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<document id="8D717E5A6215C625EC9B1FFD6F4C6EB8" ID-CLB-Dataset="47893" ID-DOI="10.5281/zenodo.196531" ID-GBIF-Dataset="72970506-484a-42c6-a2e5-352474111175" ID-ISSN="1175-5326" ID-Zenodo-Dep="196531" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460245843076" checkinUser="plazi" docAuthor="Yamada, Kazutaka, Yasunaga, Tomohide &amp; Miyamoto, Syôiti" docDate="2010" docId="03D387CEFF945C73FF7EFB96907EC7ED" docLanguage="en" docName="zt02530p028.pdf" docOrigin="Zootaxa 2530" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Montandoniola thripodes Bergroth 1916" docType="treatment" docVersion="16" lastPageNumber="24" masterDocId="FFEAFFB6FF955C76FFE9FFEB925CC551" masterDocTitle="A review of Japanese species of the genus Montandoniola (Hemiptera: Heteroptera: Anthocoridae)" masterLastPageNumber="28" masterPageNumber="19" pageNumber="20" updateTime="1732899683585" updateUser="ExternalLinkService">
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<mods:title id="B489FF9C3BC4CD3B3F2D662D187AC761">A review of Japanese species of the genus Montandoniola (Hemiptera: Heteroptera: Anthocoridae)</mods:title>
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<mods:namePart id="A00E9058A111A911479E6FA366DFB7B4">Yamada, Kazutaka</mods:namePart>
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<mods:namePart id="D33A9D51D3BECB57978AE78984948D04">Yasunaga, Tomohide</mods:namePart>
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<mods:namePart id="7CE9DB269B880C8213F88043978C32EE">Miyamoto, Syôiti</mods:namePart>
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<mods:title id="41BA06893F7BDC197A8E072761706E22">Zootaxa</mods:title>
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<mods:date id="8E5938E29F735203AC6333EF5AC76075">2010</mods:date>
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<treatment id="03D387CEFF945C73FF7EFB96907EC7ED" ID-DOI="http://doi.org/10.5281/zenodo.3510444" ID-GBIF-Taxon="119390478" ID-Zenodo-Dep="3510444" LSID="urn:lsid:plazi:treatment:03D387CEFF945C73FF7EFB96907EC7ED" httpUri="http://treatment.plazi.org/id/03D387CEFF945C73FF7EFB96907EC7ED" lastPageId="5" lastPageNumber="24" pageId="1" pageNumber="20" scope_family="Anthocoridae" scope_order="Hemiptera">
<subSubSection id="C3606553FF945C77FF7EFB969333C1EB" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8BC536D8FF945C77FF7EFB9690D7C1C6" blockId="1.[151,651,1149,1210]" box="[151,651,1149,1175]" pageId="1" pageNumber="20">
<heading id="D08D81B4FF945C77FF7EFB9690D7C1C6" bold="true" box="[151,651,1149,1175]" fontSize="11" level="1" pageId="1" pageNumber="20" reason="1">
<taxonomicName id="4C7A4D5BFF945C77FF7EFB9690D7C1C6" authority="Bergroth, 1916" authorityName="Bergroth" authorityYear="1916" box="[151,651,1149,1175]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF945C77FF7EFB9690D7C1C6" bold="true" box="[151,651,1149,1175]" pageId="1" pageNumber="20">
<emphasis id="B90EEACAFF945C77FF7EFB969395C1C6" bold="true" box="[151,457,1149,1175]" italics="true" pageId="1" pageNumber="20">Montandoniola thripodes</emphasis>
<bibRefCitation id="EFEB4B29FF945C77FE38FB9690D7C1C6" author="Bergroth" box="[465,651,1149,1175]" pageId="1" pageNumber="27" refString="Bergroth, E. (1916) New and little known Heteropterous Hemiptera in the United States National Museum. Proceedings of the United States National Museum, 51 (2150), 247 - 249." type="journal article" year="1916">Bergroth, 1916</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC536D8FF945C77FF7EFB4B9333C1EB" blockId="1.[151,651,1149,1210]" box="[151,367,1184,1210]" pageId="1" pageNumber="20">
(
<figureCitation id="13412A5DFF945C77FF76FB4B9353C1EB" box="[159,271,1184,1210]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 14</figureCitation>
,
<figureCitation id="13412A5DFF945C77FEF5FB4B9360C1EB" box="[284,316,1184,1210]" captionStart="FIGURES 14 16" captionStartId="3.[151,269,1151,1175]" captionTargetBox="[324,1158,178,1109]" captionTargetId="figure@3.[278,1310,158,1130]" captionTargetPageId="3" captionText="FIGURES 14 16. Female left fore tibiae. 14, M. thripodes; 15, M. pictipennis; 16, M. kerzhneri sp. nov. Arrows indicate minute teeth. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/196533/files/figure.png" pageId="1" pageNumber="20">14</figureCitation>
,
<figureCitation id="13412A5DFF945C77FEA1FB4B933AC1EB" box="[328,358,1184,1210]" captionStart="FIGURES 17 22" captionStartId="4.[151,273,1942,1966]" captionTargetBox="[187,1374,397,1885]" captionTargetId="figure@4.[154,1433,351,1921]" captionTargetPageId="4" captionText="FIGURES 17 22. Male (17 20) and female (21, 22) genitalia of M. pictipennis (17 20), M. thripodes (21) and M. kerzhneri sp. nov. (22). 17, Pygophore with paramere, dorsal view; 18, paramere, dorsal view; 19, 20, ditto, two different orientations; 21, 22, copulatory tube, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/196534/files/figure.png" pageId="1" pageNumber="20">21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF945C77FF7EFB019610C0F0" pageId="1" pageNumber="20" type="reference_group">
<paragraph id="8BC536D8FF945C77FF7EFB019308C010" blockId="1.[151,1436,1258,1441]" pageId="1" pageNumber="20">
<treatmentCitationGroup id="AB6A11F6FF945C77FF7EFB019308C010" pageId="1" pageNumber="20">
<taxonomicName id="4C7A4D5BFF945C77FF7EFB0190DEC050" authority="Bergroth, 1916: 233" authorityName="Bergroth" authorityPageNumber="233" authorityYear="1916" box="[151,642,1258,1281]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF945C77FF7EFB0193FAC050" box="[151,422,1258,1281]" italics="true" pageId="1" pageNumber="20">Montandoniola thripodes</emphasis>
<treatmentCitation id="0ADB10C9FF945C77FE45FB0190DEC050" author="Bergroth" box="[428,642,1258,1281]" page="233" pageId="1" pageNumber="20" year="1916">
<bibRefCitation id="EFEB4B29FF945C77FE45FB01900CC050" author="Bergroth" box="[428,592,1258,1281]" pageId="1" pageNumber="27" refString="Bergroth, E. (1916) New and little known Heteropterous Hemiptera in the United States National Museum. Proceedings of the United States National Museum, 51 (2150), 247 - 249." type="journal article" year="1916">Bergroth, 1916</bibRefCitation>
: 233
</treatmentCitation>
</taxonomicName>
(sp. n.);
<treatmentCitation id="0ADB10C9FF945C77FD36FB0191EDC050" author="Carayon" box="[735,945,1258,1281]" page="542" pageId="1" pageNumber="20" year="1961">
<bibRefCitation id="EFEB4B29FF945C77FD36FB019123C050" author="Carayon" box="[735,895,1258,1281]" pageId="1" pageNumber="27" refString="Carayon, J. (1961) Chapter XII, Hemiptera (Heteroptera): Anthocoridae. In: Hansstrom, B., Brinck, P. &amp; Rudebeck, G. (Eds.), South African Animal Life. Result of the Lund University Expedition in 1950 - 1951. 8. Almqvist &amp; Wiksell, Uppsala, pp. 533 - 557." type="journal article" year="1961">Carayon, 1961</bibRefCitation>
: 542
</treatmentCitation>
(note);
<treatmentCitation id="0ADB10C9FF945C77FBECFB019713C050" author="Carayon" box="[1029,1359,1258,1281]" page="208" pageId="1" pageNumber="20" year="1962">
<bibRefCitation id="EFEB4B29FF945C77FBECFB019741C050" author="Carayon" box="[1029,1309,1258,1281]" pageId="1" pageNumber="27" refString="Carayon, J. &amp; Ramade, F. (1962) Note sur la presence en France et en Italie de Montandoniola moraguesi (Puton) avec quelques observations sur cet Heteroptere Anthocoride. Bulletin de la Societe entomologique de France, 67, 207 - 211." type="journal article" year="1962">Carayon &amp; Ramade, 1962</bibRefCitation>
: 208
</treatmentCitation>
(note);
<treatmentCitation id="0ADB10C9FF945C77FF2EFAE193CCC070" author="Herring" box="[199,400,1290,1313]" page="93" pageId="1" pageNumber="20" year="1966">
<bibRefCitation id="EFEB4B29FF945C77FF2EFAE1933AC070" author="Herring" box="[199,358,1290,1313]" pageId="1" pageNumber="27" refString="Herring, J. L. (1966) The current name for an Anthocorid predator of the Cuban Laurel Thrips (Hemiptera: Anthocoridae). Proceedings of the Entomlogical Society of Washington, 68 (2), 93." type="journal article" year="1966">Herring, 1966</bibRefCitation>
: 93
</treatmentCitation>
[syn. with
<taxonomicName id="4C7A4D5BFF945C77FDF0FAE19149C070" authority="Puton" authorityName="Puton" box="[537,789,1290,1313]" class="Insecta" genus="Montandoniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="moraguesi">
<emphasis id="B90EEACAFF945C77FDF0FAE190EFC070" box="[537,691,1290,1313]" italics="true" pageId="1" pageNumber="20">M. moraguesi</emphasis>
(Puton)
</taxonomicName>
]. Restored by
<treatmentCitation id="0ADB10C9FF945C77FC54FAE19751C070" ID-DOI="http://doi.org/10.5281/zenodo.3501909" ID-Zenodo-Dep="3501909" author="Pluot-Sigwalt" box="[957,1293,1290,1313]" httpUri="http://treatment.plazi.org/id/03810A72B870FF8EFF7B7EE2B209FA19" page="33" pageId="1" pageNumber="20" year="2009">
<bibRefCitation id="EFEB4B29FF945C77FC54FAE196BEC070" author="Pluot-Sigwalt" box="[957,1250,1290,1313]" pageId="1" pageNumber="27" refString="Pluot-Sigwalt, D., Streito, J. C. &amp; Matocq, A. (2009) Is Montandoniola moraguesi (Puton, 1896) a mixture of different species? (Hemiptera: Heteroptera: Anthocoridae). Zootaxa, 2208, 25 - 43." type="journal article" year="2009">
Pluot-Sigwalt
<emphasis id="B90EEACAFF945C77FBB2FAE196CBC070" box="[1115,1175,1290,1313]" italics="true" pageId="1" pageNumber="20">et al.</emphasis>
, 2009
</bibRefCitation>
: 33
</treatmentCitation>
(illustration, redefinition).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC536D8FF945C77FF7EFAA19197C030" blockId="1.[151,1436,1258,1441]" box="[151,971,1354,1377]" pageId="1" pageNumber="20">
<treatmentCitationGroup id="AB6A11F6FF945C77FF7EFAA19197C030" box="[151,971,1354,1377]" pageId="1" pageNumber="20">
<taxonomicName id="4C7A4D5BFF945C77FF7EFAA1900DC030" authority="Esaki, 1931" authorityName="Esaki" authorityYear="1931" box="[151,593,1354,1377]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF945C77FF7EFAA193EBC030" box="[151,439,1354,1377]" italics="true" pageId="1" pageNumber="20">Montandoniola pictipennis</emphasis>
(
<bibRefCitation id="EFEB4B29FF945C77FE2CFAA19015C030" author="Esaki" box="[453,585,1354,1377]" pageId="1" pageNumber="27" refString="Esaki, T. (1931) Undescribed Hemiptera from Japan and Formosa. Annotationes Zoologicae Japonenses, 13 (3), 261 - 269." type="journal article" year="1931">Esaki, 1931</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0ADB10C9FF945C77FDB2FAA190AAC030" author="Hiura" box="[603,758,1354,1377]" page="2" pageId="1" pageNumber="20" year="1959">
<bibRefCitation id="EFEB4B29FF945C77FDB2FAA19082C030" author="Hiura" box="[603,734,1354,1377]" pageId="1" pageNumber="27" refString="Hiura, I. (1959) Contribution to the knowledge of the Anthocoridae from Japan and its adjacent territories (Hemiptera- Heteroptera) 1. Bulletin of the Osaka Museum of Natural History, 11, 1 - 10." type="journal article" year="1959">Hiura, 1959</bibRefCitation>
: 2
</treatmentCitation>
(misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC536D8FF945C77FF7EFA819610C0F0" blockId="1.[151,1436,1258,1441]" pageId="1" pageNumber="20">
<treatmentCitationGroup id="AB6A11F6FF945C77FF7EFA819610C0F0" pageId="1" pageNumber="20">
<taxonomicName id="4C7A4D5BFF945C77FF7EFA819000C0D0" authority="Puton, 1896" authorityName="Puton" authorityYear="1896" box="[151,604,1386,1409]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="moraguesi">
<emphasis id="B90EEACAFF945C77FF7EFA8193EBC0D0" box="[151,439,1386,1409]" italics="true" pageId="1" pageNumber="20">Montandoniola moraguesi</emphasis>
(Puton, 1896)
</taxonomicName>
:
<treatmentCitation id="0ADB10C9FF945C77FD83FA819174C0D0" author="Hiura" box="[618,808,1386,1409]" page="122" pageId="1" pageNumber="20" year="1979">
<bibRefCitation id="EFEB4B29FF945C77FD83FA8190AEC0D0" author="Hiura" box="[618,754,1386,1409]" pageId="1" pageNumber="27" refString="Hiura, I. (1979) Hemiptera: Heteroptera. In: Ito, S. et al. (Eds.), Coloured Illustrations of the Insects of Japan Vol. II. Hoikusha, Osaka, pp. 95 - 129. (in Japanese)" type="book chapter" year="1979">Hiura, 1979</bibRefCitation>
: 122
</treatmentCitation>
(note, figure);
<treatmentCitation id="0ADB10C9FF945C77FC3FFA819704C0D0" author="Miyamoto" box="[982,1368,1386,1409]" page="167" pageId="1" pageNumber="20" year="1989">
<bibRefCitation id="EFEB4B29FF945C77FC3FFA81977DC0D0" author="Miyamoto" box="[982,1313,1386,1409]" pageId="1" pageNumber="27" refString="Miyamoto, S. &amp; Yasunaga, T. (1989) Hemiptera, Heteroptera. In: Hirashima, Y. (Ed), A check list of Japanese insects. 2. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, pp. 151 - 188." type="journal article" year="1989">Miyamoto &amp; Yasunaga, 1989</bibRefCitation>
: 167
</treatmentCitation>
(list);
<treatmentCitation id="0ADB10C9FF945C77FF2EFA6193FAC0F0" author="Yasunaga" box="[199,422,1418,1441]" page="169" pageId="1" pageNumber="20" year="1993">
<bibRefCitation id="EFEB4B29FF945C77FF2EFA619328C0F0" author="Yasunaga" box="[199,372,1418,1441]" pageId="1" pageNumber="28" refString="Yasunaga, T. (1993) Anthocoridae. In: Tomokuni, M. (Ed.), A field guide to Japanese Bugs - Terrestrial Heteropterans. Zenkoku Noson Kyoiku Kyokai, Tokyo, pp. 167 - 169, pls. 26 - 28. (in Japanese)" type="book chapter" year="1993">Yasunaga, 1993</bibRefCitation>
: 169
</treatmentCitation>
(note, figure);
<treatmentCitation id="0ADB10C9FF945C77FDAEFA61917AC0F0" author="Yasunaga" box="[583,806,1418,1441]" page="287" pageId="1" pageNumber="20" year="2001">
<bibRefCitation id="EFEB4B29FF945C77FDAEFA6190A8C0F0" author="Yasunaga" box="[583,756,1418,1441]" pageId="1" pageNumber="28" refString="Yasunaga, T. (2001) Family Anthocoridae Fieber, 1836 flower bugs, minute pirate bugs. In: Yasunaga, T. et al. (Eds.), A field guide to Japanese Bugs II - Terrestrial Heteropterans. Zenkoku Noson Kyoiku Kyokai, Tokyo, pp. 278 - 303, pls. 85 - 89. (in Japanese)" type="book chapter" year="2001">Yasunaga, 2001</bibRefCitation>
: 287
</treatmentCitation>
(note) (misidentifications).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF945C74FF7EFA2492B3C416" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20" type="diagnosis">
<paragraph id="8BC536D8FF945C77FF7EFA2492B6C2A5" blockId="1.[151,1437,1487,2036]" pageId="1" pageNumber="20">
<emphasis id="B90EEACAFF945C77FF7EFA24934BC0B8" bold="true" box="[151,279,1487,1513]" pageId="1" pageNumber="20">Diagnosis.</emphasis>
In addition to the redefinition of
<taxonomicName id="4C7A4D5BFF945C77FD73FA3B916CC0B8" box="[666,816,1488,1513]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF945C77FD73FA3B916CC0B8" box="[666,816,1488,1513]" italics="true" pageId="1" pageNumber="20">M. thripodes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFEB4B29FF945C77FCB3FA3B96CFC0BB" box="[858,1171,1488,1514]" pageId="1" pageNumber="27" refString="Pluot-Sigwalt, D., Streito, J. C. &amp; Matocq, A. (2009) Is Montandoniola moraguesi (Puton, 1896) a mixture of different species? (Hemiptera: Heteroptera: Anthocoridae). Zootaxa, 2208, 25 - 43." type="journal article">
Pluot-Sigwalt
<emphasis id="B90EEACAFF945C77FBEBFA3B9660C0B8" box="[1026,1084,1488,1513]" italics="true" pageId="1" pageNumber="20">et al.</emphasis>
(2009)
</bibRefCitation>
, we herein provide the following revised diagnostic characters for Japanese specimens: Head (
<figureCitation id="13412A5DFF945C77FC30FA1E9615C35E" box="[985,1097,1525,1551]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 14</figureCitation>
) dark brown to black; length excluding neck as long as width across eyes. Antennal segments I and II dark brown to black; segments III and IV pale yellow; segment IV somewhat fuscous (
<figureCitation id="13412A5DFF945C77FCF6F9AB91CBC30B" box="[799,919,1600,1626]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 1, 3</figureCitation>
); segment II (
<figureCitation id="13412A5DFF945C77FBACF9AB96E6C30B" box="[1093,1210,1600,1626]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 1, 3</figureCitation>
) greatly enlarged, slightly longer than head width across eyes; segment IV weakly flattened. Labium (
<figureCitation id="13412A5DFF945C77FB9DF98E96B9C32E" box="[1140,1253,1637,1663]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 2, 4</figureCitation>
) dark brown to black, extending near fore coxae; apex of segment III pale yellow; segment IV pale yellow, except for dark apex. Pronotum (
<figureCitation id="13412A5DFF945C77FE8DF95B9389C39B" box="[356,469,1712,1738]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 1, 3</figureCitation>
) dark brown to black; anterior margin about as wide as or slightly narrower than mesal length; lateral margin nearly straight; lateral carina well developed anteriorly; posterior margin concave, 2.372.55 times as wide as anterior margin. Scutellum (
<figureCitation id="13412A5DFF945C77FC7BF911965DC245" box="[914,1025,1786,1812]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 1, 3</figureCitation>
) overall black. Hemelytra (
<figureCitation id="13412A5DFF945C77FAADF91192FAC26B" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 1, 3</figureCitation>
) dark brown to black with whitish central patch on most of clavus and corium; membrane mesially with distinct dark stripe, except for basal semi-transparent area. Legs (
<figureCitation id="13412A5DFF945C77FC79F8AE91A2C20E" box="[912,1022,1861,1887]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 2, 4</figureCitation>
) dark brown to black; fore and mid tibiae pale yellow, except for dark extreme bases; hind tibiae with paler extreme apex; male fore tibiae armed ventrally with a row of 1622 minute, fuscous teeth, female without minute teeth (
<figureCitation id="13412A5DFF945C77FB8CF87B96E1C2FB" box="[1125,1213,1936,1962]" captionStart="FIGURES 14 16" captionStartId="3.[151,269,1151,1175]" captionTargetBox="[324,1158,178,1109]" captionTargetId="figure@3.[278,1310,158,1130]" captionTargetPageId="3" captionText="FIGURES 14 16. Female left fore tibiae. 14, M. thripodes; 15, M. pictipennis; 16, M. kerzhneri sp. nov. Arrows indicate minute teeth. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/196533/files/figure.png" pageId="1" pageNumber="20">Fig. 14</figureCitation>
); tarsi pale yellow, with dark apex. Ostiolar peritreme angular posteriorly. Abdomen (
<figureCitation id="13412A5DFF945C77FC5DF85E967BC29E" box="[948,1063,1973,1999]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="1" pageNumber="20">Figs. 2, 4</figureCitation>
) black and tinged with reddish brown.
</paragraph>
<paragraph id="8BC536D8FF975C74FF2CFF7392B3C416" blockId="2.[151,1436,152,327]" pageId="2" pageNumber="21">Male genitalia: Pygophore covered with many long setae on right side and 810 very long, stout setae on posteroventral surface, of which the longest are longer than length of pygophore, with scattered short setae anterior to genital opening; flagellum straight, long, thin, shorter than twice width of cone, not surpassing left edge of pygophore, basally very close to cone; cone very thin and acute apicad, with conspicuous indentation at base.</paragraph>
</subSubSection>
<caption id="DF056650FF975C74FF7EF8F193F2C2E0" httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="2" pageNumber="21" targetBox="[192,1414,421,1785]" targetPageId="2">
<paragraph id="8BC536D8FF975C74FF7EF8F193F2C2E0" blockId="2.[151,1436,1818,1969]" pageId="2" pageNumber="21">
<emphasis id="B90EEACAFF975C74FF7EF8F1930CC263" bold="true" box="[151,336,1818,1842]" pageId="2" pageNumber="21">FIGURES 113.</emphasis>
Japanese
<taxonomicName id="4C7A4D5BFF975C74FE52F8F19003C260" box="[443,607,1818,1841]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEACAFF975C74FE52F8F19003C260" box="[443,607,1818,1841]" italics="true" pageId="2" pageNumber="21">Montandoniola</emphasis>
</taxonomicName>
species. 14,
<taxonomicName id="4C7A4D5BFF975C74FD1CF8F19122C260" box="[757,894,1818,1841]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF975C74FD1CF8F19122C260" box="[757,894,1818,1841]" italics="true" pageId="2" pageNumber="21">M. thripodes</emphasis>
</taxonomicName>
, male (1, 2) and female (3, 4); 5, 6,
<taxonomicName id="4C7A4D5BFF975C74FB14F8F197C9C260" box="[1277,1429,1818,1841]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF975C74FB14F8F197C9C260" box="[1277,1429,1818,1841]" italics="true" pageId="2" pageNumber="21">M. pictipennis</emphasis>
</taxonomicName>
, male; 713,
<taxonomicName id="4C7A4D5BFF975C74FEF5F8D093F9C203" box="[284,421,1851,1874]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
<emphasis id="B90EEACAFF975C74FEF5F8D093F9C203" box="[284,421,1851,1874]" italics="true" pageId="2" pageNumber="21">M. kerzhneri</emphasis>
</taxonomicName>
<emphasis id="B90EEACAFF975C74FE45F8D1905DC203" bold="true" box="[428,513,1850,1874]" pageId="2" pageNumber="21">
<taxonomicNameLabel id="A23D57B1FF975C74FE45F8D1905DC203" box="[428,513,1850,1874]" pageId="2" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype. 18, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 18; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11.
</paragraph>
</caption>
<caption id="DF056650FF965C75FF7EFB9493A0C1E6" httpUri="https://zenodo.org/record/196533/files/figure.png" pageId="3" pageNumber="22" targetBox="[324,1158,178,1109]" targetPageId="3">
<paragraph id="8BC536D8FF965C75FF7EFB9493A0C1E6" blockId="3.[151,1435,1151,1207]" pageId="3" pageNumber="22">
<emphasis id="B90EEACAFF965C75FF7EFB949300C1C6" bold="true" box="[151,348,1151,1175]" pageId="3" pageNumber="22">FIGURES 1416.</emphasis>
Female left fore tibiae. 14,
<taxonomicName id="4C7A4D5BFF965C75FD68FB6B9155C1C6" box="[641,777,1152,1175]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF965C75FD68FB6B9155C1C6" box="[641,777,1152,1175]" italics="true" pageId="3" pageNumber="22">M. thripodes</emphasis>
</taxonomicName>
; 15,
<taxonomicName id="4C7A4D5BFF965C75FCD4FB6B9189C1C6" box="[829,981,1152,1175]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF965C75FCD4FB6B9189C1C6" box="[829,981,1152,1175]" italics="true" pageId="3" pageNumber="22">M. pictipennis</emphasis>
</taxonomicName>
; 16,
<taxonomicName id="4C7A4D5BFF965C75FBE3FB6B96CCC1C6" box="[1034,1168,1152,1175]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="22" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
<emphasis id="B90EEACAFF965C75FBE3FB6B96CCC1C6" box="[1034,1168,1152,1175]" italics="true" pageId="3" pageNumber="22">M. kerzhneri</emphasis>
</taxonomicName>
<emphasis id="B90EEACAFF965C75FB7EFB9496B0C1C6" bold="true" box="[1175,1260,1151,1175]" pageId="3" pageNumber="22">
<taxonomicNameLabel id="A23D57B1FF965C75FB7EFB9496B0C1C6" box="[1175,1260,1151,1175]" pageId="3" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Arrows indicate minute teeth. Scale bars: 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3606553FF965C75FF2CFB0E9049C325" pageId="3" pageNumber="22" type="description">
<paragraph id="8BC536D8FF965C75FF2CFB0E933CC01B" blockId="3.[151,1437,1253,2026]" pageId="3" pageNumber="22">
<emphasis id="B90EEACAFF965C75FF2CFB0E900EC1AE" bold="true" box="[197,594,1253,1279]" pageId="3" pageNumber="22">Description of female genitalia.</emphasis>
Copulatory tube (
<figureCitation id="13412A5DFF965C75FCC5FB0E91DFC1AE" box="[812,899,1253,1279]" captionStart="FIGURES 17 22" captionStartId="4.[151,273,1942,1966]" captionTargetBox="[187,1374,397,1885]" captionTargetId="figure@4.[154,1433,351,1921]" captionTargetPageId="4" captionText="FIGURES 17 22. Male (17 20) and female (21, 22) genitalia of M. pictipennis (17 20), M. thripodes (21) and M. kerzhneri sp. nov. (22). 17, Pygophore with paramere, dorsal view; 18, paramere, dorsal view; 19, 20, ditto, two different orientations; 21, 22, copulatory tube, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/196534/files/figure.png" pageId="3" pageNumber="22">Fig. 21</figureCitation>
) mesally fused on intersegmental membrane between sterna VII and VIII, very close to base of ovipositor, shortened, cylindroid, about twice as long as maximum width.
</paragraph>
<paragraph id="8BC536D8FF965C75FF2CFABE9049C325" blockId="3.[151,1437,1253,2026]" pageId="3" pageNumber="22">
<emphasis id="B90EEACAFF965C75FF2CFABE93DEC03E" bold="true" box="[197,386,1365,1391]" pageId="3" pageNumber="22">Measurements</emphasis>
[% (n = 10)/ Ψ (n = 10)]. Body length 2.703.00/ 2.953.25; head length (excl. neck) 0.380.41/ 0.380.41; head width across eyes 0.390.41/ 0.380.41; vertex width 0.170.19/ 0.190.21; width between ocelli 0.110.13/ 0.130.16; length of antennal segments I-IV 0.120.14/ 0.130.15, 0.400.44/ 0.380.47, 0.230.28/ 0.240.28, and 0.280.31/ 0.260.31; length of labial segments II-IV 0.110.13/ 0.120.14, 0.310.36/ 0.330.39, and 0.230.25/ 0.230.26; anterior pronotal width 0.340.36/ 0.340.37; mesal pronotal length 0.330.36/ 0.370.40; basal pronotal width 0.770.89/ 0.850.96; length of embolial margin 0.690.81/ 0.750.81; length of cuneal margin 0.460.54/ 0.500.61; maximum width across hemelytra 0.780.92/ 0.890.98.
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF965C75FF2CF96B9605C2CE" pageId="3" pageNumber="22" type="materials_examined">
<paragraph id="8BC536D8FF965C75FF2CF96B9605C2CE" blockId="3.[151,1437,1253,2026]" pageId="3" pageNumber="22">
<emphasis id="B90EEACAFF965C75FF2CF96B93E0C3CB" bold="true" box="[197,444,1664,1690]" pageId="3" pageNumber="22">Material examined.</emphasis>
<collectingCountry id="F36D7648FF965C75FE2DF96B907EC3CB" box="[452,546,1664,1690]" name="Japan" pageId="3" pageNumber="22">JAPAN</collectingCountry>
: Ryukyus: [Tokara Isls.] 1Ψ, Takarajima Is.,
<date id="FFC41018FF965C75FBD0F96B96A8C3CB" box="[1081,1268,1664,1690]" pageId="3" pageNumber="22" value="1953-05-26" valueMax="1953-06-01" valueMin="1953-05-26">26.v.1.vi.1953</date>
, S. Miyamoto (
<collectionCode id="ED6BAE1DFF965C75FF76F94E935EC3EE" box="[159,258,1701,1727]" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Sam Noble Oklahoma Museum of Natural History" pageId="3" pageNumber="22">OMNH</collectionCode>
). [Amami-Ôshima Is.] 3Ψ (one shown in
<figureCitation id="13412A5DFF965C75FD15F94E910AC3EE" box="[764,854,1701,1727]" captionStart="FIGURES 14 16" captionStartId="3.[151,269,1151,1175]" captionTargetBox="[324,1158,178,1109]" captionTargetId="figure@3.[278,1310,158,1130]" captionTargetPageId="3" captionText="FIGURES 14 16. Female left fore tibiae. 14, M. thripodes; 15, M. pictipennis; 16, M. kerzhneri sp. nov. Arrows indicate minute teeth. Scale bars: 0.2 mm." httpUri="https://zenodo.org/record/196533/files/figure.png" pageId="3" pageNumber="22">Fig. 14</figureCitation>
), Sokaru, Setouchi-chô,
<date id="FFC41018FF965C75FB6AF94E975BC3EE" box="[1155,1287,1701,1727]" pageId="3" pageNumber="22" value="2002-09-12">12.ix.2002</date>
, K. Yamada (
<collectionCode id="ED6BAE1DFF965C75FF76F92192A6C3B5" box="[159,250,1738,1764]" pageId="3" pageNumber="22">TKPM</collectionCode>
). [Okinawa Is.] 3%3Ψ (one shown in
<figureCitation id="13412A5DFF965C75FD2DF9219167C3B5" box="[708,827,1738,1764]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="3" pageNumber="22">Figs. 3, 4</figureCitation>
), Hedo, Kunigami-son,
<date id="FFC41018FF965C75FB88F92196B4C3B5" box="[1121,1256,1738,1764]" pageId="3" pageNumber="22" value="2002-03-24">24.iii.2002</date>
, T. Ueda &amp; K. Yamada (
<collectionCode id="ED6BAE1DFF965C75FEE5F91B9301C25B" box="[268,349,1776,1802]" pageId="3" pageNumber="22">TKPM</collectionCode>
); 2 % (one shown in
<figureCitation id="13412A5DFF965C75FDB4F91B908EC25B" box="[605,722,1776,1802]" captionStart="FIGURES 1 13" captionStartId="2.[151,269,1818,1842]" captionTargetBox="[192,1414,421,1785]" captionTargetId="figure@2.[153,1433,383,1794]" captionTargetPageId="2" captionText="FIGURES 1 13. Japanese Montandoniola species. 1 4, M. thripodes, male (1, 2) and female (3, 4); 5, 6, M. pictipennis, male; 7 13, M. kerzhneri sp. nov., female, holotype. 1 8, Habitus, dorsal (1, 3, 5, 7) and lateral (2, 4, 6, 8) views; 9, 10, head and pronotum, dorsal (9) and lateral (10) views; 11, ostiolar peritreme and evaporative area of left metapleura, lateroventral view; 12, left fore leg, outer view; 13, left mid leg, dorsal view. Scale bars: 1.0 mm for 1 8; 0.2 mm for 9, 10, 12, 13; 0.1 mm for 11." httpUri="https://zenodo.org/record/196532/files/figure.png" pageId="3" pageNumber="22">Figs. 1, 2</figureCitation>
) 3Ψ (one shown in
<figureCitation id="13412A5DFF965C75FC2FF91B967CC25B" box="[966,1056,1776,1802]" captionStart="FIGURES 17 22" captionStartId="4.[151,273,1942,1966]" captionTargetBox="[187,1374,397,1885]" captionTargetId="figure@4.[154,1433,351,1921]" captionTargetPageId="4" captionText="FIGURES 17 22. Male (17 20) and female (21, 22) genitalia of M. pictipennis (17 20), M. thripodes (21) and M. kerzhneri sp. nov. (22). 17, Pygophore with paramere, dorsal view; 18, paramere, dorsal view; 19, 20, ditto, two different orientations; 21, 22, copulatory tube, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/196534/files/figure.png" pageId="3" pageNumber="22">Fig. 21</figureCitation>
), same locality,
<date id="FFC41018FF965C75FB03F91B972DC25B" box="[1258,1393,1776,1802]" pageId="3" pageNumber="22" value="2002-03-25">25.iii.2002</date>
, K. Yamada (
<collectionCode id="ED6BAE1DFF965C75FEE5F8FE9301C27E" box="[268,349,1813,1839]" pageId="3" pageNumber="22">TKPM</collectionCode>
); 3 %14Ψ, Shurisakiyama, Naha-shi,
<date id="FFC41018FF965C75FCCFF8FE91FDC27E" box="[806,929,1813,1839]" pageId="3" pageNumber="22" value="1993-05-19">19.v.1993</date>
, T. Yasunaga (
<collectionCode id="ED6BAE1DFF965C75FBBFF8FE96F3C27E" box="[1110,1199,1813,1839]" pageId="3" pageNumber="22">TYCN</collectionCode>
); 3%3Ψ, 4 nymphs, Naha-shi,
<date id="FFC41018FF965C75FEF8F8D193D8C205" box="[273,388,1850,1876]" pageId="3" pageNumber="22" value="1993-01-30">30.i.1993</date>
, N. Shingaki (
<collectionCode id="ED6BAE1DFF965C75FDD9F8D190DBC205" box="[560,647,1850,1876]" pageId="3" pageNumber="22">TYCN</collectionCode>
). [Kume Is.] 1%, Higa,
<date id="FFC41018FF965C75FC49F8D1967FC205" box="[928,1059,1850,1876]" pageId="3" pageNumber="22" value="1977-03-21">21.iii.1977</date>
, Y. Hori (
<collectionCode id="ED6BAE1DFF965C75FB71F8D196B3C205" box="[1176,1263,1850,1876]" pageId="3" pageNumber="22">TYCN</collectionCode>
). [Ishigaki Is.] 1Ψ, Urasoko-rindô,
<date id="FFC41018FF965C75FE7CF88B904AC22B" box="[405,534,1888,1914]" pageId="3" pageNumber="22" value="1998-10-29">29.x.1998</date>
, K. Takahashi (
<collectionCode id="ED6BAE1DFF965C75FD35F88B9166C22B" box="[732,826,1888,1914]" pageId="3" pageNumber="22">TKPM</collectionCode>
). [Iriomote Is.] 1Ψ, Takana,
<date id="FFC41018FF965C75FB4DF88B9741C22B" box="[1188,1309,1888,1914]" pageId="3" pageNumber="22" value="2003-04-06">6.iv.2003</date>
, M. Takai (
<collectionCode id="ED6BAE1DFF965C75FF76F86E92ABC2CE" box="[159,247,1925,1951]" pageId="3" pageNumber="22">TKPM</collectionCode>
). [Yonaguni Is.] 1%2Ψ, Mt. Kuburadake,
<date id="FFC41018FF965C75FD32F86E910DC2CE" box="[731,849,1925,1951]" pageId="3" pageNumber="22" value="2000-05-18">18.v.2000</date>
, T. Ishikawa (
<collectionCode id="ED6BAE1DFF965C75FC1CF86E9611C2CE" box="[1013,1101,1925,1951]" pageId="3" pageNumber="22">TKPM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF965C72FF2CF8419726C586" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22" type="distribution">
<paragraph id="8BC536D8FF965C72FF2CF8419726C586" blockId="3.[151,1437,1253,2026]" lastBlockId="4.[151,1436,152,327]" lastPageId="4" lastPageNumber="23" pageId="3" pageNumber="22">
<emphasis id="B90EEACAFF965C75FF2CF841932CC295" bold="true" box="[197,368,1962,1988]" pageId="3" pageNumber="22">Distribution.</emphasis>
Hong-Kong (
<bibRefCitation id="EFEB4B29FF965C75FDF7F8419080C295" author="Bergroth" box="[542,732,1962,1988]" pageId="3" pageNumber="27" refString="Bergroth, E. (1916) New and little known Heteropterous Hemiptera in the United States National Museum. Proceedings of the United States National Museum, 51 (2150), 247 - 249." type="journal article" year="1916">Bergroth 1916</bibRefCitation>
);
<collectingCountry id="F36D7648FF965C75FD1AF8419166C295" box="[755,826,1962,1988]" name="Japan" pageId="3" pageNumber="22">Japan</collectingCountry>
[the Ryukyus: Tokara (Takarajima Is.), Amami- Ôshima, Okinawa, Kume, Ishigaki, Iriomote and Yonaguni Islands]. This species is recorded from
<collectingCountry id="F36D7648FF965C75FAC7F83B972DC2BB" box="[1326,1393,2000,2026]" name="Japan" pageId="3" pageNumber="22">Japan</collectingCountry>
for the first time.
<bibRefCitation id="EFEB4B29FF915C72FED4FF73938AC5E3" author="Hiura" box="[317,470,152,178]" pageId="4" pageNumber="27" refString="Hiura, I. (1959) Contribution to the knowledge of the Anthocoridae from Japan and its adjacent territories (Hemiptera- Heteroptera) 1. Bulletin of the Osaka Museum of Natural History, 11, 1 - 10." type="journal article" year="1959">Hiura (1959)</bibRefCitation>
recorded
<taxonomicName id="4C7A4D5BFF915C72FDA4FF7390A9C5E0" box="[589,757,152,177]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF915C72FDA4FF7390A9C5E0" box="[589,757,152,177]" italics="true" pageId="4" pageNumber="23">M. pictipennis</emphasis>
</taxonomicName>
from Tokara Islands on the basis of a female collected in 1953 by S. Miyamoto. However, we examined it and confirmed that the specimen is actually
<taxonomicName id="4C7A4D5BFF915C72FB34FF56972FC587" box="[1245,1395,189,214]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF915C72FB34FF56972FC587" box="[1245,1395,189,214]" italics="true" pageId="4" pageNumber="23">M. thripodes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF915C73FF2CFF099036C473" lastPageId="5" lastPageNumber="24" pageId="4" pageNumber="23" type="discussion">
<paragraph id="8BC536D8FF915C72FF2CFF099641C416" blockId="4.[151,1436,152,327]" pageId="4" pageNumber="23">
<emphasis id="B90EEACAFF915C72FF2CFF099363C5AD" bold="true" box="[197,319,226,252]" pageId="4" pageNumber="23">Remarks.</emphasis>
<taxonomicName id="4C7A4D5BFF915C72FEAEFF09902AC5AA" box="[327,630,226,251]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF915C72FEAEFF09902AC5AA" box="[327,630,226,251]" italics="true" pageId="4" pageNumber="23">Montandoniola thripodes</emphasis>
</taxonomicName>
was originally described from Hong-Kong by
<bibRefCitation id="EFEB4B29FF915C72FB41FF099730C5AD" author="Bergroth" box="[1192,1388,226,252]" pageId="4" pageNumber="27" refString="Bergroth, E. (1916) New and little known Heteropterous Hemiptera in the United States National Museum. Proceedings of the United States National Museum, 51 (2150), 247 - 249." type="journal article" year="1916">Bergroth (1916)</bibRefCitation>
. No additional specimen has been recognized since the original description. In this study, we found it on the Ryukyu Islands of
<collectingCountry id="F36D7648FF915C72FE9DFEC693EBC416" box="[372,439,301,327]" name="Japan" pageId="4" pageNumber="23">Japan</collectingCountry>
and described the female genitalia for the first time.
</paragraph>
<caption id="DF056650FF915C72FF7EF87D91DEC2A2" httpUri="https://zenodo.org/record/196534/files/figure.png" pageId="4" pageNumber="23" targetBox="[187,1374,397,1885]" targetPageId="4">
<paragraph id="8BC536D8FF915C72FF7EF87D91DEC2A2" blockId="4.[151,1437,1942,2035]" pageId="4" pageNumber="23">
<emphasis id="B90EEACAFF915C72FF7EF87D9338C2FF" bold="true" box="[151,356,1942,1966]" pageId="4" pageNumber="23">FIGURES 1722.</emphasis>
Male (1720) and female (21, 22) genitalia of
<taxonomicName id="4C7A4D5BFF915C72FC9CF87D9649C2FC" box="[885,1045,1942,1965]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF915C72FC9CF87D9649C2FC" box="[885,1045,1942,1965]" italics="true" pageId="4" pageNumber="23">M. pictipennis</emphasis>
</taxonomicName>
(1720),
<taxonomicName id="4C7A4D5BFF915C72FB6AF87D974DC2FC" box="[1155,1297,1942,1965]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF915C72FB6AF87D974DC2FC" box="[1155,1297,1942,1965]" italics="true" pageId="4" pageNumber="23">M. thripodes</emphasis>
</taxonomicName>
(21) and
<taxonomicName id="4C7A4D5BFF915C72FA97F87D92A0C281" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="23" phylum="Arthropoda" rank="species" species="kerzhneri" status="sp. nov.">
<emphasis id="B90EEACAFF915C72FA97F87D92A0C281" italics="true" pageId="4" pageNumber="23">M. kerzhneri</emphasis>
</taxonomicName>
<emphasis id="B90EEACAFF915C72FEEBF852930BC280" bold="true" box="[258,343,1977,2001]" pageId="4" pageNumber="23">
<taxonomicNameLabel id="A23D57B1FF915C72FEEBF852930BC280" box="[258,343,1977,2001]" pageId="4" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(22). 17, Pygophore with paramere, dorsal view; 18, paramere, dorsal view; 19, 20, ditto, two different orientations; 21, 22, copulatory tube, dorsal view. Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC536D8FF905C73FF2CFF739036C473" blockId="5.[151,1437,152,700]" pageId="5" pageNumber="24">
The copulatory tube of
<taxonomicName id="4C7A4D5BFF905C73FE33FF73902CC5E0" box="[474,624,152,177]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF905C73FE33FF73902CC5E0" box="[474,624,152,177]" italics="true" pageId="5" pageNumber="24">M. thripodes</emphasis>
</taxonomicName>
somewhat resembles that in
<taxonomicName id="4C7A4D5BFF905C73FC2CFF739736C5E3" authority="Streito &amp; Matocq, 2009" authorityName="Streito &amp; Matocq" authorityYear="2009" box="[965,1386,152,178]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B90EEACAFF905C73FC2CFF739614C5E0" box="[965,1096,152,177]" italics="true" pageId="5" pageNumber="24">M. confusa</emphasis>
<bibRefCitation id="EFEB4B29FF905C73FBB9FF739736C5E3" author="Streito" box="[1104,1386,152,178]" pageId="5" pageNumber="27" refString="Pluot-Sigwalt, D., Streito, J. C. &amp; Matocq, A. (2009) Is Montandoniola moraguesi (Puton, 1896) a mixture of different species? (Hemiptera: Heteroptera: Anthocoridae). Zootaxa, 2208, 25 - 43." type="journal article" year="2009">Streito &amp; Matocq, 2009</bibRefCitation>
</taxonomicName>
, but can be distinguished by its location and length: located very near base of ovipositor in
<taxonomicName id="4C7A4D5BFF905C73FB21FF569738C587" box="[1224,1380,189,214]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF905C73FB21FF569738C587" box="[1224,1380,189,214]" italics="true" pageId="5" pageNumber="24">M. thripodes</emphasis>
</taxonomicName>
, but mesally located in middle of sternum VII in
<taxonomicName id="4C7A4D5BFF905C73FD76FF09917EC5AA" box="[671,802,226,251]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B90EEACAFF905C73FD76FF09917EC5AA" box="[671,802,226,251]" italics="true" pageId="5" pageNumber="24">M. confusa</emphasis>
</taxonomicName>
; and, about twice as long as wide in
<taxonomicName id="4C7A4D5BFF905C73FB3BFF099734C5AA" box="[1234,1384,226,251]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF905C73FB3BFF099734C5AA" box="[1234,1384,226,251]" italics="true" pageId="5" pageNumber="24">M. thripodes</emphasis>
</taxonomicName>
, but slightly longer than wide in
<taxonomicName id="4C7A4D5BFF905C73FE09FEE3903EC470" box="[480,610,264,289]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B90EEACAFF905C73FE09FEE3903EC470" box="[480,610,264,289]" italics="true" pageId="5" pageNumber="24">M. confusa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF905C73FF2FFEC69742C776" pageId="5" pageNumber="24" type="reference_group">
<paragraph id="8BC536D8FF905C73FF2FFEC69742C776" blockId="5.[151,1437,152,700]" pageId="5" pageNumber="24">
<taxonomicName id="4C7A4D5BFF905C73FF2FFEC693A8C417" box="[198,500,301,326]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF905C73FF2FFEC693A8C417" box="[198,500,301,326]" italics="true" pageId="5" pageNumber="24">Montandoniola thripodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7A4D5BFF905C73FDD8FEC69080C417" box="[561,732,301,326]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF905C73FDD8FEC69080C417" box="[561,732,301,326]" italics="true" pageId="5" pageNumber="24">M. pictipennis</emphasis>
</taxonomicName>
have erroneously been reported as
<taxonomicName id="4C7A4D5BFF905C73FB63FEC69773C417" box="[1162,1327,301,326]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="moraguesi">
<emphasis id="B90EEACAFF905C73FB63FEC69773C417" box="[1162,1327,301,326]" italics="true" pageId="5" pageNumber="24">M. moraguesi</emphasis>
</taxonomicName>
in
<collectingCountry id="F36D7648FF905C73FAB0FEC697C0C416" box="[1369,1436,301,327]" name="Japan" pageId="5" pageNumber="24">Japan</collectingCountry>
(
<bibRefCitation id="EFEB4B29FF905C73FF76FEB9936FC43D" author="Hiura" box="[159,307,338,364]" pageId="5" pageNumber="27" refString="Hiura, I. (1959) Contribution to the knowledge of the Anthocoridae from Japan and its adjacent territories (Hemiptera- Heteroptera) 1. Bulletin of the Osaka Museum of Natural History, 11, 1 - 10." type="journal article" year="1959">Hiura 1959</bibRefCitation>
;
<bibRefCitation id="EFEB4B29FF905C73FEA8FEB99058C43D" author="Yasunaga" box="[321,516,338,364]" pageId="5" pageNumber="28" refString="Yasunaga, T. (1993) Anthocoridae. In: Tomokuni, M. (Ed.), A field guide to Japanese Bugs - Terrestrial Heteropterans. Zenkoku Noson Kyoiku Kyokai, Tokyo, pp. 167 - 169, pls. 26 - 28. (in Japanese)" type="book chapter" year="1993">Yasunaga 1993</bibRefCitation>
); these three species are similar in general appearance and may be easily confused with one another. However,
<taxonomicName id="4C7A4D5BFF905C73FD98FE93914CC4C0" box="[625,784,376,401]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="thripodes">
<emphasis id="B90EEACAFF905C73FD98FE93914CC4C0" box="[625,784,376,401]" italics="true" pageId="5" pageNumber="24">M. thripodes</emphasis>
</taxonomicName>
may be easily distinguished from
<taxonomicName id="4C7A4D5BFF905C73FB2BFE939729C4C0" box="[1218,1397,376,401]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF905C73FB2BFE939729C4C0" box="[1218,1397,376,401]" italics="true" pageId="5" pageNumber="24">M. pictipennis</emphasis>
</taxonomicName>
by antennal segment II being slightly longer than the head width across eyes (in
<taxonomicName id="4C7A4D5BFF905C73FBC4FE76968BC4E7" box="[1069,1239,413,438]" class="Insecta" genus="Montandoniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="pictipennis">
<emphasis id="B90EEACAFF905C73FBC4FE76968BC4E7" box="[1069,1239,413,438]" italics="true" pageId="5" pageNumber="24">M. pictipennis</emphasis>
</taxonomicName>
, about 0.8 times as long as head width across eyes), the pale yellow mid tibiae, except for the darkened extreme base (cf. mid tibiae dark brown, except for the pale yellow apical half), and the significantly different shape of the male paramere and female copulatory tube (see
<bibRefCitation id="EFEB4B29FF905C73FD61FDE691ECC776" author="Pluot-Sigwalt" box="[648,944,525,551]" pageId="5" pageNumber="27" refString="Pluot-Sigwalt, D., Streito, J. C. &amp; Matocq, A. (2009) Is Montandoniola moraguesi (Puton, 1896) a mixture of different species? (Hemiptera: Heteroptera: Anthocoridae). Zootaxa, 2208, 25 - 43." type="journal article" year="2009">
Pluot-Sigwalt
<emphasis id="B90EEACAFF905C73FCD8FDE69137C777" box="[817,875,525,550]" italics="true" pageId="5" pageNumber="24">et al.</emphasis>
2009
</bibRefCitation>
, Figs. 33, 36, 39;
<figureCitation id="13412A5DFF905C73FB6BFDE6974CC776" box="[1154,1296,525,551]" captionStart="FIGURES 17 22" captionStartId="4.[151,273,1942,1966]" captionTargetBox="[187,1374,397,1885]" captionTargetId="figure@4.[154,1433,351,1921]" captionTargetPageId="4" captionText="FIGURES 17 22. Male (17 20) and female (21, 22) genitalia of M. pictipennis (17 20), M. thripodes (21) and M. kerzhneri sp. nov. (22). 17, Pygophore with paramere, dorsal view; 18, paramere, dorsal view; 19, 20, ditto, two different orientations; 21, 22, copulatory tube, dorsal view. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/196534/files/figure.png" pageId="5" pageNumber="24">Figs. 1721</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3606553FF905C73FF2CFDD9907EC7ED" pageId="5" pageNumber="24" type="biology_ecology">
<paragraph id="8BC536D8FF905C73FF2CFDD9907EC7ED" blockId="5.[151,1437,152,700]" pageId="5" pageNumber="24">
<emphasis id="B90EEACAFF905C73FF2CFDD99370C71D" bold="true" box="[197,300,562,588]" pageId="5" pageNumber="24">Biology.</emphasis>
No information on biology of this species is currently available. In our field observations on Amami-Ôshima and Okinawa Islands, most specimens were found to be associated with leaf-curl galls induced by the Cuban laurel thrips,
<taxonomicName id="4C7A4D5BFF905C73FDA8FD96964AC7C6" authority="Marchal, 1908" authorityName="Marchal" authorityYear="1908" box="[577,1046,637,663]" class="Insecta" family="Phlaeothripidae" genus="Gynaikothrips" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="species" species="ficorum">
<emphasis id="B90EEACAFF905C73FDA8FD969112C7C7" box="[577,846,637,662]" italics="true" pageId="5" pageNumber="24">Gynaikothrips ficorum</emphasis>
(Marchal, 1908)
</taxonomicName>
(
<taxonomicName id="4C7A4D5BFF905C73FBCFFD969691C7C6" box="[1062,1229,637,663]" class="Insecta" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="order">Thysanoptera</taxonomicName>
:
<taxonomicName id="4C7A4D5BFF905C73FB30FD9697C8C7C6" box="[1241,1428,637,663]" class="Insecta" family="Phlaeothripidae" kingdom="Animalia" order="Thysanoptera" pageId="5" pageNumber="24" phylum="Arthropoda" rank="family">Phlaeothripidae</taxonomicName>
) on
<taxonomicName id="4C7A4D5BFF905C73FF52FD499042C7ED" box="[187,542,674,700]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="24" phylum="Mollusca" rank="species" species="microcarpa">
<emphasis id="B90EEACAFF905C73FF52FD4993D0C7EA" box="[187,396,674,699]" italics="true" pageId="5" pageNumber="24">Ficus microcarpa</emphasis>
(Moraceae)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>