571 lines
85 KiB
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571 lines
85 KiB
XML
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<mods:title id="A57F35CEC3D412D5F3AE6A6639DEA460">Taxonomic revision of the genus Oplognathus MacLeay, 1819 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)</mods:title>
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<mods:namePart id="2D9CF0E71C5876EC1F4E28A8B2B878FC">Carvalho, Tamara G.</mods:namePart>
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<mods:affiliation id="38CB6A6AB5C09AC791F160B1B9947DC9">Programa de Pós-Graduação em Entomologia da UFRPE, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, 52171 - 900, Recife, PE, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="C940902A2BA7E27C528F040B7E9C3D99" type="email">lunatamaracarvalho@gmail.com</mods:nameIdentifier>
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<mods:namePart id="14400736A613943936B101260327E22C">Seidel, Matthias</mods:namePart>
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<mods:affiliation id="09512EB8E0861107AE4093B933C59047">Centrum für Naturkunde (CeNak), Universität Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:nameIdentifier id="AB73A59F180879FF8FC02F067872EC0F" type="email">matthias.seidel@uni-hamburg.de;matth.seidel@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5A00C3052D898A9B9B5B08B792264A2E">Grossi, Paschoal C.</mods:namePart>
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<mods:affiliation id="E48A0C065B0ADD834B4F91F50F006531">Programa de Pós-Graduação em Entomologia da UFRPE, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, 52171 - 900, Recife, PE, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="52937D32394609E10709335CB3309815" type="email">paschoalgrossi@gmail.com</mods:nameIdentifier>
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<mods:title id="4EBB219DB7896575213EB8ACDAC93A81">European Journal of Taxonomy</mods:title>
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<mods:date id="2826F90C8E21CC7B488916A294E86591">2021</mods:date>
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<treatment id="03D38784E1574537FDCFF96BFAC4F91D" ID-DOI="http://doi.org/10.5281/zenodo.5572904" ID-GBIF-Taxon="188606702" ID-Zenodo-Dep="5572904" LSID="urn:lsid:plazi:treatment:03D38784E1574537FDCFF96BFAC4F91D" httpUri="http://treatment.plazi.org/id/03D38784E1574537FDCFF96BFAC4F91D" lastPageId="5" lastPageNumber="67" pageId="2" pageNumber="64">
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<subSubSection id="C3606519E1574530FDCFF96BFC43F8ED" box="[580,1008,1782,1810]" pageId="2" pageNumber="64" type="nomenclature">
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<paragraph id="8BC53692E1574530FDCFF96BFC43F8ED" blockId="2.[580,1008,1782,1810]" box="[580,1008,1782,1810]" pageId="2" pageNumber="64">
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<heading id="D08D81FEE1574530FDCFF96BFC43F8ED" box="[580,1008,1782,1810]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="64" reason="2">
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Genus
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<taxonomicName id="4C7A4D11E1574530FD1DF96AFC43F8ED" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[662,1008,1782,1810]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA80E1574530FD1DF96AFC81F8ED" bold="true" box="[662,818,1782,1809]" italics="true" pageId="2" pageNumber="64">Oplognathus</emphasis>
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<bibRefCitation id="EFEB4B63E1574530FCB1F964FC43F8ED" author="MacLeay WS" box="[826,1008,1783,1810]" pageId="2" pageNumber="64" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
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||
</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606519E1574531FF36F8A4FE42FE28" lastPageId="3" lastPageNumber="65" pageId="2" pageNumber="64" type="reference_group">
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<paragraph id="8BC53692E1574530FF36F8A4FD6DF8AF" blockId="2.[189,734,1848,1875]" box="[189,734,1848,1875]" pageId="2" pageNumber="64">
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<treatmentCitationGroup id="AB6A11BCE1574530FF36F8A4FD6DF8AF" box="[189,734,1848,1875]" pageId="2" pageNumber="64">
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<treatmentCitation id="0ADB1083E1574530FF36F8A4FDFDF8AF" author="MacLeay WS" box="[189,590,1848,1875]" page="159" pageId="2" pageNumber="64" year="1819">
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<taxonomicName id="4C7A4D11E1574530FF36F8A4FDFDF8AF" authority="MacLeay, 1819: 159" authorityName="MacLeay" authorityPageNumber="159" authorityYear="1819" box="[189,590,1848,1875]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA80E1574530FF36F8A4FEE6F8AE" box="[189,341,1848,1874]" italics="true" pageId="2" pageNumber="64">Oplognathus</emphasis>
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||
<bibRefCitation id="EFEB4B63E1574530FED7F8A5FDFDF8AF" author="MacLeay WS" box="[348,590,1848,1875]" pageId="2" pageNumber="64" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819: 159</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
(
|
||
<figureCitation id="13412A17E1574530FDD5F8A4FD62F8AF" box="[606,721,1848,1875]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="2" pageNumber="64">Fig. 1a–g</figureCitation>
|
||
).
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8BC53692E1574531FF36F8E6FAC4FEBB" blockId="2.[189,1398,1914,1976]" lastBlockId="3.[185,1399,265,468]" lastPageId="3" lastPageNumber="65" pageId="2" pageNumber="64">
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||
<treatmentCitationGroup id="AB6A11BCE1574531FF36F8E6FAC4FEBB" lastPageId="3" lastPageNumber="65" pageId="2" pageNumber="64">
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||
<taxonomicName id="4C7A4D11E1574530FF36F8E6FEE0F868" authorityName="MacLeay" authorityYear="1819" box="[189,339,1914,1940]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="64" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1574530FF36F8E6FEE0F868" box="[189,339,1914,1940]" italics="true" pageId="2" pageNumber="64">Oplognathus</emphasis>
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||
</taxonomicName>
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||
–
|
||
<treatmentCitation id="0ADB1083E1574530FEF0F8E6FDD7F869" author="Latreille M." box="[379,612,1914,1941]" page="553" pageId="2" pageNumber="64" year="1829">
|
||
<bibRefCitation id="EFEB4B63E1574530FEF0F8E6FDD7F869" author="Latreille M." box="[379,612,1914,1941]" pageId="2" pageNumber="64" refId="ref11186" refString="Latreille M. 1829. Le Regne Animal distribue d'apres son organisation, pour servir de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee, tome IV. Chez Deterville Libraire, Paris. https: // doi. org / 10.5962 / bhl. title. 49223" type="book" year="1829">Latreille 1829: 553</bibRefCitation>
|
||
</treatmentCitation>
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||
(catalogue). —
|
||
<treatmentCitation id="0ADB1083E1574530FCBBF8E6FBA2F868" author="Laporte F." box="[816,1041,1914,1940]" page="124" pageId="2" pageNumber="64" year="1840">
|
||
<bibRefCitation id="EFEB4B63E1574530FCBBF8E6FBA2F868" author="Laporte F." box="[816,1041,1914,1940]" pageId="2" pageNumber="64" refId="ref11167" refString="Laporte F. 1840. Histoire naturelle des Insectes Coleopteres, tome deuxieme. Dumenil Editeur, Paris." type="book" year="1840">Laporte 1840: 124</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
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||
<treatmentCitation id="0ADB1083E1574530FB56F8E6FED3F844" author="Machatschke J. W." page="157" pageId="2" pageNumber="64" year="1970">
|
||
<bibRefCitation id="EFEB4B63E1574530FB56F8E6FED3F844" author="Machatschke J. W." pageId="2" pageNumber="64" pagination="157 - 158" refId="ref11237" refString="Machatschke J. W. 1970. Rutelinae: synonymische bemerkungen. The Entomologist's Monthly Magazine 105: 157 - 158." type="journal article" year="1970">Machatschke 1970: 157</bibRefCitation>
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||
</treatmentCitation>
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||
(synonymy observation); 1972: 5 (catalogue). —
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||
<treatmentCitation id="0ADB1083E1574530FC2DF801FAA3F844" author="Ratcliffe B. C. & Jameson M. L." box="[934,1296,1949,1976]" page="135" pageId="2" pageNumber="64" year="1989">
|
||
<bibRefCitation id="EFEB4B63E1574530FC2DF801FAA3F844" author="Ratcliffe B. C. & Jameson M. L." box="[934,1296,1949,1976]" pageId="2" pageNumber="64" pagination="135 - 144" refId="ref11509" refString="Ratcliffe B. C. & Jameson M. L. 1989. A synopsis of the genus Areoda (Coleoptera: Scarabaeidae: Rutelinae). The Coleopterists Bulletin 43: 135 - 144." type="journal article" year="1989">Ratcliffe & Jameson 1989: 135</bibRefCitation>
|
||
</treatmentCitation>
|
||
(general characterization). —
|
||
<treatmentCitation id="0ADB1083E1564531FE6BFE95FD71FED8" author="Jameson M. L." box="[480,706,265,292]" page="415" pageId="3" pageNumber="65" year="1990">
|
||
<bibRefCitation id="EFEB4B63E1564531FE6BFE95FD71FED8" author="Jameson M. L." box="[480,706,265,292]" pageId="3" pageNumber="65" pagination="377 - 422" refId="ref10997" refString="Jameson M. L. 1990. Revision, phylogeny and biogeography of the genera Parabyrsopolis Ohaus and Viridimicus, new genus (Coleoptera: Scarabaeidae: Rutelinae). The Coleopterists Bulletin 44: 377 - 422." type="journal article" year="1990">Jameson 1990: 415</bibRefCitation>
|
||
</treatmentCitation>
|
||
(phylogeny). —
|
||
<treatmentCitation id="0ADB1083E1564531FC01FE95FBFCFEDF" author="Krajcik M." box="[906,1103,265,291]" page="90" pageId="3" pageNumber="65" year="2007">
|
||
<bibRefCitation id="EFEB4B63E1564531FC01FE95FBFCFEDF" author="Krajcik M." box="[906,1103,265,291]" pageId="3" pageNumber="65" refId="ref11075" refString="Krajcik M. 2007. Anima. X. Rutelinae of the world. Checklist of Scarabaeoidea of the world. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Plzen, Czech Republic." type="book" year="2007">Krajcik 2007: 90</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="0ADB1083E1564531FA87FE95FDB3FEBA" author="Grossi P. C. & Vaz-de-Mello F. Z." page="221" pageId="3" pageNumber="65" year="2015">
|
||
<bibRefCitation id="EFEB4B63E1564531FA87FE95FDB3FEBA" author="Grossi P. C. & Vaz-de-Mello F. Z." pageId="3" pageNumber="65" pagination="221 - 226" refId="ref10842" refString="Grossi P. C. & Vaz-de-Mello F. Z. 2015. Moronius miguelangeli new genus and new species of Areodina from western Brazil (Melolonthidae, Rutelinae, Rutelini). Dugesiana 22: 221 - 226." type="journal article" year="2015">Grossi & Vaz-de-Mello2015: 221</bibRefCitation>
|
||
</treatmentCitation>
|
||
(collecting report).—
|
||
<treatmentCitation id="0ADB1083E1564531FD76FEB0FBF4FEBA" author="Carvalho T. G. & Grossi P. C." box="[765,1095,300,326]" page="369" pageId="3" pageNumber="65" year="2018">
|
||
<bibRefCitation id="EFEB4B63E1564531FD76FEB0FBF4FEBA" author="Carvalho T. G. & Grossi P. C." box="[765,1095,300,326]" pageId="3" pageNumber="65" pagination="369 - 372" refId="ref10745" refString="Carvalho T. G. & Grossi P. C. 2018. Description of a second species of the genus Moronius Grossi & Vazde-Mello, 2015 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini). Zootaxa 4434: 369 - 372. https: // doi. org / 10.11646 / zootaxa. 4434.2.7" type="journal article" year="2018">Carvalho& Grossi2018:369</bibRefCitation>
|
||
</treatmentCitation>
|
||
(general characterization).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF32FED3FE42FE28" blockId="3.[185,1399,265,468]" pageId="3" pageNumber="65">
|
||
<treatmentCitationGroup id="AB6A11BCE1564531FF32FED3FE42FE28" pageId="3" pageNumber="65">
|
||
<taxonomicName id="4C7A4D11E1564531FF32FED3FEECFE95" authorityName="Burmeister" authorityYear="1844" box="[185,351,335,361]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1564531FF32FED3FEECFE95" box="[185,351,335,361]" italics="true" pageId="3" pageNumber="65">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADB1083E1564531FE0BFED3FD33FE95" author="Burmeister H." box="[384,640,335,361]" page="428" pageId="3" pageNumber="65" year="1844">
|
||
<bibRefCitation id="EFEB4B63E1564531FE0BFED3FD33FE95" author="Burmeister H." box="[384,640,335,361]" pageId="3" pageNumber="65" refId="ref10710" refString="Burmeister H. 1844. Handbuch der Entomologie, vol. 4, part 1. TCF Enslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1844">Burmeister 1844: 428</bibRefCitation>
|
||
</treatmentCitation>
|
||
(re-description). —
|
||
<treatmentCitation id="0ADB1083E1564531FCFAFED3FBDEFE96" author="Lacordaire M. T." box="[881,1133,335,362]" page="365" pageId="3" pageNumber="65" year="1856">
|
||
<bibRefCitation id="EFEB4B63E1564531FCFAFED3FBDEFE96" author="Lacordaire M. T." box="[881,1133,335,362]" pageId="3" pageNumber="65" refId="ref11112" refString="Lacordaire M. T. 1856. Histoire naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres propose jusqu'ici dans cet ordre d'insectes, tome III. Librairie Encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8864" type="book" year="1856">Lacordaire 1856: 365</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue). —
|
||
<treatmentCitation id="0ADB1083E1564531FAA0FED3FED2FE71" author="Ohaus F." page="322" pageId="3" pageNumber="65" year="1905">
|
||
<bibRefCitation id="EFEB4B63E1564531FAA0FED3FED2FE71" author="Ohaus F." pageId="3" pageNumber="65" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus 1905: 322</bibRefCitation>
|
||
</treatmentCitation>
|
||
(species description); 1912: 650 (species description); 1914a: 301 (species description); 1915: 257 (general characterization); 1918: 11 (catalogue); 1934: 41 (re-description). —
|
||
<treatmentCitation id="0ADB1083E1564531FB68FE0AFEEDFE28" author="Blackwelder R. E." page="235" pageId="3" pageNumber="65" year="1944">
|
||
<bibRefCitation id="EFEB4B63E1564531FB68FE0AFEEDFE28" author="Blackwelder R. E." pageId="3" pageNumber="65" pagination="189 - 341" refId="ref10657" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1564531FF36FE67FCBAFDBF" pageId="3" pageNumber="65" type="type_taxon">
|
||
<paragraph id="8BC53692E1564531FF36FE67FEEBFDE8" blockId="3.[189,344,506,533]" box="[189,344,506,533]" pageId="3" pageNumber="65">
|
||
<heading id="D08D81FEE1564531FF36FE67FEEBFDE8" bold="true" box="[189,344,506,533]" fontSize="11" level="3" pageId="3" pageNumber="65" reason="3">
|
||
<emphasis id="B90EEA80E1564531FF36FE67FEEBFDE8" bold="true" box="[189,344,506,533]" pageId="3" pageNumber="65">
|
||
<typeStatus id="54C18830E1564531FF36FE67FF49FDE9" box="[189,250,507,533]" pageId="3" pageNumber="65">Type</typeStatus>
|
||
species
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF36FDB5FCBAFDBF" blockId="3.[189,777,553,579]" box="[189,777,553,579]" pageId="3" pageNumber="65">
|
||
<taxonomicName id="4C7A4D11E1564531FF36FDB5FDE8FDBF" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[189,603,553,579]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E1564531FF36FDB5FE2FFDBF" box="[189,412,553,579]" italics="true" pageId="3" pageNumber="65">Oplognathus kirbii</emphasis>
|
||
<bibRefCitation id="EFEB4B63E1564531FE28FDB5FDE8FDBF" author="MacLeay WS" box="[419,603,553,579]" pageId="3" pageNumber="65" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1564531FF36FDF6FC19FCBC" pageId="3" pageNumber="65" type="diagnosis">
|
||
<paragraph id="8BC53692E1564531FF36FDF6FE85FD78" blockId="3.[189,310,618,644]" box="[189,310,618,644]" pageId="3" pageNumber="65">
|
||
<heading id="D08D81FEE1564531FF36FDF6FE85FD78" bold="true" box="[189,310,618,644]" fontSize="11" level="3" pageId="3" pageNumber="65" reason="3">
|
||
<emphasis id="B90EEA80E1564531FF36FDF6FE85FD78" bold="true" box="[189,310,618,644]" pageId="3" pageNumber="65">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF36FD05FC19FCBC" blockId="3.[189,1399,665,832]" pageId="3" pageNumber="65">
|
||
Males with quadrangular, trilobate clypeus (
|
||
<figureCitation id="13412A17E1564531FD5CFD05FCFCFD48" box="[727,847,665,692]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="3" pageNumber="65">Fig. 3a–d</figureCitation>
|
||
), rounded in females (
|
||
<figureCitation id="13412A17E1564531FBEDFD05FAB1FD4F" box="[1126,1282,665,691]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="3" pageNumber="65">Fig. 1b, e, g</figureCitation>
|
||
); clypeus extending beyond labrum in ventral view in both sexes; mandibles with three teeth in incisive area (
|
||
<figureCitation id="13412A17E1564531FF4DFD43FEFCFD06" box="[198,335,735,762]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="3" pageNumber="65">Fig. 4a, e, i</figureCitation>
|
||
); maxillae with six teeth (
|
||
<figureCitation id="13412A17E1564531FD0FFD43FCB8FD06" box="[644,779,735,762]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="3" pageNumber="65">Fig. 4b, f, j</figureCitation>
|
||
); antennae with 10 antennomeres (
|
||
<figureCitation id="13412A17E1564531FB22FD43FAADFD06" box="[1193,1310,735,762]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="3" pageNumber="65">Fig. 3a–d</figureCitation>
|
||
); elytra with 10 striae (
|
||
<figureCitation id="13412A17E1564531FEE7FC9EFE6CFCE1" box="[364,479,770,797]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="3" pageNumber="65">Fig. 1a–g</figureCitation>
|
||
); mesoventral process present (
|
||
<figureCitation id="13412A17E1564531FCC4FC9EFC0EFCE1" box="[847,957,770,797]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="3" pageNumber="65">Fig. 8a–f</figureCitation>
|
||
); parameres asymmetrical (
|
||
<figureCitation id="13412A17E1564531FB74FC9EFADAFCE1" box="[1279,1385,770,797]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="13.[189,232,1958,1984]" captionStartId-1="15.[189,232,1921,1947]" captionStartId-2="17.[189,232,1924,1950]" captionTargetBox-0="[273,1281,606,1893]" captionTargetBox-1="[277,1269,617,1876]" captionTargetBox-2="[270,1309,269,1863]" captionTargetId-0="figure-86@13.[189,1398,583,1920]" captionTargetId-1="figure-111@15.[189,1398,583,1882]" captionTargetId-2="figure-16@17.[243,1344,265,1886]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionTargetPageId-2="17" captionText-0="Fig. 5. Aedeagus in dorsal (a, e) y lateral view (b–c, d, f). a–c. Oplognathus kirbii MacLeay, 1819 (ZMHB). d–f. Oplognathus bahianus Ohaus, 1912, syntype, ♂ (ZMHB). Scale bar = 1 mm." captionText-1="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." captionText-2="Fig. 7. Variation in male genitalia. a–d. Oplognathus kirbii MacLeay, 1819 (a-b: CEMT; c-d: ZMHB). e–h. O. helmenreichi Ohaus, 1905 (ZMHB). i–m. O. bahianus Ohaus, 1912 (CERPE). Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514432" figureDoi-1="http://doi.org/10.5281/zenodo.5514434" figureDoi-2="http://doi.org/10.5281/zenodo.5514436" httpUri-0="https://zenodo.org/record/5514432/files/figure.png" httpUri-1="https://zenodo.org/record/5514434/files/figure.png" httpUri-2="https://zenodo.org/record/5514436/files/figure.png" pageId="3" pageNumber="65">Figs 5–7</figureCitation>
|
||
), each paramere with two apical lobes, and one lateral projection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1564536FF36FCFBFD5CFBB5" lastPageId="4" lastPageNumber="66" pageId="3" pageNumber="65" type="description">
|
||
<paragraph id="8BC53692E1564531FF36FCFBFED8FC7D" blockId="3.[189,363,871,897]" box="[189,363,871,897]" pageId="3" pageNumber="65">
|
||
<heading id="D08D81FEE1564531FF36FCFBFED8FC7D" bold="true" box="[189,363,871,897]" fontSize="11" level="3" pageId="3" pageNumber="65" reason="3">
|
||
<emphasis id="B90EEA80E1564531FF36FCFBFED8FC7D" bold="true" box="[189,363,871,897]" pageId="3" pageNumber="65">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF61FC0AFE99FC4C" blockId="3.[189,1041,918,979]" box="[234,298,918,944]" pageId="3" pageNumber="65">
|
||
<emphasis id="B90EEA80E1564531FF61FC0AFE99FC4C" bold="true" box="[234,298,918,944]" pageId="3" pageNumber="65">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF36FC25FBA2FC2F" blockId="3.[189,1041,918,979]" box="[189,1041,953,979]" pageId="3" pageNumber="65">
|
||
SIZE. Total length:
|
||
<quantity id="4C829B77E1564531FE13FC25FDF5FC2F" box="[408,582,953,979]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.2399999999999998" metricValueMin="1.7399999999999998" pageId="3" pageNumber="65" unit="mm" value="19.9" valueMax="22.4" valueMin="17.4">17.4–22.4 mm</quantity>
|
||
; width across prothorax: 8.0–
|
||
<quantity id="4C829B77E1564531FC29FC25FBBFFC2F" box="[930,1036,953,979]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="3" pageNumber="65" unit="mm" value="11.4">11.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF36FC66FD13FBA0" blockId="3.[189,1398,1018,1116]" pageId="3" pageNumber="65">
|
||
COLOUR. General aspect light yellow varying from yellowish to reddish metallic green, with golden and green reflexes; ventral surface metallic green; pronotum sometimes with black maculae; elytral margins metallic green or copper (
|
||
<figureCitation id="13412A17E1564531FE60FBDDFD21FBA0" box="[491,658,1089,1116]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="3" pageNumber="65">Fig. 1a, c–d, f</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564531FF36FB1FFDEAFA00" blockId="3.[189,1399,1154,1533]" pageId="3" pageNumber="65">
|
||
HEAD (
|
||
<figureCitation id="13412A17E1564531FE99FB1EFEEEFB60" box="[274,349,1154,1181]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="3" pageNumber="65">Fig. 3</figureCitation>
|
||
). Reddish brown and/or metallic green. Clypeus subquadrate, surface rugose, setose, apex trilobate, apical and lateral margins elevated; clypeus produced beyond labrum in ventral view. Frontoclypeal suture usually distinct, straight or sinuous. Frons predominantly rectangular, surface glabrous, distinctly punctate, punctures dense; ocular canthus intruding a third of the eye’s length, with scattered yellow setae. Labrum transverse, widely bilobed, setose, never visible from above. Mandibles laminar, rounded, intimately joined to labrum, with three incisive teeth (
|
||
<figureCitation id="13412A17E1564531FB5AFAAEFAD4FAB1" box="[1233,1383,1330,1357]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="3" pageNumber="65">Fig. 4a, e, i</figureCitation>
|
||
). Maxillae setose, galea with six teeth, palpi 4-segmented, last palpomere longer than other three combined, varying from cylindrical to fusiform (
|
||
<figureCitation id="13412A17E1564531FCA1FAE4FBAFFA6F" box="[810,1052,1400,1427]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="3" pageNumber="65">Fig. 4b–c, f–g, j, l</figureCitation>
|
||
). Mentum sinuous, bilobed or notched, ventral surface setose; labial palpus inserted dorsally on mentum, 3-segmented; last palpomere enlarged (
|
||
<figureCitation id="13412A17E1564531FE30FA23FDE8FA25" box="[443,603,1471,1497]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="3" pageNumber="65">Fig. 4d, h, m</figureCitation>
|
||
). Antenna with 10 antennomeres, antennal club 1.3 to 1.8× longer than antennomeres 2–7 combined.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1564536FF36F9B8FE0CFE0F" blockId="3.[188,1400,1571,2021]" lastBlockId="4.[189,1400,265,499]" lastPageId="4" lastPageNumber="66" pageId="3" pageNumber="65">
|
||
THORAX. Pronotum convex, transverse, about 1.8 times wider than longer, beaded; pronotal bead incomplete posteriorly at middle; surface varying from sparse to densely punctate; punctures small to moderate; maculae varying from two distinct anterolateral spots to two complete elongated maculae; some specimens with spots and maculae obsolete or absent. Each elytron with 10 distinctly punctate striae, punctures fine to moderate, interstriae almost smooth, punctures visible from 10× of magnification; humeri rounded, smooth; elytral apex straight. Hind wings full developed with a group of setae at AA 1+2, setae long and fine, longer than width of first axillar sclerite; AP 3+4 softly setose at base, with fine and minute setae; R 3+4 sclerotized, disc with a distinct row of short setae. Mesoventral process acute, elongated, but never extended beyond anterior procoxal base (
|
||
<figureCitation id="13412A17E1564531FBDEF8A1FB2CF8AB" box="[1109,1183,1853,1879]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="3" pageNumber="65">Fig. 8</figureCitation>
|
||
). Metafemur with anterior and posterior margins rounded, distinctly developed that other femora, with at least twice the mesofemoral width. Protibia tridentate, teeth reducing in size from apex to base, proximal tooth sometimes obsolete; inner apex with one tiny spur; ventral surface distinctly dilated, convex. Mesotibia with surface rugose, and two oblique carinae: proximal carina with 2–7 spinules, and distal carina with 4–9 spinules, distal carina longer; mesotibial apex truncate with 7–13 spinules and two spurs, inner spur slightly longer. Metatibia wider than mesotibia, with inner margin distinctly rounded, apical spinules varying from 14–22; spurs distinctly short, and flattened, apex less acute than spinules; proximal carina with 2–8 spinules and distal carina with 6–11 spinules. Protarsomere V thickened; inner claw stronger with a basal obtuse tooth. Meso- and metatarsomere ornamented with lateral setae, and inner apical spinules. Meso- and metatarsal claws curved, simple, unequal in thickness; unguitractor plate present with two apical setae.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FD86FD30FD43" blockId="4.[188,1399,538,703]" pageId="4" pageNumber="66">
|
||
ABDOMEN. Pygidium flat to weakly convex, subtrapezoidal, transverse, surface rugose and setose; longer setae often on posterior sides. Ventrites metallic green, slightly convex, surface weakly punctate, punctures sparse, with scattered setae, sternites V and VI about twice as long as sternite IV. Parameres asymmetrical, each paramere with 1–2 distal lobes, left lateral expanded with a long projection, apical bifurcation wide, V-shaped (
|
||
<figureCitation id="13412A17E1514536FD81FD38FDC6FD43" box="[522,629,676,703]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="13.[189,232,1958,1984]" captionStartId-1="15.[189,232,1921,1947]" captionStartId-2="17.[189,232,1924,1950]" captionTargetBox-0="[273,1281,606,1893]" captionTargetBox-1="[277,1269,617,1876]" captionTargetBox-2="[270,1309,269,1863]" captionTargetId-0="figure-86@13.[189,1398,583,1920]" captionTargetId-1="figure-111@15.[189,1398,583,1882]" captionTargetId-2="figure-16@17.[243,1344,265,1886]" captionTargetPageId-0="13" captionTargetPageId-1="15" captionTargetPageId-2="17" captionText-0="Fig. 5. Aedeagus in dorsal (a, e) y lateral view (b–c, d, f). a–c. Oplognathus kirbii MacLeay, 1819 (ZMHB). d–f. Oplognathus bahianus Ohaus, 1912, syntype, ♂ (ZMHB). Scale bar = 1 mm." captionText-1="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." captionText-2="Fig. 7. Variation in male genitalia. a–d. Oplognathus kirbii MacLeay, 1819 (a-b: CEMT; c-d: ZMHB). e–h. O. helmenreichi Ohaus, 1905 (ZMHB). i–m. O. bahianus Ohaus, 1912 (CERPE). Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5514432" figureDoi-1="http://doi.org/10.5281/zenodo.5514434" figureDoi-2="http://doi.org/10.5281/zenodo.5514436" httpUri-0="https://zenodo.org/record/5514432/files/figure.png" httpUri-1="https://zenodo.org/record/5514434/files/figure.png" httpUri-2="https://zenodo.org/record/5514436/files/figure.png" pageId="4" pageNumber="66">Figs 5–7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF61FD7AFE5FFCFC" blockId="4.[189,782,741,803]" box="[234,492,741,768]" pageId="4" pageNumber="66">
|
||
<emphasis id="B90EEA80E1514536FF61FD7AFEF6FCFC" bold="true" box="[234,325,742,768]" pageId="4" pageNumber="66">Female</emphasis>
|
||
(
|
||
<figureCitation id="13412A17E1514536FEDEFD79FE50FCFC" box="[341,483,741,768]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="4" pageNumber="66">Fig. 1b, e, g</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FC94FCBDFCDF" blockId="4.[189,782,741,803]" box="[189,782,776,803]" pageId="4" pageNumber="66">Similar to male, differing in the following aspects:</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FCD5FBA3FC98" blockId="4.[189,1040,841,868]" box="[189,1040,841,868]" pageId="4" pageNumber="66">
|
||
SIZE. Total length:
|
||
<quantity id="4C829B77E1514536FE13FCD5FDF5FC98" box="[408,582,841,868]" metricMagnitude="-2" metricUnit="m" metricValue="1.9649999999999999" metricValueMax="2.21" metricValueMin="1.72" pageId="4" pageNumber="66" unit="mm" value="19.65" valueMax="22.1" valueMin="17.2">17.2–22.1 mm</quantity>
|
||
; width across prothorax:
|
||
<quantity id="4C829B77E1514536FCE5FCD5FBBFFC98" box="[878,1036,841,868]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" metricValueMax="11.1" metricValueMin="7.5" pageId="4" pageNumber="66" unit="mm" value="9.3" valueMax="11.1" valueMin="7.5">7.5–11.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FC16FBF8FC58" blockId="4.[189,1099,906,932]" box="[189,1099,906,932]" pageId="4" pageNumber="66">HEAD. Apex rounded, never with projections; antennal club distinctly smaller.</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FC57FA89FBFB" blockId="4.[189,1398,971,1032]" pageId="4" pageNumber="66">THORAX. Pronotum more densely punctate. Elytra more convex, with dilated external margins, and elytral epipleuron wider in ventral view; legs thinner, anterior claws subequal, simple, not dilated.</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FBB3FD5CFBB5" blockId="4.[189,751,1070,1097]" box="[189,751,1070,1097]" pageId="4" pageNumber="66">ABDOMEN. Sternum and pygidium more convex.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1514536FF36FBF3FE2AFB01" pageId="4" pageNumber="66" type="distribution">
|
||
<paragraph id="8BC53692E1514536FF36FBF3FEE5FB75" blockId="4.[189,342,1135,1161]" box="[189,342,1135,1161]" pageId="4" pageNumber="66">
|
||
<heading id="D08D81FEE1514536FF36FBF3FEE5FB75" bold="true" box="[189,342,1135,1161]" fontSize="11" level="3" pageId="4" pageNumber="66" reason="3">
|
||
<emphasis id="B90EEA80E1514536FF36FBF3FEE5FB75" bold="true" box="[189,342,1135,1161]" pageId="4" pageNumber="66">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FB01FE2AFB01" blockId="4.[189,1398,1181,1277]" pageId="4" pageNumber="66">
|
||
<emphasis id="B90EEA80E1514536FF36FB01FEB9FB4B" bold="true" box="[189,266,1181,1207]" pageId="4" pageNumber="66">
|
||
<collectingCountry id="F36D7602E1514536FF36FB01FEB9FB4B" box="[189,266,1181,1207]" name="Brazil" pageId="4" pageNumber="66">Brazil</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion id="49BEF870E1514536FEAFFB01FEDBFB4B" box="[292,360,1181,1207]" country="Brazil" name="Bahia" pageId="4" pageNumber="66">Bahia</collectingRegion>
|
||
: Condeúba;
|
||
<collectingRegion id="49BEF870E1514536FD9DFB01FD7AFB4B" box="[534,713,1181,1207]" country="Brazil" name="Espirito Santo" pageId="4" pageNumber="66">Espírito Santo</collectingRegion>
|
||
: Palmital;
|
||
<collectingRegion id="49BEF870E1514536FCE9FB01FBB8FB4B" box="[866,1035,1181,1207]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="66">Minas Gerais</collectingRegion>
|
||
: Águas Vermelhas, Berizal, Manhumirim, Grão Mogol, Leme do Prado;
|
||
<collectingRegion id="49BEF870E1514536FD4BFB5CFCA3FB26" box="[704,784,1216,1242]" country="Brazil" name="Parana" pageId="4" pageNumber="66">Paraná</collectingRegion>
|
||
: Curitiba;
|
||
<collectingRegion id="49BEF870E1514536FC06FB5CFB87FB26" box="[909,1076,1216,1242]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="66">Rio de Janeiro</collectingRegion>
|
||
:
|
||
<collectingRegion id="49BEF870E1514536FBC8FB5CFB5CFB26" box="[1091,1263,1216,1242]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="66">Rio de Janeiro</collectingRegion>
|
||
;
|
||
<collectingRegion id="49BEF870E1514536FB73FB5CFADEFB26" box="[1272,1389,1216,1242]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="66">São Paulo</collectingRegion>
|
||
:
|
||
<collectingRegion id="49BEF870E1514536FF36FB7FFE80FB01" box="[189,307,1251,1277]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="66">São Paulo</collectingRegion>
|
||
(Fig. 9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1514537FF36FAB8FE66FBBF" lastPageId="5" lastPageNumber="67" pageId="4" pageNumber="66" type="discussion">
|
||
<paragraph id="8BC53692E1514536FF36FAB8FE6BFAC1" blockId="4.[189,472,1315,1342]" box="[189,472,1315,1342]" pageId="4" pageNumber="66">
|
||
<heading id="D08D81FEE1514536FF36FAB8FE6BFAC1" bold="true" box="[189,472,1315,1342]" fontSize="11" level="3" pageId="4" pageNumber="66" reason="3">
|
||
<emphasis id="B90EEA80E1514536FF36FAB8FE6BFAC1" bold="true" box="[189,472,1315,1342]" pageId="4" pageNumber="66">Nomenclatural history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1514536FF36FACDFBF6F820" blockId="4.[188,1401,1361,2012]" pageId="4" pageNumber="66">
|
||
<taxonomicName id="4C7A4D11E1514536FF36FACDFEE6FA97" authorityName="MacLeay" authorityYear="1819" box="[189,341,1361,1387]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FF36FACDFEE6FA97" box="[189,341,1361,1387]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EFEB4B63E1514536FDB1FACEFCB2FA90" author="MacLeay WS" box="[570,769,1362,1388]" pageId="4" pageNumber="66" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay (1819)</bibRefCitation>
|
||
without ‘H’ at the beginning of the name, but later
|
||
<bibRefCitation id="EFEB4B63E1514536FF36FAE8FE2BFA72" author="Burmeister H." box="[189,408,1396,1422]" pageId="4" pageNumber="66" refId="ref10710" refString="Burmeister H. 1844. Handbuch der Entomologie, vol. 4, part 1. TCF Enslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1844">Burmeister (1844)</bibRefCitation>
|
||
redescribed the genus as “
|
||
<taxonomicName id="4C7A4D11E1514536FD6DFAE8FC3FFA72" authorityName="Burmeister" authorityYear="1844" box="[742,908,1396,1422]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FD6DFAE8FC3FFA72" box="[742,908,1396,1422]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
”, justifying that it would be the correct spelling for the name.
|
||
<bibRefCitation id="EFEB4B63E1514536FE43FA0BFD2EFA4E" author="Lacordaire M. T." box="[456,669,1431,1458]" pageId="4" pageNumber="66" refId="ref11112" refString="Lacordaire M. T. 1856. Histoire naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres propose jusqu'ici dans cet ordre d'insectes, tome III. Librairie Encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8864" type="book" year="1856">Lacordaire (1856)</bibRefCitation>
|
||
also pointed the “error” of MacLeay, and uses
|
||
<taxonomicName id="4C7A4D11E1514536FB5AFA0BFAC4FA4D" authorityName="Burmeister" authorityYear="1844" box="[1233,1399,1431,1457]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FB5AFA0BFAC4FA4D" box="[1233,1399,1431,1457]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
as a valid name, indicating “
|
||
<emphasis id="B90EEA80E1514536FD8DFA25FD29FA2F" box="[518,666,1465,1491]" italics="true" pageId="4" pageNumber="66">Aplognathus</emphasis>
|
||
” as original wrong spelling, but not specifying the cause of the error. Since then, the name
|
||
<taxonomicName id="4C7A4D11E1514536FD82FA40FD1CFA0A" authorityName="Burmeister" authorityYear="1844" box="[521,687,1500,1526]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FD82FA40FD1CFA0A" box="[521,687,1500,1526]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
has been widely used, including the description of the new species, generating more synonyms, such as
|
||
<taxonomicName id="4C7A4D11E1514536FD58FA63FC5DF9E5" authorityName="Ohaus" authorityYear="1912" box="[723,1006,1535,1561]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E1514536FD58FA63FC5DF9E5" box="[723,1006,1535,1561]" italics="true" pageId="4" pageNumber="66">Hoplognathus bahianus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A4D11E1514536FBA2FA63FADCF9E5" authorityName="Ohaus" authorityYear="1905" box="[1065,1391,1535,1561]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E1514536FBA2FA63FADCF9E5" box="[1065,1391,1535,1561]" italics="true" pageId="4" pageNumber="66">Hoplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
. Only
|
||
<bibRefCitation id="EFEB4B63E1514536FF75F9BDFE5FF9C0" author="Machatschke J. W." box="[254,492,1569,1596]" pageId="4" pageNumber="66" pagination="157 - 158" refId="ref11237" refString="Machatschke J. W. 1970. Rutelinae: synonymische bemerkungen. The Entomologist's Monthly Magazine 105: 157 - 158." type="journal article" year="1970">Machatschke (1970)</bibRefCitation>
|
||
comments that
|
||
<taxonomicName id="4C7A4D11E1514536FD28F9BDFCFAF9C7" authorityName="Burmeister" authorityYear="1844" box="[675,841,1569,1595]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FD28F9BDFCFAF9C7" box="[675,841,1569,1595]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
is a synonym, but when using
|
||
<taxonomicName id="4C7A4D11E1514536FB27F9BDFAF7F9C7" authorityName="MacLeay" authorityYear="1819" box="[1196,1348,1569,1595]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FB27F9BDFAF7F9C7" box="[1196,1348,1569,1595]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
, the spelling appears as “
|
||
<emphasis id="B90EEA80E1514536FE27F9D8FDE1F9A2" box="[428,594,1604,1630]" italics="true" pageId="4" pageNumber="66">Oplongnathus</emphasis>
|
||
”. Already in his catalogue,
|
||
<bibRefCitation id="EFEB4B63E1514536FC18F9D8FB31F9A2" author="Machatschke J. W." box="[915,1154,1604,1630]" pageId="4" pageNumber="66" pagination="1 - 361" refId="ref11259" refString="Machatschke J. W. 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361." type="journal article" year="1972">Machatschke (1972)</bibRefCitation>
|
||
cites “
|
||
<taxonomicName id="4C7A4D11E1514536FB59F9D8FAD9F9A2" authorityName="MacLeay" authorityYear="1819" box="[1234,1386,1604,1630]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FB59F9D8FAD9F9A2" box="[1234,1386,1604,1630]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
” using the original spelling, which proves to be a typing error in the previous publications.
|
||
<bibRefCitation id="EFEB4B63E1514536FB7AF9FBFECAF958" author="Ratcliffe B. C. & Jameson M. L." pageId="4" pageNumber="66" pagination="135 - 144" refId="ref11509" refString="Ratcliffe B. C. & Jameson M. L. 1989. A synopsis of the genus Areoda (Coleoptera: Scarabaeidae: Rutelinae). The Coleopterists Bulletin 43: 135 - 144." type="journal article" year="1989">Ratcliffe & Jameson (1989)</bibRefCitation>
|
||
note this fact and show the correct use “
|
||
<taxonomicName id="4C7A4D11E1514536FCEBF915FBDCF958" authority="MacLeay" authorityName="MacLeay" authorityYear="1819" box="[864,1135,1673,1700]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FCEBF915FC4BF95F" box="[864,1016,1673,1699]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
MacLeay
|
||
</taxonomicName>
|
||
(not
|
||
<taxonomicName id="4C7A4D11E1514536FB3BF915FAE5F95F" authorityName="Burmeister" authorityYear="1844" box="[1200,1366,1673,1699]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FB3BF915FAE5F95F" box="[1200,1366,1673,1699]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
as in
|
||
<bibRefCitation id="EFEB4B63E1514536FF54F930FE1AF93A" author="Burmeister H." box="[223,425,1708,1734]" pageId="4" pageNumber="66" refId="ref10710" refString="Burmeister H. 1844. Handbuch der Entomologie, vol. 4, part 1. TCF Enslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1844">Burmeister 1844</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB4B63E1514536FE3CF930FDFAF93A" author="Ohaus F." box="[439,585,1708,1734]" pageId="4" pageNumber="66" pagination="1 - 241" refId="ref11437" refString="Ohaus F. 1918. Scarabaeidae: Euchirinae, Phaenomerinae, Rutelinae. In: Junk W. & Schenkling S. (eds) Coleopterorum Catalogus 66: 1 - 241. W. Junk, Berlin." type="journal article" year="1918">Ohaus 1918</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEB4B63E1514536FDDCF930FD26F93A" author="Ohaus F." box="[599,661,1708,1734]" pageId="4" pageNumber="66" pagination="1 - 172" refId="ref11476" refString="Ohaus F. 1934. Coleoptera Lamelicornia fam. Scarabaeidae, sub. Rutelinae. In: Wytsman P. (ed.) Genera Insectorum 199 a: 1 - 172." type="book chapter" year="1934">1934</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB4B63E1514536FD2FF930FC31F93A" author="Blackwelder R. E." box="[676,898,1708,1734]" pageId="4" pageNumber="66" pagination="189 - 341" refId="ref10657" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944</bibRefCitation>
|
||
)”, because according to Article 23 (ICZN 1999), the Principle of Priority, the oldest name should be used. In the original description,
|
||
<bibRefCitation id="EFEB4B63E1514536FA81F953FEBFF8F0" author="MacLeay WS" pageId="4" pageNumber="66" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay (1819)</bibRefCitation>
|
||
does not detail the etymology of
|
||
<taxonomicName id="4C7A4D11E1514536FD2DF96DFC8DF8F7" authorityName="MacLeay" authorityYear="1819" box="[678,830,1777,1803]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FD2DF96DFC8DF8F7" box="[678,830,1777,1803]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
, but the Hoplo prefix, from the Greek Hoplon, means ‘any tool or implement of armour and shield’ while Gnathus, from the Greek, means ‘mandible, mouth’, possibly due to the distinct shape of the clypeus of the males. Perhaps, this explains the insistence of later authors (
|
||
<bibRefCitation id="EFEB4B63E1514536FE7FF8C5FD0EF888" author="Burmeister H." box="[500,701,1881,1908]" pageId="4" pageNumber="66" refId="ref10710" refString="Burmeister H. 1844. Handbuch der Entomologie, vol. 4, part 1. TCF Enslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1844">Burmeister 1844</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB4B63E1514536FD43F8C5FC3AF888" author="Lacordaire M. T." box="[712,905,1881,1908]" pageId="4" pageNumber="66" refId="ref11112" refString="Lacordaire M. T. 1856. Histoire naturelle des Insectes. Genera des Coleopteres ou expose methodique et critique de tous les genres propose jusqu'ici dans cet ordre d'insectes, tome III. Librairie Encyclopedique de Roret, Paris. https: // doi. org / 10.5962 / bhl. title. 8864" type="book" year="1856">Lacordaire 1856</bibRefCitation>
|
||
) to use the spelling
|
||
<taxonomicName id="4C7A4D11E1514536FBE5F8C5FAA7F88F" authorityName="Burmeister" authorityYear="1844" box="[1134,1300,1881,1907]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FBE5F8C5FAA7F88F" box="[1134,1300,1881,1907]" italics="true" pageId="4" pageNumber="66">Hoplognathus</emphasis>
|
||
</taxonomicName>
|
||
, instead of
|
||
<taxonomicName id="4C7A4D11E1514536FF56F8E0FEC6F86A" authorityName="MacLeay" authorityYear="1819" box="[221,373,1916,1942]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FF56F8E0FEC6F86A" box="[221,373,1916,1942]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
, as correct. However, as there is no written indication of this, and the addition of ‘H’ is not a clear, obvious and necessary correction of the name,
|
||
<taxonomicName id="4C7A4D11E1514536FCFBF803FBBBF845" authorityName="MacLeay" authorityYear="1819" box="[880,1032,1951,1977]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1514536FCFBF803FBBBF845" box="[880,1032,1951,1977]" italics="true" pageId="4" pageNumber="66">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
must be maintained, following the original description and not its subsequent spelling (ICZN 1999, Art. 33).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1504537FF36FE95FAC4FE95" blockId="5.[187,1399,265,362]" pageId="5" pageNumber="67">
|
||
Although Burmeister corrected the name of the genus, this does not mean that Ohaus intended to describe
|
||
<taxonomicName id="4C7A4D11E1504537FF30FEB0FE7AFEBA" authorityName="Ohaus" authorityYear="1912" box="[187,457,300,326]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E1504537FF30FEB0FE7AFEBA" box="[187,457,300,326]" italics="true" pageId="5" pageNumber="67">Oplognathus bahianus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A4D11E1504537FD80FEB0FCF6FEBA" authorityName="Ohaus" authorityYear="1905" box="[523,837,300,326]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E1504537FD80FEB0FCF6FEBA" box="[523,837,300,326]" italics="true" pageId="5" pageNumber="67">Oplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
in a genus other than
|
||
<taxonomicName id="4C7A4D11E1504537FBD6FEB0FB46FEBA" authorityName="MacLeay" authorityYear="1819" box="[1117,1269,300,326]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1504537FBD6FEB0FB46FEBA" box="[1117,1269,300,326]" italics="true" pageId="5" pageNumber="67">Oplognathus</emphasis>
|
||
</taxonomicName>
|
||
. Thus, the author’s name is written here without brackets, since the species is basically in its original combination.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1504537FF36FE0DFD0BFD5D" blockId="5.[189,1399,401,673]" pageId="5" pageNumber="67">
|
||
<taxonomicName id="4C7A4D11E1504537FF36FE0DFE2DFE57" authorityName="MacLeay" authorityYear="1819" box="[189,414,401,427]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E1504537FF36FE0DFE2DFE57" box="[189,414,401,427]" italics="true" pageId="5" pageNumber="67">Oplognathus kirbii</emphasis>
|
||
</taxonomicName>
|
||
was cited by
|
||
<bibRefCitation id="EFEB4B63E1504537FDCCFE0DFD4AFE57" author="Laporte F." box="[583,761,401,427]" pageId="5" pageNumber="67" refId="ref11167" refString="Laporte F. 1840. Histoire naturelle des Insectes Coleopteres, tome deuxieme. Dumenil Editeur, Paris." type="book" year="1840">Laporte (1840)</bibRefCitation>
|
||
as
|
||
<taxonomicName id="4C7A4D11E1504537FCACFE0DFBBFFE57" authorityName="MacLeay" authorityYear="1819" box="[807,1036,401,427]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbyi">
|
||
<emphasis id="B90EEA80E1504537FCACFE0DFBBFFE57" box="[807,1036,401,427]" italics="true" pageId="5" pageNumber="67">Oplognathus kirbyi</emphasis>
|
||
</taxonomicName>
|
||
and since then, both spellings were widely used, in addition to
|
||
<taxonomicName id="4C7A4D11E1504537FDC7FE28FC8FFE32" box="[588,828,436,462]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E1504537FDC7FE28FC8FFE32" box="[588,828,436,462]" italics="true" pageId="5" pageNumber="67">Hoplognathus kirbii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A4D11E1504537FCF1FE28FBDDFE32" authorityName="McLeay" box="[890,1134,436,462]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbyi">
|
||
<emphasis id="B90EEA80E1504537FCF1FE28FBDDFE32" box="[890,1134,436,462]" italics="true" pageId="5" pageNumber="67">Hoplognathus kirbyi</emphasis>
|
||
</taxonomicName>
|
||
, it is common to find four spellings for this species throughout the literature. There are no details for replacing ‘i’ with ‘y’, but we can suppose
|
||
<bibRefCitation id="EFEB4B63E1504537FE33FE67FDDFFDE9" author="Laporte F." box="[440,620,507,533]" pageId="5" pageNumber="67" refId="ref11167" refString="Laporte F. 1840. Histoire naturelle des Insectes Coleopteres, tome deuxieme. Dumenil Editeur, Paris." type="book" year="1840">Laporte (1840)</bibRefCitation>
|
||
to make this change on the assumption that MacLeay intended to honour the naturalist William Kirby. ICZN (1999, Article 58.2) states that the use of ‘i’ and ‘y’ are ‘variant spellings’ deemed to be identical. However, as
|
||
<bibRefCitation id="EFEB4B63E1504537FCC2FDDDFBBEFDA7" author="MacLeay WS" box="[841,1037,577,603]" pageId="5" pageNumber="67" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay (1819)</bibRefCitation>
|
||
makes the description with no detail about etymology, it is prudent to keep the original spelling
|
||
<taxonomicName id="4C7A4D11E1504537FC4BFDF8FB2CFD82" authorityName="MacLeay" authorityYear="1819" box="[960,1183,612,638]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E1504537FC4BFDF8FB2CFD82" box="[960,1183,612,638]" italics="true" pageId="5" pageNumber="67">Oplognathus kirbii</emphasis>
|
||
</taxonomicName>
|
||
and not to use the subsequent spellings (ICZN 1999, Art. 33).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1504537FF36FD55FE66FBBF" blockId="5.[187,1399,713,1091]" pageId="5" pageNumber="67">
|
||
<taxonomicName id="4C7A4D11E1504537FF36FD55FE41FD1F" authorityName="Ohaus" authorityYear="1905" box="[189,498,713,739]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E1504537FF36FD55FE41FD1F" box="[189,498,713,739]" italics="true" pageId="5" pageNumber="67">Oplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
was described for the first time based on a single specimen in the collection of W.J.C. Weber by
|
||
<bibRefCitation id="EFEB4B63E1504537FE0CFD70FD94FCFB" author="Ohaus F." box="[391,551,748,775]" pageId="5" pageNumber="67" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus (1905)</bibRefCitation>
|
||
with this spelling (for the specific epithet) and the locality
|
||
<collectingRegion id="49BEF870E1504537FB58FD70FAC1FCFA" box="[1235,1394,748,774]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="5" pageNumber="67">Buenos Aires</collectingRegion>
|
||
,
|
||
<collectingCountry id="F36D7602E1504537FF36FC93FE85FCD5" box="[189,310,783,809]" name="Argentina" pageId="5" pageNumber="67">Argentina</collectingCountry>
|
||
. Later, Ohaus wrote that he found (
|
||
<bibRefCitation id="EFEB4B63E1504537FD59FC93FC2FFCD5" author="Ohaus F." box="[722,924,783,809]" pageId="5" pageNumber="67" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a: 22</bibRefCitation>
|
||
) several specimens of this species in the Museum of Vienna, but he described them as a variety “
|
||
<taxonomicName id="4C7A4D11E1504537FCBDFCAEFC45FCB0" authorityName="Ohaus" authorityYear="1914" box="[822,1014,818,844]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
var.
|
||
<emphasis id="B90EEA80E1504537FCECFCAEFC45FCB0" box="[871,1014,818,844]" italics="true" pageId="5" pageNumber="67">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
”. These specimens were labelled “Helmr.”, realizing that the name of the collector was Helmreichen and not Helmenreich, as published in
|
||
<bibRefCitation id="EFEB4B63E1504537FF50FCE5FECDFC6F" author="Ohaus F." box="[219,382,888,915]" pageId="5" pageNumber="67" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus (1905)</bibRefCitation>
|
||
. Although
|
||
<bibRefCitation id="EFEB4B63E1504537FE77FCE5FD1BFC6F" author="Ohaus F." box="[508,680,889,915]" pageId="5" pageNumber="67" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
has described the variety as “
|
||
<taxonomicName id="4C7A4D11E1504537FB8BFCE5FEF8FC4A" authorityName="Ohaus" authorityYear="1914" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="variety" species="helmreicheni" variety="maculicollis">
|
||
<emphasis id="B90EEA80E1504537FB8BFCE5FAF6FC6F" box="[1024,1349,889,915]" italics="true" pageId="5" pageNumber="67">Hoplognathus helmreicheni</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E1504537FF37FC00FEF8FC4A" box="[188,331,924,950]" italics="true" pageId="5" pageNumber="67">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
” in the original publication, the correct name is
|
||
<taxonomicName id="4C7A4D11E1504537FCE7FC00FAC7FC4A" authorityName="Ohaus" authorityYear="1914" box="[876,1396,924,950]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E1504537FCE7FC00FB03FC4A" box="[876,1200,924,950]" italics="true" pageId="5" pageNumber="67">Hoplognathus helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E1504537FB6EFC00FAC7FC4A" box="[1253,1396,924,950]" italics="true" pageId="5" pageNumber="67">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
(as written on the original label). In this case,
|
||
<bibRefCitation id="EFEB4B63E1504537FD5FFC23FC32FC25" author="Ohaus F." box="[724,897,959,985]" pageId="5" pageNumber="67" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
made an unjustified emendation almost 10 years after the original publication of the species, not being recognized by the Code to validate a name (ICZN 1999, Art. 19.1, 33.2). Still, the latter spelling is used as valid, violating the Principle of Priority (ICZN 1999, Art. 23.9).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1504537FF36FBF6FC8DFB78" box="[189,830,1130,1157]" pageId="5" pageNumber="67" type="description">
|
||
<paragraph id="8BC53692E1504537FF36FBF6FC8DFB78" blockId="5.[189,830,1130,1157]" box="[189,830,1130,1157]" pageId="5" pageNumber="67">
|
||
<heading id="D08D81FEE1504537FF36FBF6FC8DFB78" bold="true" box="[189,830,1130,1157]" fontSize="11" level="3" pageId="5" pageNumber="67" reason="3">
|
||
<emphasis id="B90EEA80E1504537FF36FBF6FC8DFB78" bold="true" box="[189,830,1130,1157]" pageId="5" pageNumber="67">
|
||
Key to adult species of
|
||
<taxonomicName id="4C7A4D11E1504537FE56FBF6FC8DFB78" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[477,830,1130,1157]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA80E1504537FE56FBF6FDCAFB79" bold="true" box="[477,633,1130,1157]" italics="true" pageId="5" pageNumber="67">Oplognathus</emphasis>
|
||
<bibRefCitation id="EFEB4B63E1504537FD0BFBF6FC8DFB78" author="MacLeay WS" box="[640,830,1130,1156]" pageId="5" pageNumber="67" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E1504537FF36FB05FAC4F91D" pageId="5" pageNumber="67" type="key">
|
||
<key id="FFEB61A3E1504537FF36FB05FAC4F91D" pageId="5" pageNumber="67">
|
||
<keyStep id="308E2E37E1504537FF36FB05FAC4FA55" pageId="5" pageNumber="67">
|
||
<paragraph id="8BC53692E1504537FF36FB05FAC4FB05" blockId="5.[189,1399,1177,1761]" pageId="5" pageNumber="67">
|
||
<keyLead id="308B95A7E1504537FF36FB05FAC4FB05" pageId="5" pageNumber="67">1. Males with weakly trilobed clypeus, sides usually parallel; last maxillary palpomere cylindrical; mentum longer than wider, anterior margin sinuous; mesoventral process long (exceeding the anterior margin on mesocoxae, never exceeding the procoxae); (if female, clypeus rounded) ....... 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1504537FF4AFA9FFAC4FA55" blockId="5.[189,1399,1177,1761]" pageId="5" pageNumber="67">
|
||
<keyLead id="308B95A7E1504537FF4AFA9FFAC4FA55" pageId="5" pageNumber="67">
|
||
– Males with strongly trilobed clypeus, sides clearly divergent (
|
||
<figureCitation id="13412A17E1504537FC53FA9EFB80FAE1" box="[984,1075,1282,1309]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="5" pageNumber="67">Fig. 3b</figureCitation>
|
||
); last maxillary palpomere fusiform (
|
||
<figureCitation id="13412A17E1504537FED4FAB9FE05FABC" box="[351,438,1317,1344]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="5" pageNumber="67">
|
||
Fig.
|
||
<quantity id="4C829B77E1504537FE1EFABAFE05FABC" box="[405,438,1318,1344]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="67" unit="g" value="4.0">4g</quantity>
|
||
</figureCitation>
|
||
); mentum as long as wide, anterior margin notched (
|
||
<figureCitation id="13412A17E1504537FB97FAB9FBC0FABC" box="[1052,1139,1317,1344]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="5" pageNumber="67">Fig. 4h</figureCitation>
|
||
); mesoventral process short (not exceeding the anterior margin on mesocoxae), slightly flat (
|
||
<figureCitation id="13412A17E1504537FBB2FAD5FB01FA9F" box="[1081,1202,1353,1379]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="5" pageNumber="67">Fig. 8b, e</figureCitation>
|
||
); (if female, the clypeus rounded, and last maxillary palpomere cylindrical).
|
||
<collectingCountry id="F36D7602E1504537FC26FAF0FC4AFA7A" box="[941,1017,1388,1414]" name="Brazil" pageId="5" pageNumber="67">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="49BEF870E1504537FB8FFAF0FBF8FA7A" box="[1028,1099,1388,1414]" country="Brazil" name="Bahia" pageId="5" pageNumber="67">Bahia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49BEF870E1504537FBDCFAF0FB46FA7A" box="[1111,1269,1388,1414]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="67">Minas Gerais</collectingRegion>
|
||
............... ....................................................................................................................
|
||
<taxonomicName id="4C7A4D11E1504537FBCDFA13FAC4FA55" authority="Ohaus, 1912" authorityName="Ohaus" authorityYear="1912" box="[1094,1399,1422,1449]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E1504537FBCDFA13FB69FA55" bold="true" box="[1094,1242,1422,1449]" italics="true" pageId="5" pageNumber="67">O. bahianus</emphasis>
|
||
<bibRefCitation id="EFEB4B63E1504537FB69FA13FAC4FA55" author="Ohaus F." box="[1250,1399,1423,1449]" pageId="5" pageNumber="67" pagination="650" refId="ref11341" refString="Ohaus F. 1912. Hoplognathus bahianus n. sp. (Col. Rutel.). Deutsche Entomologische Zeitschrift 1912: 650." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="308E2E37E1504537FF36FA4DFAC4F91D" pageId="5" pageNumber="67">
|
||
<paragraph id="8BC53692E1504537FF36FA4DFAC4F9A9" blockId="5.[189,1399,1177,1761]" pageId="5" pageNumber="67">
|
||
<keyLead id="308B95A7E1504537FF36FA4DFAC4F9A9" pageId="5" pageNumber="67">
|
||
2. Body distinctly bicoloured. Pronotum dark green with yellow maculae. Elytra reddish brown to orange yellow; size varying from
|
||
<quantity id="4C829B77E1504537FD0EFA68FD42F9F2" box="[645,753,1524,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="5" pageNumber="67" unit="mm" value="17.2">17.2 mm</quantity>
|
||
to
|
||
<quantity id="4C829B77E1504537FC97FA68FC34F9F2" box="[796,903,1524,1550]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="5" pageNumber="67" unit="mm" value="18.6">18.6 mm</quantity>
|
||
in length (
|
||
<figureCitation id="13412A17E1504537FB84FA68FB30F9F2" box="[1039,1155,1524,1550]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="5" pageNumber="67">Fig. 1c–e</figureCitation>
|
||
); anterior margin of mentum slightly sinuous (
|
||
<figureCitation id="13412A17E1504537FDA6F98BFD20F9CD" box="[557,659,1559,1585]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="5" pageNumber="67">
|
||
Fig.
|
||
<quantity id="4C829B77E1504537FDE1F98BFD20F9CD" box="[618,659,1559,1585]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="67" unit="m" value="4.0">4m</quantity>
|
||
</figureCitation>
|
||
); females without dilated epipleuron, simple.
|
||
<collectingCountry id="F36D7602E1504537FB5BF98BFAAFF9CD" box="[1232,1308,1559,1585]" name="Brazil" pageId="5" pageNumber="67">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="49BEF870E1504537FAA6F98BFE8AF9A8" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="67">Minas Gerais</collectingRegion>
|
||
,
|
||
<collectingRegion id="49BEF870E1504537FECDF9A6FE25F9A8" box="[326,406,1594,1620]" country="Brazil" name="Parana" pageId="5" pageNumber="67">Paraná</collectingRegion>
|
||
......................................................................................
|
||
<taxonomicName id="4C7A4D11E1504537FB9CF9A6FAC4F9A9" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[1047,1399,1593,1621]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E1504537FB9CF9A6FB68F9A8" bold="true" box="[1047,1243,1593,1620]" italics="true" pageId="5" pageNumber="67">O. helmenreichi</emphasis>
|
||
<bibRefCitation id="EFEB4B63E1504537FB69F9A6FAC4F9A9" author="Ohaus F." box="[1250,1399,1594,1621]" pageId="5" pageNumber="67" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E1504537FF4AF9C2FAC4F91D" blockId="5.[189,1399,1177,1761]" pageId="5" pageNumber="67">
|
||
<keyLead id="308B95A7E1504537FF4AF9C2FAC4F91D" pageId="5" pageNumber="67">
|
||
– Body colour uniformly yellowish golden, sometimes with spots or maculae on pronotum. Elytra golden yellow, with greenish reflections; size varying from
|
||
<quantity id="4C829B77E1504537FC10F91CFBA2F967" box="[923,1041,1664,1691]" metricMagnitude="-2" metricUnit="m" metricValue="1.9149999999999998" pageId="5" pageNumber="67" unit="mm" value="19.15">19.15 mm</quantity>
|
||
to 22.0 mm in length (
|
||
<figureCitation id="13412A17E1504537FA9DF91CFF4FF943" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="5" pageNumber="67">Fig. 1a– b</figureCitation>
|
||
); females with distinctly dilated epipleuron.
|
||
<collectingCountry id="F36D7602E1504537FC87F938FCEBF942" box="[780,856,1700,1726]" name="Brazil" pageId="5" pageNumber="67">Brazil</collectingCountry>
|
||
:
|
||
<collectingRegion id="49BEF870E1504537FCEFF938FBBDF942" box="[868,1038,1700,1726]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="67">Espírito Santo</collectingRegion>
|
||
;
|
||
<collectingRegion id="49BEF870E1504537FB90F938FB0EF942" box="[1051,1213,1700,1726]" country="Brazil" name="Minas Gerais" pageId="5" pageNumber="67">Minas Gerais</collectingRegion>
|
||
;
|
||
<collectingRegion id="49BEF870E1504537FB42F938FAAEF942" box="[1225,1309,1700,1726]" country="Brazil" name="Parana" pageId="5" pageNumber="67">Paraná</collectingRegion>
|
||
;
|
||
<collectingRegion id="49BEF870E1504537FAA2F938FEF7F91D" country="Brazil" name="Rio de Janeiro" pageId="5" pageNumber="67">Rio de Janeiro</collectingRegion>
|
||
;
|
||
<collectingRegion id="49BEF870E1504537FEC4F95BFE76F91D" box="[335,453,1735,1761]" country="Brazil" name="Sao Paulo" pageId="5" pageNumber="67">São Paulo</collectingRegion>
|
||
........................................................................................
|
||
<taxonomicName id="4C7A4D11E1504537FBDFF95BFAC4F91D" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[1108,1399,1734,1761]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E1504537FBDFF95BFB0AF91D" bold="true" box="[1108,1209,1734,1761]" italics="true" pageId="5" pageNumber="67">O. kirbii</emphasis>
|
||
<bibRefCitation id="EFEB4B63E1504537FB4BF95BFAC4F91D" author="MacLeay WS" box="[1216,1399,1735,1761]" pageId="5" pageNumber="67" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |