412 lines
54 KiB
XML
412 lines
54 KiB
XML
<document id="DA92887193453DA5711F20AB7D36AE2E" ID-DOI="10.5281/zenodo.11263268" ID-ISSN="1525-9153" ID-Zenodo-Dep="11263268" ID-ZooBank="D6FE38AE-C4AF-40BE-AD69-93F197FB91E1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1714486077319" checkinUser="felipe" docAuthor="Rodríguez, Lily O. & Mamani, Luis" docDate="2020" docId="03D387CFFF82866AFF7DA796FC99F817" docLanguage="en" docName="AmphibReptileCons.14.1.e227.pdf" docOrigin="Amphibian & Reptile Conservation (e 227) 14 (1)" docStyle="DocumentStyle:D97478CDCB68366E5D4AECD282BFFB6F.1:AmphibReptileConse.2015-.journal_article.0cover" docStyleId="D97478CDCB68366E5D4AECD282BFFB6F" docStyleName="AmphibReptileConse.2015-.journal_article.0cover" docStyleVersion="1" docTitle="Petracola pajatensis Rodríguez & Mamani 2020, sp. nov." docType="treatment" docVersion="2" lastPageNumber="144" masterDocId="FFEAFFB7FF83866EFFF0A36EFFF6FF8A" masterDocTitle="A new species of Petracola (Squamata: Gymnophthalmidae) from Río Abiseo National Park, San Martín, Peru" masterLastPageNumber="146" masterPageNumber="140" pageNumber="141" updateTime="1716476409225" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="323AD917020B3F6C6472B4ED9509B5F7">
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<mods:title id="C0490C23D354C75BA41289292939CEED">A new species of Petracola (Squamata: Gymnophthalmidae) from Río Abiseo National Park, San Martín, Peru</mods:title>
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</mods:titleInfo>
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<mods:name id="4CFA0BBCDF7FFF5E7CAB47CBEA64A9C4" type="personal">
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<mods:roleTerm id="AE13D719245F1C4E40D7B80CAC7F0990">Author</mods:roleTerm>
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<mods:namePart id="0C2DE13166748665F0E83D8745BCF12F">Rodríguez, Lily O.</mods:namePart>
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<mods:affiliation id="5EB9FC3EC29F714DFDCA889C9DAAD35B">Museo de Historia Natural, Universidad Nacional Mayor de San Marcos (MUSM), Lima, PERÚ</mods:affiliation>
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<mods:nameIdentifier id="18A867920906714B25B2211B635715D5" type="email">lrodriguez@cima.org.pe</mods:nameIdentifier>
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</mods:name>
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<mods:roleTerm id="53A7E62A3EE704EF7400580744D620D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CC29902B41B6BD99F9E0943B7526B9AA">Mamani, Luis</mods:namePart>
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<mods:affiliation id="443AB715527D759C067467297BE4301A">Museo de Biodiversidad del Perú (MUBI), Urbanización Mariscal Gamarra A- 61, Zona 2, Cusco, PERÚ & Museo de Historia Natural de la Universidad Nacional de San Antonio Abad del Cusco (MHNC), Plaza de Armas s / n (Paraninfo Universitario), Cusco, PERÚ & Programa de Magister en Ciencias con mención en Zoología, Departamento de Zoología, Facultad de Ciencias Naturales y Oceanográficas, Universidad de Concepción, Barrio Universitario S / N, Casilla 160 C, Concepción, CHILE</mods:affiliation>
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<mods:typeOfResource id="084FC06637738C360449389F38413EF0">text</mods:typeOfResource>
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<mods:title id="52694DF1E3F27E5313C77B2D14EDE041">Amphibian & Reptile Conservation</mods:title>
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</mods:titleInfo>
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<mods:part id="5C471C9E23BC29BB7F014D79766A54C2">
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<mods:date id="4C5F212979A40BE4ED060FD7D11C364D">2020</mods:date>
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<mods:title id="759DC0E208C7D3A4FF41D9A7A9C4477D">e 227</mods:title>
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</mods:detail>
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<mods:detail id="0955B1B85B0C33FB050672257C9C905E" type="pubDate">
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<mods:number id="2FDD2D60CD5340A9609AEFDCDD16F052">2020-03-28</mods:number>
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</mods:detail>
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<mods:detail id="37A8DC2286DB0F1D19FE2281F4C32E1B" type="volume">
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<mods:number id="A39829BCC9B07B0AA07DC4B5D5CED591">14</mods:number>
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<mods:detail id="4D12665F1352AB5A5C6AFB8B354CAC17" type="issue">
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<mods:number id="77A1419192BC0AC6E3AB6AAB0D6BDE80">1</mods:number>
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<mods:classification id="509D98519FF99F079ADB185DC92C45A6">journal article</mods:classification>
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<mods:identifier id="6B341EF925C780A934C27C673676A617" type="DOI">10.5281/zenodo.11263268</mods:identifier>
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<mods:identifier id="CA0866667A5164B15DCBAB6B06A6F6C2" type="ISSN">1525-9153</mods:identifier>
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<treatment id="03D387CFFF82866AFF7DA796FC99F817" ID-DOI="http://doi.org/10.5281/zenodo.11263280" ID-Zenodo-Dep="11263280" LSID="urn:lsid:plazi:treatment:03D387CFFF82866AFF7DA796FC99F817" httpUri="http://treatment.plazi.org/id/03D387CFFF82866AFF7DA796FC99F817" lastPageId="4" lastPageNumber="144" pageId="1" pageNumber="141">
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<subSubSection id="C3606552FF82866FFF7DA796FE31FA9A" box="[141,455,1272,1296]" pageId="1" pageNumber="141" type="nomenclature">
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<paragraph id="8BC536D9FF82866FFF7DA796FE31FA9A" blockId="1.[141,455,1272,1296]" box="[141,455,1272,1296]" pageId="1" pageNumber="141">
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<heading id="D08D81B5FF82866FFF7DA796FE31FA9A" box="[141,455,1272,1296]" fontSize="10" level="2" pageId="1" pageNumber="141" reason="4">
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<emphasis id="B90EEACBFF82866FFF7DA796FE31FA9A" bold="true" box="[141,455,1272,1296]" pageId="1" pageNumber="141">
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<taxonomicName id="4C7A4D5AFF82866FFF7DA796FE9DFA9A" ID-CoL="8H6PH" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[141,363,1272,1296]" class="Repitilia" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="141" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
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<emphasis id="B90EEACBFF82866FFF7DA796FE9DFA9A" bold="true" box="[141,363,1272,1296]" italics="true" pageId="1" pageNumber="141">Petracola pajatensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D57B0FF82866FFE81A796FE31FA9A" box="[369,455,1272,1296]" pageId="1" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3606552FF82866FFF7DA653FD12FAC4" box="[141,740,1341,1358]" pageId="1" pageNumber="141" type="description">
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<paragraph id="8BC536D9FF82866FFF7DA653FD12FAC4" blockId="1.[141,740,1341,1358]" box="[141,740,1341,1358]" pageId="1" pageNumber="141">
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<emphasis id="B90EEACBFF82866FFF7DA653FD12FAC4" bold="true" box="[141,740,1341,1358]" pageId="1" pageNumber="141">
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<uri id="FFEB3ADBFF82866FFF7DA653FD12FAC4" box="[141,740,1341,1358]" pageId="1" pageNumber="141">
