pageId="7" pageNumber="1042">
|
||
Unfortunately, we also did not have the opportunity to analyse the type specimen, as it was not found in the museum in Paris (L. Desutter, pers. comm.). The status of this species is additionally complicated by the uncertain location of the type locality. Even in Pljevlja, the closest larger settlement, in northern
|
||
<collectingCountry id="F349AF88FFDCFFA5FF3CFB87FEFCFBA8" box="[164,303,1116,1137]" name="Serbia" pageId="7" pageNumber="1042">Montenegro</collectingCountry>
|
||
, not many people have heard of a very small and remote village of Bjeloševina. We have been very lucky to find the village and the cave and to collect specimens of two
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE57FB63FD68FB17" box="[463,699,1208,1230]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B92A330AFFDCFFA5FE57FB63FD8DFB14" box="[463,606,1208,1229]" italics="true" pageId="7" pageNumber="1042">Troglophilus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
there. One was
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE88FB0CFE5BFB32" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[272,392,1238,1260]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDCFFA5FE88FB0CFE5BFB32" box="[272,392,1238,1260]" italics="true" pageId="7" pageNumber="1042">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
and the other was a species that was well described by
|
||
<bibRefCitation id="EFCF92E9FFDCFFA5FE2EFB2EFD85FAD2" author="Maran J" box="[438,598,1269,1291]" pageId="7" pageNumber="1042" pagination="387 - 393" refId="ref11056" refString="Maran J. 1958. Beitrag zur Kenntnis der geographischen Variabilitat von Troglophilus neglectus Krauss. Orthoptera Raphidophoridae. Acta Entomologica Musei Nationalis Pragae 32: 387 - 393." type="journal article" year="1958">Maran (1958)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C5E949BFFDCFFA5FD1FFB2EFED3FAF1" authorityName="Maran" authorityYear="1958" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="serbicus">
|
||
<emphasis id="B92A330AFFDCFFA5FD1FFB2EFED3FAF1" italics="true" pageId="7" pageNumber="1042">T. neglectus serbicus</emphasis>
|
||
</taxonomicName>
|
||
. The similar sympatric appearance of these two, the only
|
||
<taxonomicName id="4C5E949BFFDCFFA5FED9FAE9FE03FA9E" authorityName="Krauss" authorityYear="1879" box="[321,464,1330,1351]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92A330AFFDCFFA5FED9FAE9FE03FA9E" box="[321,464,1330,1351]" italics="true" pageId="7" pageNumber="1042">Troglophilus</emphasis>
|
||
</taxonomicName>
|
||
taxa in the wider region of that part of
|
||
<collectingCountry id="F349AF88FFDCFFA5FEAFFA8AFD9BFABE" box="[311,584,1361,1383]" name="Serbia" pageId="7" pageNumber="1042">Montenegro and Serbia</collectingCountry>
|
||
is common. The dilemma we had was which species of the mentioned two is the synonym of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE23FA55FD86FA7A" authorityName=": HARZ" authorityYear="1969" box="[443,597,1422,1443]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDCFFA5FE23FA55FD86FA7A" box="[443,597,1422,1443]" italics="true" pageId="7" pageNumber="1042">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
. Two points in the description of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE1FFA76FDF2FA18" authorityName=": HARZ" authorityYear="1969" box="[391,545,1452,1474]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDCFFA5FE1FFA76FDF2FA18" box="[391,545,1452,1474]" italics="true" pageId="7" pageNumber="1042">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
brought us to the conclusion that it is
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE0CFA17FDAEFA39" authorityName="Maran" authorityYear="1958" box="[404,637,1483,1505]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="serbicus">
|
||
<emphasis id="B92A330AFFDCFFA5FE0CFA17FDAEFA39" box="[404,637,1483,1505]" italics="true" pageId="7" pageNumber="1042">T. neglectus serbicus</emphasis>
|
||
</taxonomicName>
|
||
. The length of the ovipositor in the description completely matches the length of the ovipositor of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FD1FF9D2FED3F9E5" authorityName="Maran" authorityYear="1958" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="serbicus">
|
||
<emphasis id="B92A330AFFDCFFA5FD1FF9D2FED3F9E5" italics="true" pageId="7" pageNumber="1042">T. neglectus serbicus</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is much longer in
|
||
<taxonomicName id="4C5E949BFFDCFFA5FD14F9FCFCD7F9E5" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[652,772,1575,1596]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDCFFA5FD14F9FCFCD7F9E5" box="[652,772,1575,1596]" italics="true" pageId="7" pageNumber="1042">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
. Although this character can vary, we have never found specimens of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FE15F9BEFDD6F9A0" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[397,517,1636,1658]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDCFFA5FE15F9BEFDD6F9A0" box="[397,517,1636,1658]" italics="true" pageId="7" pageNumber="1042">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
with the ovipositor of similar length as in
|
||
<typeStatus id="54E551BAFFDCFFA5FE50F958FDFAF940" box="[456,553,1667,1689]" pageId="7" pageNumber="1042" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FDF2F958FCD7F941" authorityName=": HARZ" authorityYear="1969" box="[618,772,1667,1688]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDCFFA5FDF2F958FCD7F941" box="[618,772,1667,1688]" italics="true" pageId="7" pageNumber="1042">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
. Another detail is the description of the margin of the tenth tergite, which could be treated as the
|
||
<taxonomicName id="4C5E949BFFDCFFA5FD0BF91BFCD8F90C" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[659,779,1728,1749]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDCFFA5FD0BF91BFCD8F90C" box="[659,779,1728,1749]" italics="true" pageId="7" pageNumber="1042">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
type. However, the observation of truncated apical margin ‘
|
||
<emphasis id="B92A330AFFDCFFA5FE9AF926FEBDF8E8" italics="true" pageId="7" pageNumber="1042">
|
||
10
|
||
<superScript id="7C2B4250FFDCFFA5FE86F920FEF7F8DE" attach="left" box="[286,292,1787,1799]" fontSize="5" pageId="7" pageNumber="1042">e</superScript>
|
||
tergite abdominal un peu tronqué á l’apex, á angles arrondis
|
||
</emphasis>
|
||
’ (
|
||
<bibRefCitation id="EFCF92E9FFDCFFA5FE1BF8C7FDFEF8EB" author="Chopard L" box="[387,557,1820,1842]" pageId="7" pageNumber="1042" pagination="137 - 139" refId="ref10416" refString="Chopard L. 1934. Diagnoses d'Orthopteres cavernicoles nouveaux. Bulletin de la Societe entomologique de France 39: 137 - 139." type="journal article" year="1934">Chopard, 1934</bibRefCitation>
|
||
) resembles greatly the anatomy of tergites we see in some populations of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FF3CF882FE42F8B7" authorityName="Maran" authorityYear="1958" box="[164,401,1881,1902]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="serbicus">
|
||
<emphasis id="B92A330AFFDCFFA5FF3CF882FE42F8B7" box="[164,401,1881,1902]" italics="true" pageId="7" pageNumber="1042">T. neglectus serbicus</emphasis>
|
||
</taxonomicName>
|
||
from western
|
||
<collectingCountry id="F349AF88FFDCFFA5FDD2F882FD47F8B6" box="[586,660,1881,1903]" name="Serbia" pageId="7" pageNumber="1042">Serbia</collectingCountry>
|
||
(
|
||
<figureCitation id="1365F39DFFDCFFA5FD32F882FD2DF8B6" box="[682,766,1881,1903]" captionStart="Figure 2" captionStartId="4.[144,223,1236,1255]" captionTargetBox="[409,1159,203,1201]" captionTargetId="figure-247@4.[399,1167,196,1206]" captionTargetPageId="4" captionText="Figure 2. Bayesian inference for the Troglophilus species tree based on the analysis of the 16S sequences. Posterior probability values are shown above the branches when they are> 0.5. Bootstrap values (2000 pseudoreplicates) given by neighbour joining (NJ) and maximum parsimony (MP) are presented below the branches when they are>50% (NJ/MP)." figureDoi="http://doi.org/10.5281/zenodo.5442327" httpUri="https://zenodo.org/record/5442327/files/figure.png" pageId="7" pageNumber="1042">Fig. 2F</figureCitation>
|
||
). Unfortunately, we have not found females at the type locality. The margin of the tenth tergite in females of
|
||
<taxonomicName id="4C5E949BFFDCFFA5FCA2FC39FC61FC2F" baseAuthorityName="Kollar" baseAuthorityYear="1833" box="[826,946,993,1015]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="cavicola">
|
||
<emphasis id="B92A330AFFDCFFA5FCA2FC39FC61FC2F" box="[826,946,993,1015]" italics="true" pageId="7" pageNumber="1042">T. cavicola</emphasis>
|
||
</taxonomicName>
|
||
is always in the form of a shallow concave cavity (
|
||
<figureCitation id="1365F39DFFDCFFA5FC15FBDBFC23FBCF" box="[909,1008,1024,1046]" captionStart="Figure 12" captionStartId="15.[164,243,1316,1335]" captionTargetBox="[323,1283,196,1286]" captionTargetId="figure-192@15.[323,1283,196,1286]" captionTargetPageId="15" captionText="Figure 12. A–C, Troglophilus cavicola: A, male 18 mm, tenth tergite, dorsal view (Bjeloševina, Montenegro); B, female 18.5 mm, tenth tergite, dorsal view (Bukovik, Macedonia); C, male 18 mm, phallus with titillator (arrow), lateral view (Bjeloševina, Montenegro). D–I ovipositor, lateral view: D, Troglophilus lazaropolensis, female 16.5 mm (Lazaropole, Macedonia); E, Troglophilus cavicola, female 18.5 mm (Bjeloševina, Montenegro); F, Troglophilus zorae sp. nov., paratype female 18 mm (Matka, Macedonia); G, Troglophilus brevicauda, female 15 mm (Olovo, Bosnia); H, Troglophilus ovuliformis, female 19.5 mm (D. Seoca, Montenegro); I, Troglophilus neglectus, female 15 mm (N. Breznica, Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5442347" httpUri="https://zenodo.org/record/5442347/files/figure.png" pageId="7" pageNumber="1042">Fig. 12B</figureCitation>
|
||
). This all led us to the conclusion that
|
||
<taxonomicName id="4C5E949BFFDCFFA5FCA2FBC4FC07FBED" authorityName=": HARZ" authorityYear="1969" box="[826,980,1055,1076]" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="species" species="brevicauda">
|
||
<emphasis id="B92A330AFFDCFFA5FCA2FBC4FC07FBED" box="[826,980,1055,1076]" italics="true" pageId="7" pageNumber="1042">T. brevicauda</emphasis>
|
||
</taxonomicName>
|
||
is a valid species and that
|
||
<taxonomicName id="4C5E949BFFDCFFA5FA85FBC4FC45FB8B" authorityName="Maran" authorityYear="1958" class="Insecta" family="Rhaphidophoridae" genus="Troglophilus" kingdom="Animalia" order="Orthoptera" pageId="7" pageNumber="1042" phylum="Arthropoda" rank="subSpecies" species="neglectus" subSpecies="serbicus">
|
||
<emphasis id="B92A330AFFDCFFA5FA85FBC4FC45FB8B" italics="true" pageId="7" pageNumber="1042">T. neglectus serbicus</emphasis>
|
||
</taxonomicName>
|
||
is its junior synonym.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |