138 lines
14 KiB
XML
138 lines
14 KiB
XML
<document id="ADEEAA3A2BA93BF47B5B5EC7B2118BC9" ID-DOI="10.1649/0010-065x(2000)054[0239:atootg]2.0.co;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="4900237" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622827576800" checkinUser="carolina" docAuthor="Carpaneto, Giuseppe M., Mignani, Roberto & Piattella, Emanuele" docDate="2000" docId="03DE1747D8588B56FEA215121E8CFC77" docLanguage="en" docName="TheColeopBull.54.2.239-247.pdf.imf" docOrigin="The Coleopterists Bulletin 54 (2)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Pseudoathyreus kordofanus" docType="treatment" docVersion="3" lastPageNumber="240" masterDocId="FFE76F3FD8588B57FF9B13741F23FFC0" masterDocTitle="A Taxonomic Overview Of The Genus Pseudoathyreus Howden And Martinez, And Redescription Of Two Related Species From Southern Africa (Coleoptera: Geotrupidae: Bolboceratinae)" masterLastPageNumber="247" masterPageNumber="239" pageNumber="239" updateTime="1698955049550" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="89B325F5FA2AFBDC741B6429E5427CE5">A Taxonomic Overview Of The Genus Pseudoathyreus Howden And Martinez, And Redescription Of Two Related Species From Southern Africa (Coleoptera: Geotrupidae: Bolboceratinae)</mods:title>
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<mods:name id="B269E25D0B73D114A9AE049AEFF486A0" type="personal">
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<mods:roleTerm id="A9EF6234A54CA652FA6D0C4F5DE2C603">Author</mods:roleTerm>
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<mods:namePart id="B8BC31B83F1BAF2513684D7A55C97417">Carpaneto, Giuseppe M.</mods:namePart>
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<mods:affiliation id="48ABE88634B332678BCE9A11F0B0BB3C">Dipartimento di Biologia Università degli Studi Roma Tre Viale G. Marconi 446 00146 Roma, ITALY</mods:affiliation>
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<mods:namePart id="7310A4B30DA3CF64A5399D484FAF7A67">Mignani, Roberto</mods:namePart>
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<mods:affiliation id="AD6ED7969C7BF3D46D059D78473E73DA">Dipartimento di Fisica ‘‘ E. Amaldi’ ’ Università degli Studi Roma Tre Via della Vasca Navale 84 00146 Roma, ITALY</mods:affiliation>
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<mods:roleTerm id="18C2CA1936D801208C64DB7A82F44016">Author</mods:roleTerm>
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<mods:namePart id="210D3A9B28EADB165DC244C626E77D4A">Piattella, Emanuele</mods:namePart>
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<mods:affiliation id="EC324DE20D22F85F73D11DD955669A24">Dipartimento di Biologia Animale e dell’Uomo Università ‘‘ La Sapienza’ ’ Viale dell’Università 32 I 00185 Roma, ITALY</mods:affiliation>
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<mods:title id="BF87D879D121509F4D9B19E9B560EF6C">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="C7CD6A3C578E92FEBFC910709E421B98">
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<mods:date id="17D93FEF05DC0286C19790A1A6C85CEF">2000</mods:date>
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<mods:number id="4C3F8129378E4CC7A3D31A59A7536827">2000-06-01</mods:number>
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<mods:number id="C8DA2E22B29340F8284BD01A5EEE98AE">54</mods:number>
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<mods:number id="30E6047527E280C13915B5246E099EC8">2</mods:number>
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<mods:start id="C575CE951695A4D92C71BD57B2FDE57D">239</mods:start>
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<mods:end id="7CA4F17F78C4984C643F6596D7C2AC3A">247</mods:end>
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<mods:identifier id="97C32759405E4D8E84117BDBA136558D" type="DOI">10.1649/0010-065x(2000)054[0239:atootg]2.0.co;2</mods:identifier>
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<mods:identifier id="E1573462A3CF5D492B3BF8EF9DE47028" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="C36DF5DAD8588B57FEA215121D88F96A" pageId="0" pageNumber="239" type="nomenclature">
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<paragraph id="8BC8A651D8588B57FEA215121C83F9BA" blockId="0.[313,928,1637,1659]" box="[313,928,1637,1659]" pageId="0" pageNumber="239">
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<heading id="D080113DD8588B57FEA215121C83F9BA" box="[313,928,1637,1659]" centered="true" fontSize="9" level="2" pageId="0" pageNumber="239" reason="2">
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<taxonomicName id="4C77DDD2D8588B57FEA215121D79F9BA" authority="(Klug 1843)" baseAuthorityName="Klug" baseAuthorityYear="1843" box="[313,602,1637,1659]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="239" phylum="Arthropoda" rank="species" species="kordofanus">
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<emphasis id="B9037A43D8588B57FEA215121EECF9BB" box="[313,463,1638,1659]" italics="true" pageId="0" pageNumber="239">P. kordofanus</emphasis>
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(Klug 1843)
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</taxonomicName>
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(
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<collectingCountry id="F360E6C1D8588B57FD7A15111C0BF9BA" box="[737,808,1637,1658]" name="Sudan" pageId="0" pageNumber="239">Sudan</collectingCountry>
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: Kordofan)
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</heading>
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</paragraph>
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<paragraph id="8BC8A651D8588B57FD1315E31D88F96A" blockId="0.[648,683,1687,1706]" box="[648,683,1687,1706]" pageId="0" pageNumber="239">239</paragraph>
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</subSubSection>
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<subSubSection id="C36DF5DAD8598B56FEAA12621C3BFEEA" box="[305,792,277,299]" pageId="1" pageNumber="240" type="description">
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<paragraph id="8BC8A651D8598B56FEAA12621C3BFEEA" blockId="1.[305,1021,277,352]" box="[305,792,277,299]" pageId="1" pageNumber="240">
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<taxonomicName id="4C77DDD2D8598B56FEAA12621C3BFEEA" authority="(Castelnau 1840) (India)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[305,792,277,299]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="orientalis">
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<emphasis id="B9037A43D8598B56FEAA12621E96FEEB" box="[305,437,278,299]" italics="true" pageId="1" pageNumber="240">P. orientalis</emphasis>
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(Castelnau 1840) (
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<collectingCountry id="F360E6C1D8598B56FD4312611C32FEEA" box="[728,785,277,298]" name="India" pageId="1" pageNumber="240">India</collectingCountry>
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)
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DF5DAD8598B56FEAA12441CDBFDCD" pageId="1" pageNumber="240" type="reference_group">
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<paragraph id="8BC8A651D8598B56FEAA12441C8AFEA0" blockId="1.[305,1021,277,352]" pageId="1" pageNumber="240">
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<taxonomicName id="4C77DDD2D8598B56FEAA12441D49FE85" authority="(Castelnau 1840)" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[305,618,304,325]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="porcatus">
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<emphasis id="B9037A43D8598B56FEAA12441E8FFE85" box="[305,428,304,325]" italics="true" pageId="1" pageNumber="240">P. porcatus</emphasis>
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(Castelnau 1840)
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</taxonomicName>
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(Sahel:
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<collectingCountry id="F360E6C1D8598B56FCBF12441C54FE85" box="[804,887,304,325]" name="Senegal" pageId="1" pageNumber="240">Senegal</collectingCountry>
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to
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<collectingCountry id="F360E6C1D8598B56FC0612441CD5FE85" box="[925,1014,304,325]" name="Djibouti" pageId="1" pageNumber="240">Djibouti</collectingCountry>
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)
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<taxonomicName id="4C77DDD2D8598B56FEAA123F1DB2FEA0" authority="(Peringuey 1908)" baseAuthorityName="Peringuey" baseAuthorityYear="1908" box="[305,657,331,352]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="rhodesianus">
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<emphasis id="B9037A43D8598B56FEAA123F1EF2FEA0" box="[305,465,331,352]" italics="true" pageId="1" pageNumber="240">P. rhodesianus</emphasis>
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(
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<bibRefCitation id="EFE6DBA0D8598B56FE7B123F1DA8FEA0" author="Peringuey" box="[480,651,331,352]" pageId="1" pageNumber="240" pagination="1 - 752" refId="ref4025" refString="Peringuey, L. 1908. Descriptive catalogue of the Coleoptera of South Africa (Lucanidae and Scarabaeidae). Transactions of the South African Philosophical Society 13: 1 - 752." type="journal article" year="1908">Peringuey 1908</bibRefCitation>
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)
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</taxonomicName>
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(
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<collectingCountry id="F360E6C1D8598B56FD43123F1C6EFEA0" box="[728,845,331,352]" name="Zimbabwe" pageId="1" pageNumber="240">Zimbabwe</collectingCountry>
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:
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<collectingRegion id="49B368B3D8598B56FCC2123F1C81FEA0" box="[857,930,331,352]" country="Zimbabwe" name="Harare" pageId="1" pageNumber="240">Harare</collectingRegion>
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)
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</paragraph>
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<paragraph id="8BC8A651D8598B56FE9012071CDBFDCD" blockId="1.[241,1074,371,632]" pageId="1" pageNumber="240">
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Our present knowledge of this genus does not permit us to ascertain whether some of the above mentioned species are to be synonymized. According to
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<bibRefCitation id="EFE6DBA0D8598B56FF6A12DC1E98FE7D" author="Boucomont" box="[241,443,424,445]" pageId="1" pageNumber="240" pagination="333 - 350" refId="ref3757" refString="Boucomont, A. 1911. Contribution a la classification des Geotrypidae (Col.). Annales de la Societe Entomologique de France 79 (1910): 333 - 350." type="journal article" year="1911">Boucomont (1911)</bibRefCitation>
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all the species described up to that time were to be considered synonyms of
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<emphasis id="B9037A43D8598B56FE4A12B71D71FE18" box="[465,594,451,472]" italics="true" pageId="1" pageNumber="240">
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<taxonomicName id="4C77DDD2D8598B56FE4A12B71D6DFE18" baseAuthorityName="Castelnau" baseAuthorityYear="1840" box="[465,590,451,472]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="porcatus">P. porcatus</taxonomicName>
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.
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</emphasis>
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Nevertheless, we cannot agree with his statement, because many differences of specific rank occur among the specimens we examined from several African countries and the Arabian peninsula.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36DF5DAD8598B56FE9011671E8CFC77" pageId="1" pageNumber="240" type="description">
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<paragraph id="8BC8A651D8598B56FE9011671C35FDB8" blockId="1.[241,1074,371,632]" pageId="1" pageNumber="240">
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The present work originated from the examination of several specimens of
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<taxonomicName id="4C77DDD2D8598B56FF6A11591E67FD82" baseAuthorityName="Gomes Alves" baseAuthorityYear="1957" box="[241,324,557,578]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="freyi">
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<emphasis id="B9037A43D8598B56FF6A11591E67FD82" box="[241,324,557,578]" italics="true" pageId="1" pageNumber="240">P. freyi</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C77DDD2D8598B56FE1811591D04FD82" baseAuthorityName="Peringuey" baseAuthorityYear="1908" box="[387,551,557,578]" class="Insecta" family="Bolboceratidae" genus="Pseudoathyreus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="rhodesianus">
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<emphasis id="B9037A43D8598B56FE1811591D04FD82" box="[387,551,557,578]" italics="true" pageId="1" pageNumber="240">P. rhodesianus</emphasis>
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</taxonomicName>
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(including
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<typeStatus id="54CC18F3D8598B56FD3011591DFBFD82" box="[683,728,557,578]" pageId="1" pageNumber="240">type</typeStatus>
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specimens). The study of this material led us to new insights of the taxonomic and zoogeographic status of these two rare species within the genus as follows:
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</paragraph>
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<paragraph id="8BC8A651D8598B56FF6A11FE1DC0FD15" blockId="1.[241,1073,650,858]" pageId="1" pageNumber="240">
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(1) better definition of the taxonomic characters (both external and internal morphology) distinguishing the two species which were informally considered synonyms (
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<emphasis id="B9037A43D8598B56FE7611B41D0DFD15" box="[493,558,704,725]" italics="true" pageId="1" pageNumber="240">in litt.</emphasis>
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H. F. Howden).
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</paragraph>
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<paragraph id="8BC8A651D8598B56FF6A11AE1D51FCCA" blockId="1.[241,1073,650,858]" pageId="1" pageNumber="240">(2) deeper knowledge of their intraspecific variation, including clypeopronotal ornamentation and colour;</paragraph>
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<paragraph id="8BC8A651D8598B56FF6A10641C2EFCE5" blockId="1.[241,1073,650,858]" box="[241,781,784,805]" pageId="1" pageNumber="240">(3) enlargement of their known geographic range;</paragraph>
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<paragraph id="8BC8A651D8598B56FF6A105E1C9AFC9A" blockId="1.[241,1073,650,858]" pageId="1" pageNumber="240">(4) identification of a distinct phylogenetic unit formed by these two species which are distinct from the other representatives of the genus.</paragraph>
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<paragraph id="8BC8A651D8598B56FE9010191E8CFC77" blockId="1.[241,1073,877,951]" pageId="1" pageNumber="240">Identification of acronyms used for museums holding specimens studied are given in the Acknowledgments. Private collections are indicated by the owner’s name in brackets.</paragraph>
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</subSubSection>
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</treatment>
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