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urn:lsid:zoobank.org:act:
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<uuid id="FFDC0C0CFF82866FFEAEA653FD12FAC4" box="[350,740,1341,1358]" pageId="1" pageNumber="141">E41CD69F-3F7A-41AF-B756-BF836A536EA8</uuid>
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</uri>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606552FF82866FFF7DA616FE76F93A" pageId="1" pageNumber="141" type="materials_examined">
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<paragraph id="8BC536D9FF82866FFF7DA616FF3CF9BA" blockId="1.[141,746,1400,1584]" pageId="1" pageNumber="141">
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<materialsCitation id="3B123C84FF82866FFF7DA616FF3CF9BA" collectingDate="1987-07-12" collectionCode="MUSM" collectorName="Lily O. Rodriguez" country="Peru" county="Departamento" elevation="3" latitude="-7.6369443" location="Los Chochos" longitude="-77.28" municipality="Provincia Mariscal Caceres" pageId="1" pageNumber="141" specimenCode="MUSM 3829" specimenCount="1" specimenCount-male="1" stateProvince="Departamento San Martin" typeStatus="holotype">
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<emphasis id="B90EEACBFF82866FFF7DA616FF0AFA1A" bold="true" box="[141,252,1400,1424]" pageId="1" pageNumber="141">
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<typeStatus id="54C1887BFF82866FFF7DA616FF01FA1A" box="[141,247,1400,1424]" pageId="1" pageNumber="141" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBDC9EA2FF82866FFEF1A616FE66FA1A" box="[257,400,1400,1424]" collectionCode="MUSM" pageId="1" pageNumber="141">MUSM 3829</specimenCode>
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, adult
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<specimenCount id="9D7CFD50FF82866FFE25A616FDFFFA1A" box="[469,521,1400,1424]" pageId="1" pageNumber="141" type="male">male</specimenCount>
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(
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<figureCitation id="13412A5CFF82866FFDE6A616FD8CFA1A" box="[534,634,1400,1424]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[141,176,808,830]" captionStartId-1="1.[791,826,1903,1925]" captionTargetBox-0="[153,1381,152,781]" captionTargetBox-1="[826,1363,878,1879]" captionTargetId-0="figure-187@1.[141,1394,139,799]" captionTargetId-1="figure-212@1.[791,1395,856,1898]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="Fig. 1. Holotype of Petracola pajatensis sp. nov. (MUSM 3829; male, SVL = 60.5, Tail = 76.8 mm, scale bar = 10 mm)." captionText-1="Fig. 2. Dorsal, lateral, and ventral views of head of the holotype (MUSM 3829)." figureDoi-0="http://doi.org/10.5281/zenodo.11263272" figureDoi-1="http://doi.org/10.5281/zenodo.11263274" httpUri-0="https://zenodo.org/record/11263272/files/figure.png" httpUri-1="https://zenodo.org/record/11263274/files/figure.png" pageId="1" pageNumber="141">Figs. 1–2</figureCitation>
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) from
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<location id="8EA56002FF82866FFD31A617FF1BFA3A" LSID="urn:lsid:plazi:treatment:03D387CFFF82866AFF7DA796FC99F817:8EA56002FF82866FFD31A617FF1BFA3A" country="Peru" county="Departamento" latitude="-7.6369443" municipality="Provincia Mariscal Caceres" name="Los Chochos" pageId="1" pageNumber="141" stateProvince="Departamento San Martin">Los Chochos</location>
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,
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<location id="8EA56002FF82866FFF09A6F6FDE4FA3A" LSID="urn:lsid:plazi:treatment:03D387CFFF82866AFF7DA796FC99F817:8EA56002FF82866FFF09A6F6FDE4FA3A" box="[249,530,1432,1456]" country="Peru" county="Departamento" latitude="-7.6369443" municipality="Provincia Mariscal Caceres" name="Rio Abiseo National Park" pageId="1" pageNumber="141" stateProvince="Departamento San Martin">Rio Abiseo National Park</location>
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||
,
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||
<collectingMunicipality id="6BA1ACA3FF82866FFDEEA6F6FF12FA5A" pageId="1" pageNumber="141">Provincia Mariscal Cáceres</collectingMunicipality>
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||
,
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||
<collectingRegion id="49BEF83BFF82866FFF1FA6D7FDF0FA5A" box="[239,518,1464,1488]" pageId="1" pageNumber="141">Departamento San Martín</collectingRegion>
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,
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||
<collectingCountry id="F36D7649FF82866FFDE1A6D7FDB3FA5A" box="[529,581,1465,1488]" name="Peru" pageId="1" pageNumber="141">Peru</collectingCountry>
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, approximately
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||
<elevation id="0057D1EAFF82866FFF7DA6B6FEEAFA7A" box="[141,284,1496,1520]" metricMagnitude="0" metricUnit="m" metricValue="3.23" pageId="1" pageNumber="141" unit="m" value="3.23">
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<quantity id="4C829B3CFF82866FFF7DA6B6FF1BFA7A" box="[141,237,1496,1520]" metricMagnitude="0" metricUnit="m" metricValue="3.23" pageId="1" pageNumber="141" unit="m" value="3.23">3,230 m</quantity>
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asl
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||
</elevation>
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,
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||
<quantity id="4C829B3CFF82866FFEDFA6B6FE8DFA7A" box="[303,379,1496,1520]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="141" unit="km" value="18.0">
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<locationDeviation id="9A772FFBFF82866FFEDFA6B6FE8DFA7A" box="[303,379,1496,1520]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="1" pageNumber="141" unit="km" value="18.0">18 km</locationDeviation>
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</quantity>
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airline from
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<collectingMunicipality id="6BA1ACA3FF82866FFDD0A6B7FDACFA7A" box="[544,602,1497,1520]" pageId="1" pageNumber="141">Pataz</collectingMunicipality>
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(
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<geoCoordinate id="EE4E501EFF82866FFD82A6B6FD12FA7A" box="[626,740,1496,1520]" degrees="7" direction="south" minutes="38" orientation="latitude" pageId="1" pageNumber="141" precision="15" seconds="13" value="-7.6369443">7°38’13”S</geoCoordinate>
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,
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||
<geoCoordinate id="EE4E501EFF82866FFF7DA696FEEFF99A" box="[141,281,1528,1552]" direction="west" orientation="longitude" pageId="1" pageNumber="141" value="-77.28">77°28’80”W</geoCoordinate>
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), collected by
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<collectorName id="268F530FFF82866FFE43A696FD83F99A" box="[435,629,1528,1552]" pageId="1" pageNumber="141">Lily O. Rodriguez</collectorName>
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on
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<date id="FFC41019FF82866FFD6CA696FF30F9BA" pageId="1" pageNumber="141" value="1987-07-12">
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<collectingDate id="EF80E9F1FF82866FFD6CA696FF30F9BA" pageId="1" pageNumber="141" value="1987-07-12">12 July 1987</collectingDate>
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</date>
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||
.
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BC536D9FF82866FFF7DA536FE76F93A" blockId="1.[141,745,1624,1712]" pageId="1" pageNumber="141">
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||
<materialsCitation id="3B123C84FF82866FFF7DA536FEC8F91A" collectingDate="1987-07-12" collectionCode="MUSM" collectorName="Lily O. Rodriguez" country="Peru" county="Departamento" elevation="3" latitude="-7.6369443" location="Los Chochos" municipality="Provincia Mariscal Caceres" pageId="1" pageNumber="141" specimenCode="MUSM 3830" specimenCount="3" specimenCount-female="3" stateProvince="Departamento San Martin" typeStatus="paratype">
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||
<emphasis id="B90EEACBFF82866FFF7DA536FEF0F9FA" bold="true" box="[141,262,1624,1648]" pageId="1" pageNumber="141">
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||
<typeStatus id="54C1887BFF82866FFF7DA536FEF7F9FA" box="[141,257,1624,1648]" pageId="1" pageNumber="141" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
<specimenCount id="9D7CFD50FF82866FFEFEA536FE10F9FA" box="[270,486,1624,1648]" pageId="1" pageNumber="141">Three adult females</specimenCount>
|
||
(
|
||
<specimenCode id="DBDC9EA2FF82866FFE06A536FD71F9FA" box="[502,647,1624,1648]" collectionCode="MUSM" pageId="1" pageNumber="141">MUSM 3830</specimenCode>
|
||
[
|
||
<figureCitation id="13412A5CFF82866FFD67A536FD2AF9FA" box="[663,732,1624,1648]" captionStart="Fig" captionStartId="1.[791,826,1903,1925]" captionTargetBox="[826,1363,878,1879]" captionTargetId="figure-212@1.[791,1395,856,1898]" captionTargetPageId="1" captionText="Fig. 2. Dorsal, lateral, and ventral views of head of the holotype (MUSM 3829)." figureDoi="http://doi.org/10.5281/zenodo.11263274" httpUri="https://zenodo.org/record/11263274/files/figure.png" pageId="1" pageNumber="141">Fig. 2</figureCitation>
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||
], 15986–87), and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B123C84FF82866FFEBCA517FE8AF93A" collectingDate="1987-07-12" collectionCode="MUSM" collectorName="Lily O. Rodriguez" country="Peru" county="Departamento" elevation="3" latitude="-7.6369443" location="Los Chochos" municipality="Provincia Mariscal Caceres" pageId="1" pageNumber="141" specimenCode="MUSM 15985" specimenCount="1" specimenCount-male="1" stateProvince="San Martin" typeStatus="paratype">
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||
<specimenCount id="9D7CFD50FF82866FFEBCA517FDE9F91A" box="[332,543,1656,1680]" pageId="1" pageNumber="141">one subadult male</specimenCount>
|
||
(
|
||
<specimenCode id="DBDC9EA2FF82866FFDC5A516FD2BF91A" box="[565,733,1656,1680]" collectionCode="MUSM" pageId="1" pageNumber="141">MUSM 15985</specimenCode>
|
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), same data as holotype
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</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606552FF82866CFF7DA5B6FA87FA88" lastPageId="2" lastPageNumber="142" pageId="1" pageNumber="141" type="diagnosis">
|
||
<paragraph id="8BC536D9FF82866CFF7DA5B6FE8FFA28" blockId="1.[141,746,1752,1968]" lastBlockId="2.[141,746,906,1986]" lastPageId="2" lastPageNumber="142" pageId="1" pageNumber="141">
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||
<emphasis id="B90EEACBFF82866FFF7DA5B6FF0DF97A" bold="true" box="[141,251,1752,1776]" pageId="1" pageNumber="141">Diagnosis</emphasis>
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. (1) Frontonasal and frontal scales sub-equal; (2) nasoloreal suture present, loreal scale not in contact with supralabials; (3) supraoculars two; (4) superciliaries two, discontinuous, first expanded onto dorsal surface of head; (5) postoculars two; (6) palpebral disc divided in two; (7) three supralabials anterior to the posteroventral angle of subocular; (8) two pairs of genials in contact; (9) dorsal body scales quadrangular, smooth, juxtaposed; (10) transverse dorsal rows 33–35; (11) transversal ventral rows 20–22; (12) a continuous series of small lateral scales separates dorsals from ventrals; (13) posterior cloacal plate scales 2–5; (14) anterior preanal plate scales paired; (15) femoral pores per hind limb in males 6–8, in females three; (16) preanal pores absent; (17) subdigital lamellae on finger IV eight; (18) limbs not overlapping when adpressed against body on adults; (19) pentadactyl, digits clawed; (20) coloration in liquid preservative: in males, dorsum is light-brown with numerous irregular dark-brown spots and venter is dark-brown with some small cream spots on their flanks; in females, dorsum is light-brown with some and irregular dark-brown spots, venter is brown with cream spots that form discontinuous transversal bands from the chest to the anal plate (
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<figureCitation id="13412A5CFF81866CFEDAA6E4FE9DFA28" box="[298,363,1418,1442]" captionStart="Fig" captionStartId="2.[141,175,848,870]" captionTargetBox="[156,1382,145,836]" captionTargetId="figure-558@2.[141,1394,126,846]" captionTargetPageId="2" captionText="Fig. 3. Dorsal and ventral views of the paratype of Petracola pajatensis sp. nov. (MUSM 3830; female, SVL = 65.1, Tail = 64.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.11263276" httpUri="https://zenodo.org/record/11263276/files/figure.png" pageId="2" pageNumber="142">Fig. 3</figureCitation>
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).
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</paragraph>
|
||
<caption id="DF056651FF82866FFCE7A401FC5BF829" ID-DOI="http://doi.org/10.5281/zenodo.11263274" ID-Zenodo-Dep="11263274" httpUri="https://zenodo.org/record/11263274/files/figure.png" pageId="1" pageNumber="141" startId="1.[791,826,1903,1925]" targetBox="[826,1363,878,1879]" targetPageId="1" targetType="figure">
|
||
<paragraph id="8BC536D9FF82866FFCE7A401FC5BF829" blockId="1.[791,1395,1903,1955]" pageId="1" pageNumber="141">
|
||
<emphasis id="B90EEACBFF82866FFCE7A401FCA3F80E" bold="true" box="[791,853,1903,1925]" pageId="1" pageNumber="141">Fig. 2.</emphasis>
|
||
Dorsal, lateral, and ventral views of head of the holotype (MUSM 3829).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF056651FF81866CFF7DA03EFA82FCEC" ID-DOI="http://doi.org/10.5281/zenodo.11263276" ID-Zenodo-Dep="11263276" box="[141,1396,848,870]" httpUri="https://zenodo.org/record/11263276/files/figure.png" pageId="2" pageNumber="142" startId="2.[141,175,848,870]" targetBox="[156,1382,145,836]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BC536D9FF81866CFF7DA03EFA82FCEC" blockId="2.[141,1396,848,870]" box="[141,1396,848,870]" pageId="2" pageNumber="142">
|
||
<emphasis id="B90EEACBFF81866CFF7DA03EFF3DFCEF" bold="true" box="[141,203,848,870]" pageId="2" pageNumber="142">Fig. 3.</emphasis>
|
||
Dorsal and ventral views of the paratype of
|
||
<taxonomicName id="4C7A4D5AFF81866CFD9AA03EFCDDFCEC" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[618,811,848,870]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF81866CFD9AA03EFCDDFCEC" box="[618,811,848,870]" italics="true" pageId="2" pageNumber="142">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF81866CFCC0A03EFC8DFCEF" bold="true" box="[816,891,848,869]" pageId="2" pageNumber="142">
|
||
<taxonomicNameLabel id="A23D57B0FF81866CFCC0A03EFC8DFCEF" box="[816,891,848,869]" pageId="2" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(MUSM 3830; female, SVL = 65.1, Tail = 64.3 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC536D9FF81866CFF5DA6C4FA87FA88" blockId="2.[141,746,906,1986]" lastBlockId="2.[791,1395,906,1282]" pageId="2" pageNumber="142">
|
||
<taxonomicName id="4C7A4D5AFF81866CFF5DA6C4FE79FA4B" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[173,399,1450,1474]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF81866CFF5DA6C4FE79FA4B" box="[173,399,1450,1474]" italics="true" pageId="2" pageNumber="142">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF81866CFE6CA6C4FE01FA48" bold="true" box="[412,503,1450,1474]" pageId="2" pageNumber="142">
|
||
<taxonomicNameLabel id="A23D57B0FF81866CFE6CA6C4FE01FA48" box="[412,503,1450,1474]" pageId="2" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName id="4C7A4D5AFF81866CFF20A6A4FE8CFA6B" authorityName="Echevarria and Venegas" authorityYear="2015" box="[208,378,1482,1505]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="angustisoma">
|
||
<emphasis id="B90EEACBFF81866CFF20A6A4FE8CFA6B" box="[208,378,1482,1505]" italics="true" pageId="2" pageNumber="142">P. angustisoma</emphasis>
|
||
</taxonomicName>
|
||
by having a robust body, two discontinuous superciliaries, 6–8 femoral pores per hind limb in males, maximum
|
||
<collectionCode id="ED6BAE1CFF81866CFDF5A564FDC0F9A8" box="[517,566,1546,1570]" pageId="2" pageNumber="142">SVL</collectionCode>
|
||
in males 60.5 mm, dorsum is light-brown with irregular dark brown spots not forming longitudinal stripes, venter is brown in preservative (gracile body, three discontinuous superciliaries, nine femoral pores per hind limb in males, maximum
|
||
<collectionCode id="ED6BAE1CFF81866CFEF2A5C4FEC5F948" box="[258,307,1706,1730]" pageId="2" pageNumber="142">SVL</collectionCode>
|
||
in males 43.6 mm, dorsum is brown or olive with seven discontinuous dark brown longitudinal stripes, venter is white with black semicircular black spots on anterior margin of scales); from
|
||
<taxonomicName id="4C7A4D5AFF81866CFDB1A464FD1FF8A8" baseAuthorityName="Kohler and Lehr" baseAuthorityYear="2004" box="[577,745,1802,1826]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="labioocularis">
|
||
<emphasis id="B90EEACBFF81866CFDB1A464FD1FF8A8" box="[577,745,1802,1826]" italics="true" pageId="2" pageNumber="142">P. labioocularis</emphasis>
|
||
</taxonomicName>
|
||
by having two supraoculars, absence of precloacal pores, and two pairs of genials in contact (three supraoculars, presence of precloacal pores, and usually three pairs of genials in contact); from
|
||
<taxonomicName id="4C7A4D5AFF81866CFE58A4E4FD84F828" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[424,626,1930,1954]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="ventrimaculatus">
|
||
<emphasis id="B90EEACBFF81866CFE58A4E4FD84F828" box="[424,626,1930,1954]" italics="true" pageId="2" pageNumber="142">P. ventrimaculatus</emphasis>
|
||
</taxonomicName>
|
||
by having 6–8 femoral pores in males, in preservative the venter in males is dark brown with small lateral cream spots, in females it is a combination of cream with brown forming longitudinal bands, and maximum
|
||
<collectionCode id="ED6BAE1CFF81866CFB7BA0A4FB4AFC68" box="[1163,1212,970,994]" pageId="2" pageNumber="142">SVL</collectionCode>
|
||
in males
|
||
<quantity id="4C829B3CFF81866CFAD7A0A4FA84FC6B" box="[1319,1394,970,994]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="142" unit="mm" value="60.0">60 mm</quantity>
|
||
(2–5 femoral pores in males, in males and females the venter is cream with a bold black transversal band, and maximum
|
||
<collectionCode id="ED6BAE1CFF81866CFC7CA744FC4BFBC8" box="[908,957,1066,1090]" pageId="2" pageNumber="142">SVL</collectionCode>
|
||
in males 71.05 mm); from
|
||
<taxonomicName id="4C7A4D5AFF81866CFB08A744FABBFBC8" authorityName="Kizirian, Bayefsky-Anand, Eriksson, Le, and Donnelly" authorityYear="2008" box="[1272,1357,1066,1090]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="2" pageNumber="142" phylum="Chordata" rank="species" species="waka">
|
||
<emphasis id="B90EEACBFF81866CFB08A744FABBFBC8" box="[1272,1357,1066,1090]" italics="true" pageId="2" pageNumber="142">P. waka</emphasis>
|
||
</taxonomicName>
|
||
by having two discontinuous superciliaries, two genials in contact, and venter in males is a dark brown with lateral cream spots, in females the venter is a combination of cream with brown forming a longitudinal band (four continuous superciliaries, three genials in contact, and the venter in males is white to pale yellow with brown spots).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606552FF81866AFCE7A647FED9F9B7" lastPageId="4" lastPageNumber="144" pageId="2" pageNumber="142" type="description">
|
||
<paragraph id="8BC536D9FF81866DFCE7A647FC6AFB47" blockId="2.[791,1395,1321,1986]" lastBlockId="3.[791,1395,726,1230]" lastPageId="3" lastPageNumber="143" pageId="2" pageNumber="142">
|
||
<emphasis id="B90EEACBFF81866CFCE7A647FBA8FAC8" bold="true" box="[791,1118,1321,1346]" pageId="2" pageNumber="142">
|
||
Description of the
|
||
<typeStatus id="54C1887BFF81866CFC0CA644FBA8FAC8" box="[1020,1118,1322,1346]" pageId="2" pageNumber="142" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Adult male, snout-vent length (
|
||
<collectionCode id="ED6BAE1CFF81866CFC81A624FC50FAE8" box="[881,934,1354,1378]" pageId="2" pageNumber="142">SVL</collectionCode>
|
||
) 60.5 mm, tail length 76.8 mm, head scales smooth, rounded in lateral and ventral views, without striations or rugosities; rostral scale wider than tall, in contact with frontonasal, nasals, and first supralabials; frontonasal longer that wide, longer that the frontal scale, widest posteriorly, in contact with rostral, nasal, first superciliary, and frontal; prefrontal absent; frontal longer that wide, pentagonal, in contact with first superciliary, first supraocular, and frontoparietal; frontoparietal paired, polygonal (hexagonal), in contact with frontal, supraoculars, parietals, and interparietals; supraoculars two, the first separates the first and second superciliaries, in contact with superciliaries, frontal, frontoparietals, interparietal, and postoculars; parietals longer than wide, polygonal (irregular hexagon), in contact with frontoparietals and supraoculars anteriorly, with interparietal, and temporals laterally, and with postparietals posteriorly; interparietal polygonal (hexagonal), in contact with frontoparietals anteriorly, with parietals laterally, and with postparietal posteriorly; postparietals paired, smaller than parietals, and polygonal. Nasal scale divided, longer than high, in contact with first supralabials; loreal scale present on the left side, nasoloreal suture incomplete on the right side, not in contact with the supralabials; two superciliaries, discontinuous, and first expanded onto the dorsal surface of the head; two preoculars; frenocular fused with the first subocular only on the right side; palpebral disc transparent and divided in two; three suboculars, on the right side the first subocular is fused with the frenocular; two postoculars; temporals smooth, polygonal; three supralabials anterior to posteroventral angle of suboculars. Mental wider than long, in contact with first infralabials and postmental posteriorly; postmental single, polygonal (irregular heptagonal), in contact with the first and second infralabials, and the first pair of genials; 3/2 genials, on right and left sides respectively, all in contact medially, the first genial on right side is divided; three transversal rows of pregular scales; six gular scale rows, polygonal, and smooth. Dorsal scales rectangular, longer that wide, juxtaposed, smooth, 33 transverse rows; 21 longitudinal dorsal scale rows at midbody; a continuous series of small lateral scales; reduced scales at limb insertion region present; 21 transverse ventral scale rows; 10 longitudinal ventral scale rows at midbody, the lateral scales are slightly smaller; anterior preanal plate scales paired; three posterior preanal plate scales, and a small and polygonal scale lies between the anterior and posterior preanal plate scales; scales on the tail rectangular, juxtaposed, and smooth; ventral scales quadrangular, juxtaposed, and smooth. Limbs pentadactyl; digits clawed; dorsal brachial scales polygonal, subimbricate, and smooth; ventral brachial scales rounded, subimbricate, and smooth; dorsal antebrachial scales polygonal, smooth; ventral antebrachial scales polygonal, smooth, smaller than dorsal; dorsal manus scales polygonal, smooth, subimbricate; palmar scales small, rounded, juxtaposed, and domelike; dorsal scales on fingers smooth, quadrangular, imbricate, three on finger I, five on II, six on III, five on IV, and four on
|
||
<collectionCode id="ED6BAE1CFF80866DFBE2A198FBD1FC87" box="[1042,1063,758,781]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="143" type="Museum">V</collectionCode>
|
||
; scales on anterodorsal surface of thigh polygonal, smooth, subimbricate; scales on posterior surface of thigh small, rounded, and juxtaposed; scales on ventral surface of thigh small, polygonal and juxtaposed; six femoral pores on left thigh and seven on right; scales on anterior surface of crus polygonal, smooth, juxtaposed, decreasing in size distally; scales on posterodorsal surface of crus smooth, polygonal, juxtaposed; scales on ventral surface of crus polygonal, enlarged, smooth, and subimbricate; scales on dorsal surface of toes polygonal, smooth, and subimbricate; scales on ventral surface of toes rounded, small, and domelike; dorsal scales of toes smooth, imbricate, two on toe I, five on toe II, seven on toe III, ten on toe IV, and six on toe
|
||
<collectionCode id="ED6BAE1CFF80866DFC75A7D8FC6EFB47" box="[901,920,1206,1229]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="143" type="Museum">V</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF056651FF80866DFF7DA1C5FBBAFD4B" ID-DOI="http://doi.org/10.5281/zenodo.11263278" ID-Zenodo-Dep="11263278" box="[141,1100,683,705]" httpUri="https://zenodo.org/record/11263278/files/figure.png" pageId="3" pageNumber="143" startId="3.[141,175,683,705]" targetBox="[141,1394,141,670]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BC536D9FF80866DFF7DA1C5FBBAFD4B" blockId="3.[141,1100,683,705]" box="[141,1100,683,705]" pageId="3" pageNumber="143">
|
||
<emphasis id="B90EEACBFF80866DFF7DA1C5FF3DFD4A" bold="true" box="[141,203,683,705]" pageId="3" pageNumber="143">Fig. 4.</emphasis>
|
||
Dorsolateral view of unvouchered living specimen from Los Chochos. Photo by Ken R. Young.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC536D9FF80866DFCE7A79BFC64FA87" blockId="3.[791,914,1269,1293]" box="[791,914,1269,1293]" pageId="3" pageNumber="143">
|
||
<heading id="D08D81B5FF80866DFCE7A79BFC64FA87" bold="true" box="[791,914,1269,1293]" fontSize="10" level="2" pageId="3" pageNumber="143" reason="2">
|
||
<emphasis id="B90EEACBFF80866DFCE7A79BFC64FA87" bold="true" box="[791,914,1269,1293]" pageId="3" pageNumber="143">Coloration</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC536D9FF80866DFCE7A658FAA3F884" blockId="3.[791,1395,1333,1806]" pageId="3" pageNumber="143">
|
||
<emphasis id="B90EEACBFF80866DFCE7A658FC3AFAC7" bold="true" box="[791,972,1333,1358]" pageId="3" pageNumber="143">In preservative.</emphasis>
|
||
<taxonomicName id="4C7A4D5AFF80866DFC25A658FB42FAC7" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[981,1204,1334,1358]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="3" pageNumber="143" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF80866DFC25A658FB42FAC7" box="[981,1204,1334,1358]" italics="true" pageId="3" pageNumber="143">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF80866DFB4DA658FAE3FAC4" bold="true" box="[1213,1301,1334,1358]" pageId="3" pageNumber="143">
|
||
<taxonomicNameLabel id="A23D57B0FF80866DFB4DA658FAE3FAC4" box="[1213,1301,1334,1358]" pageId="3" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
exhibits a variable coloration in adults of both sexes. The males (
|
||
<figureCitation id="13412A5CFF80866DFCEFA618FCA9FA04" box="[799,863,1398,1422]" captionStart="Fig" captionStartId="1.[141,176,808,830]" captionTargetBox="[153,1381,152,781]" captionTargetId="figure-187@1.[141,1394,139,799]" captionTargetPageId="1" captionText="Fig. 1. Holotype of Petracola pajatensis sp. nov. (MUSM 3829; male, SVL = 60.5, Tail = 76.8 mm, scale bar = 10 mm)." figureDoi="http://doi.org/10.5281/zenodo.11263272" httpUri="https://zenodo.org/record/11263272/files/figure.png" pageId="3" pageNumber="143">Fig. 1</figureCitation>
|
||
) have a light-brown dorsum with numerous and irregular dark-brown spots, with a cream continuous dorsolateral line on both sides of the body that starts from the back of the eye to the tail. This cream line is bordered on both sides by a continuous dark-brown line; the venter is dark-brown with some small cream spots on their flanks. The females (
|
||
<figureCitation id="13412A5CFF80866DFBA5A558FB63F9C4" box="[1109,1173,1590,1614]" captionStart="Fig" captionStartId="2.[141,175,848,870]" captionTargetBox="[156,1382,145,836]" captionTargetId="figure-558@2.[141,1394,126,846]" captionTargetPageId="2" captionText="Fig. 3. Dorsal and ventral views of the paratype of Petracola pajatensis sp. nov. (MUSM 3830; female, SVL = 65.1, Tail = 64.3 mm)." figureDoi="http://doi.org/10.5281/zenodo.11263276" httpUri="https://zenodo.org/record/11263276/files/figure.png" pageId="3" pageNumber="143">Fig. 3</figureCitation>
|
||
) have a light-brown dorsum with some irregular dark-brown spots, with a cream discontinuous dorsolateral line on both sides of the body that starts from the back of the eye to the tail; the venter is brown with cream spots that form continuous longitudinal bands from the chest to the anal plate; the neck and chin are a combination of cream and brown.
|
||
</paragraph>
|
||
<paragraph id="8BC536D9FF80866DFCE7A458FAA6F824" blockId="3.[791,1395,1845,1966]" pageId="3" pageNumber="143">
|
||
<emphasis id="B90EEACBFF80866DFCE7A458FC9EF8C7" bold="true" box="[791,872,1845,1870]" pageId="3" pageNumber="143">In life.</emphasis>
|
||
Based on unvouchered specimens, the dorsal coloration is brown with irregular dark spots distributed irregularly, the dorsolateral lines are obscure, and the flanks have more dark spots than the dorsum (
|
||
<figureCitation id="13412A5CFF80866DFAF2A4F8FAB5F824" box="[1282,1347,1942,1966]" captionStart="Fig" captionStartId="3.[141,175,683,705]" captionTargetBox="[141,1394,141,670]" captionTargetId="figure-637@3.[141,1394,139,674]" captionTargetPageId="3" captionText="Fig. 4. Dorsolateral view of unvouchered living specimen from Los Chochos. Photo by Ken R. Young." figureDoi="http://doi.org/10.5281/zenodo.11263278" httpUri="https://zenodo.org/record/11263278/files/figure.png" pageId="3" pageNumber="143">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF056651FF87866AFCE7A130FC6BFD1B" ID-DOI="http://doi.org/10.5281/zenodo.11263282" ID-Zenodo-Dep="11263282" httpUri="https://zenodo.org/record/11263282/files/figure.png" pageId="4" pageNumber="144" startId="4.[791,825,606,628]" targetBox="[791,1395,144,597]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BC536D9FF87866AFCE7A130FC6BFD1B" blockId="4.[791,1395,606,657]" pageId="4" pageNumber="144">
|
||
<emphasis id="B90EEACBFF87866AFCE7A130FCA1FDF9" bold="true" box="[791,855,606,628]" pageId="4" pageNumber="144">Fig. 6.</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="4C7A4D5AFF87866AFC0DA130FB36FDFE" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[1021,1216,606,628]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF87866AFC0DA130FB36FDFE" box="[1021,1216,606,628]" italics="true" pageId="4" pageNumber="144">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF87866AFB38A131FAE3FDFE" bold="true" box="[1224,1301,607,628]" pageId="4" pageNumber="144">
|
||
<taxonomicNameLabel id="A23D57B0FF87866AFB38A131FAE3FDFE" box="[1224,1301,607,628]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Photo by Ken R. Young.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF056651FF87866AFF7EA0A6FEE4FB92" ID-DOI="http://doi.org/10.5281/zenodo.11263284" ID-Zenodo-Dep="11263284" httpUri="https://zenodo.org/record/11263284/files/figure.png" pageId="4" pageNumber="144" startId="4.[142,176,968,990]" targetBox="[198,695,170,924]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BC536D9FF87866AFF7EA0A6FEE4FB92" blockId="4.[141,746,968,1048]" pageId="4" pageNumber="144">
|
||
<emphasis id="B90EEACBFF87866AFF7EA0A6FF3DFC57" bold="true" box="[142,203,968,990]" pageId="4" pageNumber="144">Fig. 5.</emphasis>
|
||
Drawings of the right side showing the condition of loreal scales in
|
||
<taxonomicName id="4C7A4D5AFF87866AFF1EA08BFE43FC71" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[238,437,997,1019]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF87866AFF1EA08BFE43FC71" box="[238,437,997,1019]" italics="true" pageId="4" pageNumber="144">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF87866AFE30A08BFDE6FC70" bold="true" box="[448,528,997,1018]" pageId="4" pageNumber="144">
|
||
<taxonomicNameLabel id="A23D57B0FF87866AFE30A08BFDE6FC70" box="[448,528,997,1018]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A) MUSM 3829, B) MUSM 3830.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC536D9FF87866AFF7DA72BFF0EFBD7" blockId="4.[141,248,1093,1117]" box="[141,248,1093,1117]" pageId="4" pageNumber="144">
|
||
<heading id="D08D81B5FF87866AFF7DA72BFF0EFBD7" bold="true" box="[141,248,1093,1117]" fontSize="10" level="2" pageId="4" pageNumber="144" reason="2">
|
||
<emphasis id="B90EEACBFF87866AFF7DA72BFF0EFBD7" bold="true" box="[141,248,1093,1117]" pageId="4" pageNumber="144">Variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC536D9FF87866AFF7DA7EBFED9F9B7" blockId="4.[141,746,1157,1597]" pageId="4" pageNumber="144">
|
||
Considerable variation is evident in the position and form of the loreal scale. The loreal scale can be incomplete, tiny, or enlarged; and can be separating, or not, the first superciliary and nasal scale; in the
|
||
<typeStatus id="54C1887BFF87866AFD7CA78BFD1FFB77" box="[652,745,1253,1277]" pageId="4" pageNumber="144" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="ED6BAE1CFF87866AFF64A66BFF11FA97" box="[148,231,1285,1309]" pageId="4" pageNumber="144">MUSM</collectionCode>
|
||
3829) the right scale is incomplete and the left does not separate the nasal and first superciliary, while in
|
||
<specimenCount id="9D7CFD50FF87866AFF5EA62BFEA3FAD7" box="[174,341,1349,1373]" count="3" pageId="4" pageNumber="144" type="generic" typeStatus="paratype">three paratypes</specimenCount>
|
||
(
|
||
<collectionCode id="ED6BAE1CFF87866AFE98A62BFE4DFAD7" box="[360,443,1349,1373]" pageId="4" pageNumber="144">MUSM</collectionCode>
|
||
3830, 15985–6) the loreal scale separated the nasal and first superciliary; and in
|
||
<specimenCount id="9D7CFD50FF87866AFF7DA6EBFEECFA17" box="[141,282,1413,1437]" count="1" pageId="4" pageNumber="144" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
||
(
|
||
<collectionCode id="ED6BAE1CFF87866AFEDBA6EBFE88FA17" box="[299,382,1413,1437]" pageId="4" pageNumber="144">MUSM</collectionCode>
|
||
15987) the first superciliary and nasal are in contact on both sides (
|
||
<figureCitation id="13412A5CFF87866AFDF8A6CBFDA3FA37" box="[520,597,1445,1469]" captionStart="Fig" captionStartId="1.[791,826,1903,1925]" captionTargetBox="[826,1363,878,1879]" captionTargetId="figure-212@1.[791,1395,856,1898]" captionTargetPageId="1" captionText="Fig. 2. Dorsal, lateral, and ventral views of head of the holotype (MUSM 3829)." figureDoi="http://doi.org/10.5281/zenodo.11263274" httpUri="https://zenodo.org/record/11263274/files/figure.png" pageId="4" pageNumber="144">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13412A5CFF87866AFD93A6CBFD84FA37" box="[611,626,1445,1469]" captionStart="Fig" captionStartId="4.[142,176,968,990]" captionTargetBox="[198,695,170,924]" captionTargetId="figure-274@4.[141,745,144,950]" captionTargetPageId="4" captionText="Fig. 5. Drawings of the right side showing the condition of loreal scales in Petracola pajatensis sp. nov. A) MUSM 3829, B) MUSM 3830." figureDoi="http://doi.org/10.5281/zenodo.11263284" httpUri="https://zenodo.org/record/11263284/files/figure.png" pageId="4" pageNumber="144">5</figureCitation>
|
||
). Variation in meristic characters is shown in
|
||
<tableCitation id="C6F80362FF87866AFDF2A6ABFDA5FA57" box="[514,595,1477,1501]" captionStart="Table 1" captionStartId="5.[141,197,142,163]" captionTargetBox="[153,1366,216,1294]" captionTargetId="graphics-275@5.[141,1394,205,1300]" captionTargetPageId="5" captionText="Table 1. Measurements and scale counts of specimens of Petracola pajatensis sp. nov. and P. ventrimaculatus examined in this paper. *only an adult male, ** the first right genial is divided, ***the anterior subocular scale is fused with freno ocular." httpUri="http://table.plazi.org/id/DF056651FF86866BFF7DA3E0FAF8FF4B" pageId="4" pageNumber="144" tableUuid="DF056651FF86866BFF7DA3E0FAF8FF4B">Table 1</tableCitation>
|
||
. No evidence of sexual dimorphism exists in the scutellation, except for the larger number of and better developed femoral pores in males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF056651FF87866AFCE7A6E6FAC9F99E" ID-DOI="http://doi.org/10.5281/zenodo.11263286" ID-Zenodo-Dep="11263286" httpUri="https://zenodo.org/record/11263286/files/figure.png" pageId="4" pageNumber="144" startId="4.[791,825,1416,1438]" targetBox="[794,1391,700,1396]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BC536D9FF87866AFCE7A6E6FAC9F99E" blockId="4.[791,1395,1416,1556]" pageId="4" pageNumber="144">
|
||
<emphasis id="B90EEACBFF87866AFCE7A6E6FCA2FA14" bold="true" box="[791,852,1416,1438]" pageId="4" pageNumber="144">Fig. 7.</emphasis>
|
||
Locations of the type localities of the species of
|
||
<taxonomicName id="4C7A4D5AFF87866AFAE6A6E7FA85FA14" authorityName="Doan and Castoe" authorityYear="2005" box="[1302,1395,1417,1438]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90EEACBFF87866AFAE6A6E7FA85FA14" box="[1302,1395,1417,1438]" italics="true" pageId="4" pageNumber="144">Petracola</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C7A4D5AFF87866AFCEEA6C9FC59FA36" authorityName="Echevarria and Venegas" authorityYear="2015" box="[798,943,1447,1468]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="angustisoma">
|
||
<emphasis id="B90EEACBFF87866AFCEEA6C9FC59FA36" box="[798,943,1447,1468]" italics="true" pageId="4" pageNumber="144">P. angustisoma</emphasis>
|
||
</taxonomicName>
|
||
= circle,
|
||
<taxonomicName id="4C7A4D5AFF87866AFBFEA6C9FB48FA31" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[1038,1214,1446,1468]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="ventrimaculatus">
|
||
<emphasis id="B90EEACBFF87866AFBFEA6C9FB48FA31" box="[1038,1214,1446,1468]" italics="true" pageId="4" pageNumber="144">P. ventrimaculatus</emphasis>
|
||
</taxonomicName>
|
||
= square,
|
||
<taxonomicName id="4C7A4D5AFF87866AFAD7A6C9FA85FA31" authorityName="Kizirian, Bayefsky-Anand, Eriksson, Le, and Donnelly" authorityYear="2008" box="[1319,1395,1446,1468]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="waka">
|
||
<emphasis id="B90EEACBFF87866AFAD7A6C9FA85FA31" box="[1319,1395,1446,1468]" italics="true" pageId="4" pageNumber="144">P. waka</emphasis>
|
||
</taxonomicName>
|
||
= diamond,
|
||
<taxonomicName id="4C7A4D5AFF87866AFC60A6AAFBDEFA52" baseAuthorityName="Kohler and Lehr" baseAuthorityYear="2004" box="[912,1064,1475,1497]" class="Squamata" family="Gymnophthalmidae" genus="Proctoporus" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="labioocularis">
|
||
<emphasis id="B90EEACBFF87866AFC60A6AAFBDEFA52" box="[912,1064,1475,1497]" italics="true" pageId="4" pageNumber="144">P. labioocularis</emphasis>
|
||
</taxonomicName>
|
||
= triangle,
|
||
<taxonomicName id="4C7A4D5AFF87866AFB50A6AAFAECFA53" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[1184,1306,1476,1497]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF87866AFB50A6AAFAECFA53" box="[1184,1306,1476,1497]" italics="true" pageId="4" pageNumber="144">P. pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF87866AFAD4A6ADFA85FA52" bold="true" box="[1316,1395,1475,1496]" pageId="4" pageNumber="144">
|
||
<taxonomicNameLabel id="A23D57B0FF87866AFAD4A6ADFA85FA52" box="[1316,1395,1475,1496]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= star). Sources:
|
||
<bibRefCitation id="EFEB4B28FF87866AFC36A68FFB08FA7C" author="Echevarria LY & Venegas PJ" box="[966,1278,1504,1527]" pageId="4" pageNumber="144" pagination="26 - 33" refId="ref3838" refString="Echevarria LY, Venegas PJ. 2015. A new elusive species of Petracola (Squamata: Gymnophthalmidae) from the Utcubamba Basin in the Andes of northern Peru. Amphibian & Reptile Conservation 9 (1) [Special Section]: 26 - 33 (e 107)." type="journal article" year="2015">Echevarría and Venegas (2015)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEB4B28FF87866AFAFDA68FFCAEF99E" author="Echevarria LY" pageId="4" pageNumber="144" refId="ref3795" refString="Echevarria LY. 2014. Revision del estado actual de la taxonomia de Petracola ventrimaculatus Boulenger, 1900 utilizando evidencias morfologicas y ecologicas. B. Sc. thesis, Universidad Nacional Agraria La Molina, Lima, Peru. 128 p." type="book" year="2014">Echevarría (2014)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEB4B28FF87866AFC91A690FBD1F99E" author="Kizirian D & Bayefsky-Anand S & Eriksson A & Le M & Donnelly M." box="[865,1063,1534,1556]" pageId="4" pageNumber="144" pagination="53 - 62" refId="ref3974" refString="Kizirian D, Bayefsky-Anand S, Eriksson A, Le M, Donnelly M. 2008. A new Petracola and a redescription of P. ventrimaculatus (Squamata: Gymnophthalmidae). Zootaxa 1700: 53 - 62." type="journal article" year="2008">Kizirian et al. (2008)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="EFEB4B28FF87866AFBA8A690FACDF99E" author="Kohler G & Lehr E." box="[1112,1339,1534,1556]" pageId="4" pageNumber="144" pagination="501 - 518" refId="ref4014" refString="Kohler G, Lehr E. 2004. Comments on Euspondylus and Proctoporus (Squamata: Gymnophthalmidae) from Peru, with the description of three new species and a key to the Peruvian species. Herpetologica 60 (4): 501 - 518." type="journal article" year="2004">Köhler and Lehr (2004)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3606552FF87866AFF7DA50BFD15F977" pageId="4" pageNumber="144" type="etymology">
|
||
<paragraph id="8BC536D9FF87866AFF7DA50BFEFEF9F7" blockId="4.[141,264,1637,1661]" box="[141,264,1637,1661]" pageId="4" pageNumber="144">
|
||
<heading id="D08D81B5FF87866AFF7DA50BFEFEF9F7" bold="true" box="[141,264,1637,1661]" fontSize="10" level="2" pageId="4" pageNumber="144" reason="2">
|
||
<emphasis id="B90EEACBFF87866AFF7DA50BFEFEF9F7" bold="true" box="[141,264,1637,1661]" pageId="4" pageNumber="144">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC536D9FF87866AFF7DA5CBFD15F977" blockId="4.[141,745,1701,1789]" pageId="4" pageNumber="144">
|
||
The specific epithet is an adjective that recognizes Gran Pajatén archaeological remains, which, like
|
||
<taxonomicName id="4C7A4D5AFF87866AFD70A5ABFF0FF977" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF87866AFD70A5ABFF0FF977" italics="true" pageId="4" pageNumber="144">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF87866AFEF0A58BFEA0F977" bold="true" box="[256,342,1765,1789]" pageId="4" pageNumber="144">
|
||
<taxonomicNameLabel id="A23D57B0FF87866AFEF0A58BFEA0F977" box="[256,342,1765,1789]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, occurs in Río Abiseo National Park.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606552FF87866AFF7DA44BFC99F817" pageId="4" pageNumber="144" type="discussion">
|
||
<paragraph id="8BC536D9FF87866AFF7DA44BFD81F8B7" blockId="4.[141,631,1829,1853]" box="[141,631,1829,1853]" pageId="4" pageNumber="144">
|
||
<heading id="D08D81B5FF87866AFF7DA44BFD81F8B7" bold="true" box="[141,631,1829,1853]" fontSize="10" level="2" pageId="4" pageNumber="144" reason="2">
|
||
<emphasis id="B90EEACBFF87866AFF7DA44BFD81F8B7" bold="true" box="[141,631,1829,1853]" pageId="4" pageNumber="144">Habitat, Ecological Notes, and Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC536D9FF87866AFF7DA40BFC99F817" blockId="4.[141,746,1893,1949]" lastBlockId="4.[791,1395,1605,1949]" pageId="4" pageNumber="144">
|
||
The
|
||
<typeStatus id="54C1887BFF87866AFF4CA40BFEEFF8F7" box="[188,281,1893,1917]" pageId="4" pageNumber="144" type="holotype">holotype</typeStatus>
|
||
was taken from the ground in grassland, in a rocky area dominated by bunchgrasses of
|
||
<taxonomicName id="4C7A4D5AFF87866AFDBAA4EBFCCEF9D6" box="[586,824,1605,1949]" class="Liliopsida" family="Poaceae" genus="Calamagrostis" kingdom="Plantae" order="Poales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B90EEACBFF87866AFDBAA4EBFD1FF817" box="[586,745,1925,1949]" italics="true" pageId="4" pageNumber="144">Calamagrostis</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7A4D5AFF87866AFCBAA528FC38F9D6" box="[842,974,1605,1629]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B90EEACBFF87866AFCBAA528FC56F9D7" box="[842,928,1606,1629]" italics="true" pageId="4" pageNumber="144">Festuca</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90EEACBFF87866AFC10A52BFBAFF9D7" box="[992,1113,1605,1629]" italics="true" pageId="4" pageNumber="144">Cortaderla</emphasis>
|
||
sp., and
|
||
<taxonomicName id="4C7A4D5AFF87866AFB3DA52BFAC4F9D6" box="[1229,1330,1605,1628]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B90EEACBFF87866AFB3DA52BFAF3F9D6" box="[1229,1285,1605,1628]" italics="true" pageId="4" pageNumber="144">Stipa</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13412A5CFF87866AFAB7A52BFCD1F9F7" captionStart="Fig" captionStartId="4.[791,825,606,628]" captionTargetBox="[791,1395,144,597]" captionTargetId="figure-275@4.[791,1395,144,597]" captionTargetPageId="4" captionText="Fig. 6. Type locality of Petracola pajatensis sp. nov. Photo by Ken R. Young." figureDoi="http://doi.org/10.5281/zenodo.11263282" httpUri="https://zenodo.org/record/11263282/files/figure.png" pageId="4" pageNumber="144">Fig. 6</figureCitation>
|
||
). The habitat of
|
||
<taxonomicName id="4C7A4D5AFF87866AFC1FA50BFB23F9F7" authority="Rodríguez & Mamani, 2020" authorityName="Rodríguez & Mamani" authorityYear="2020" box="[1007,1237,1637,1661]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="pajatensis" status="sp. nov.">
|
||
<emphasis id="B90EEACBFF87866AFC1FA50BFB23F9F7" box="[1007,1237,1637,1661]" italics="true" pageId="4" pageNumber="144">Petracola pajatensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90EEACBFF87866AFB11A50BFACBF9F7" bold="true" box="[1249,1341,1637,1661]" pageId="4" pageNumber="144">
|
||
<taxonomicNameLabel id="A23D57B0FF87866AFB11A50BFACBF9F7" box="[1249,1341,1637,1661]" pageId="4" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was previously disturbed through overgrazing and is currently undergoing secondary succession, including scattered shrubs (
|
||
<taxonomicName id="4C7A4D5AFF87866AFC2CA5ABFBBCF957" box="[988,1098,1733,1757]" class="Magnoliopsida" family="Asteraceae" genus="Baccharis" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="144" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B90EEACBFF87866AFC2CA5ABFBBCF957" box="[988,1098,1733,1757]" italics="true" pageId="4" pageNumber="144">Baccharis</emphasis>
|
||
</taxonomicName>
|
||
). Multiple nest deposition sites contained elliptical egg shells (9.5–11.0 mm). The new species is known only from Los Chochos, a small valley on the western slopes of the Cordillera Oriental, south of the Huancabamba Depression, approximately
|
||
<quantity id="4C829B3CFF87866AFCE7A40BFC9BF8F7" box="[791,877,1893,1917]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="144" unit="km" value="100.0">100 km</quantity>
|
||
south of the localities of
|
||
<taxonomicName id="4C7A4D5AFF87866AFB53A408FA85F8F7" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[1187,1395,1893,1917]" class="Squamata" family="Gymnophthalmidae" genus="Petracola" kingdom="Animalia" pageId="4" pageNumber="144" phylum="Chordata" rank="species" species="ventrimaculatus">
|
||
<emphasis id="B90EEACBFF87866AFB53A408FA85F8F7" box="[1187,1395,1893,1917]" italics="true" pageId="4" pageNumber="144">P. ventrimaculatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13412A5CFF87866AFCEFA4EBFC94F817" box="[799,866,1925,1949]" captionStart="Fig" captionStartId="4.[791,825,1416,1438]" captionTargetBox="[794,1391,700,1396]" captionTargetId="figure-276@4.[791,1395,687,1405]" captionTargetPageId="4" captionText="Fig. 7. Locations of the type localities of the species of Petracola (P. angustisoma = circle, P. ventrimaculatus = square, P. waka = diamond, P. labioocularis = triangle, P. pajatensis sp. nov. = star). Sources: Echevarría and Venegas (2015), Echevarría (2014), Kizirian et al. (2008), and Köhler and Lehr (2004)." figureDoi="http://doi.org/10.5281/zenodo.11263286" httpUri="https://zenodo.org/record/11263286/files/figure.png" pageId="4" pageNumber="144">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |