148 lines
20 KiB
XML
148 lines
20 KiB
XML
<document id="DFA8F578699DE1575C500F6A9971CD73" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629613271721" checkinUser="felipe" docAuthor="Paquin, Pierre & Dupérré, Nadine" docDate="2009" docId="03C55737FFE5FFCBFF50F9B4D091FD9B" docLanguage="en" docName="zt02002p067.pdf" docOrigin="Zootaxa 2002" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cicurina (Cicurella) venii Gertsch 1992" docType="treatment" docVersion="3" lastPageNumber="53" masterDocId="FFFC2F4FFFD6FFFFFFC7FFA4D12FFFC0" masterDocTitle="of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution" masterLastPageNumber="67" masterPageNumber="1" pageNumber="52" updateTime="1699083252512" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E30E10C73FABC039CB8BA57343AA9B3C">of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution</mods:title>
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<mods:name id="9CC5BC6B0E6F6CF876A59CC0E96B2D09" type="personal">
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<mods:namePart id="B8695DF41280B8DBD8A0513CEC4849DC">Paquin, Pierre</mods:namePart>
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<mods:affiliation id="E6735EB93D7BA3EEDF8CDF6DA502CC19">Cave and Endangered Invertebrate Research Laboratory, SWCA Environmental Consultants, 4407 Monterey Oaks Boulevard, Building 1, Suite 110, Austin, Texas, 78749, USA. E-mail: ppaquin @ swca. com & This is publication no. 15 of the Karst Biosciences and Environmental Geophysics Research Laboratories, SWCA Environmental Consultants</mods:affiliation>
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<mods:namePart id="176AC63F8C2AFE50CC9387989B1766AB">Dupérré, Nadine</mods:namePart>
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<mods:affiliation id="E96C6484F9150C341C841808DDB02BA5">American Museum of Natural History, Division of Invertebrate Zoology, Central Park West at 79 Street, New York, New York, 10024 USA. E-mail: nduperre @ amnh. org</mods:affiliation>
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<mods:typeOfResource id="C453A277BC680288E5D5DD682D9D8ED7">text</mods:typeOfResource>
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<mods:title id="5A2F574861B4D99FCF422ABF56CDCC59">Zootaxa</mods:title>
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<mods:date id="D5AD450A75C6BA91D6790CCBDE8A0DC5">2009</mods:date>
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<mods:number id="0709FFC2A9BA89B01D46459EB5A4A8BD">2009-02-04</mods:number>
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<mods:number id="7DD63AB65581B764F659DC0D6EAD947A">2002</mods:number>
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<treatment id="03C55737FFE5FFCBFF50F9B4D091FD9B" ID-DOI="http://doi.org/10.5281/zenodo.5243089" ID-Zenodo-Dep="5243089" LSID="urn:lsid:plazi:treatment:03C55737FFE5FFCBFF50F9B4D091FD9B" httpUri="http://treatment.plazi.org/id/03C55737FFE5FFCBFF50F9B4D091FD9B" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
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<paragraph id="8BD3E621FFE5FFCCFF50F9B4D354F9EA" blockId="51.[151,635,1552,1613]" box="[151,635,1552,1578]" pageId="51" pageNumber="52">
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<taxonomicName id="4C6C9DA2FFE5FFCCFF50F9B4D354F9EA" ID-CoL="VB3Z" authority="Gertsch 1992" authorityName="Gertsch" authorityYear="1992" box="[151,635,1552,1578]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="venii" subGenus="Cicurella">
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<emphasis id="B9183A33FFE5FFCCFF50F9B4D354F9EA" bold="true" box="[151,635,1552,1578]" pageId="51" pageNumber="52">
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<emphasis id="B9183A33FFE5FFCCFF50F9B4D0E0F9EA" bold="true" box="[151,463,1552,1578]" italics="true" pageId="51" pageNumber="52">Cicurina (Cicurella) venii</emphasis>
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<bibRefCitation id="EFFD9BD0FFE5FFCCFE10F9B4D354F9EA" author="Gertsch, W. J." box="[471,635,1552,1578]" pageId="51" pageNumber="52" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376B5AAFFE5FFCCFF50F997D0ADF98D" box="[151,386,1587,1613]" pageId="51" pageNumber="52" type="description">
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<paragraph id="8BD3E621FFE5FFCCFF50F997D0ADF98D" blockId="51.[151,635,1552,1613]" box="[151,386,1587,1613]" pageId="51" pageNumber="52">
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(
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<figureCitation id="1357FAA4FFE5FFCCFF58F997D06EF98D" box="[159,321,1587,1613]" captionStart="FIGURES 108–127" captionStartId="48.[151,269,1827,1851]" captionTargetBox="[268,1307,197,1812]" captionTargetId="figure-117@48.[265,1322,194,1821]" captionTargetPageId="48" captionText="FIGURES 108–127. Cleared epigynum of troglobitic Cicurina (Cicurella) holotypes. For each species, a ventral and a dorsal view of the cleared epigynum are presented; the low number refers to the ventral view and the high number refers to the dorsal view. 108–109 Cicurina troglobia Cokendolpher 2004. 110–111 Cicurina ubicki Gertsch 1992. 112–113" pageId="51" pageNumber="52">Figs 116–117</figureCitation>
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,
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<figureCitation id="1357FAA4FFE5FFCCFE8AF997D055F98D" box="[333,378,1587,1613]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="51" pageNumber="52">134</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFE5FFCCFF50F925D44FF977" pageId="51" pageNumber="52" type="reference_group">
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<paragraph id="8BD3E621FFE5FFCCFF50F925D44FF977" blockId="51.[151,1437,1664,1719]" pageId="51" pageNumber="52">
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<treatmentCitationGroup id="AB7CC10FFFE5FFCCFF50F925D44FF977" pageId="51" pageNumber="52">
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<treatmentCitation id="0ACDC030FFE5FFCCFF50F925D0D2F957" author="Gertsch, W. J." box="[151,509,1664,1688]" page="111" pageId="51" pageNumber="52" year="1992">
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<taxonomicName id="4C6C9DA2FFE5FFCCFF50F925D0D2F957" ID-CoL="VB3Z" authority="Gertsch 1992: 111" authorityName="Gertsch" authorityPageNumber="111" authorityYear="1992" box="[151,509,1664,1688]" class="Arachnida" family="Hahniidae" genus="Cicurina" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="venii">
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<emphasis id="B9183A33FFE5FFCCFF50F925D01FF958" box="[151,304,1665,1688]" italics="true" pageId="51" pageNumber="52">Cicurina venii</emphasis>
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<bibRefCitation id="EFFD9BD0FFE5FFCCFEF0F924D0D2F957" author="Gertsch, W. J." box="[311,509,1664,1687]" pageId="51" pageNumber="52" pagination="75 - 122" refId="ref40017" refString="Gertsch, W. J. (1992). Distribution patterns and speciation in North American cave spiders with a list of the troglobites and revision of the cicurinas of the subgenus Cicurella. Texas Memorial Museum Speleological Monographs, 3. Studies on the endogean fauna of North America. 2, 75 - 122." type="journal article" year="1992">Gertsch 1992: 111</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
, figs 95–96 (description of female).
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<emphasis id="B9183A33FFE5FFCCFC46F924D2B3F958" bold="true" box="[897,924,1664,1688]" pageId="51" pageNumber="52">—</emphasis>
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||
<treatmentCitation id="0ACDC030FFE5FFCCFC64F924D55BF957" author="Jackman, J. A." box="[931,1140,1664,1687]" page="162" pageId="51" pageNumber="52" year="1997">
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<bibRefCitation id="EFFD9BD0FFE5FFCCFC64F924D55BF957" author="Jackman, J. A." box="[931,1140,1664,1687]" pageId="51" pageNumber="52" refId="ref40113" refString="Jackman, J. A. (1997) A Field Guide to Spiders and Scorpions of Texas. Houston, Texas: Gulf Publishing Company. xiv + 201 pp. + 32 plates." type="book" year="1997">Jackman 1997: 162</bibRefCitation>
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</treatmentCitation>
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, 171;
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<treatmentCitation id="0ACDC030FFE5FFCCFB72F924D024F977" author="Cokendolpher, J. C." page="19" pageId="51" pageNumber="52" year="2004">
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<bibRefCitation id="EFFD9BD0FFE5FFCCFB72F924D024F977" author="Cokendolpher, J. C." pageId="51" pageNumber="52" pagination="13 - 58" refId="ref39625" refString="Cokendolpher, J. C. (2004 a) Cicurina spiders from caves in Bexar County, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, 4, 13 - 58." type="journal article" year="2004">Cokendolpher 2004a: 19–20</bibRefCitation>
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</treatmentCitation>
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, 24–25, 27–28, 34, 52–54, figs 63–64;
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||
<treatmentCitation id="0ACDC030FFE5FFCCFD69F904D50EF977" author="Reddell, J. R. & Cokendolpher, J. C." box="[686,1057,1696,1719]" page="81" pageId="51" pageNumber="52" year="2004">
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||
<bibRefCitation id="EFFD9BD0FFE5FFCCFD69F904D50EF977" author="Reddell, J. R. & Cokendolpher, J. C." box="[686,1057,1696,1719]" pageId="51" pageNumber="52" pagination="75 - 94" refId="ref41088" refString="Reddell, J. R. & Cokendolpher, J. C. (2004) The cave spiders (Araneae) of Bexar and Comal counties, Texas. Texas Memorial Museum Speleological Monographs, 6. Studies on the cave and endogean fauna of North America, IV, 75 - 94." type="journal article" year="2004">Reddell & Cokendolpher 2004: 81</bibRefCitation>
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||
</treatmentCitation>
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||
;
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||
<treatmentCitation id="0ACDC030FFE5FFCCFBEAF904D473F977" author="Paquin, P. & Hedin, M." box="[1069,1372,1696,1719]" page="3249" pageId="51" pageNumber="52" year="2004">
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||
<bibRefCitation id="EFFD9BD0FFE5FFCCFBEAF904D473F977" author="Paquin, P. & Hedin, M." box="[1069,1372,1696,1719]" pageId="51" pageNumber="52" pagination="1 - 40" refId="ref40436" refString="Paquin, P., & Hedin, M. (2004) The power and perils of ' molecular taxonomy': a case study of eyeless and endangered relatives. Report to Fish and Wildlife Services (Austin, Texas, U. S. A). 1 - 40 p." type="book chapter" year="2004">Paquin & Hedin 2004: 3249</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376B5AAFFE5FFCCFF50F943D023F8E8" pageId="51" pageNumber="52" type="materials_examined">
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||
<paragraph id="8BD3E621FFE5FFCCFF50F943D023F8E8" blockId="51.[151,1436,1767,1948]" pageId="51" pageNumber="52">
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||
<materialsCitation id="3B04EC7CFFE5FFCCFF50F943D023F8E8" collectingDate="1980-11-22" collectionCode="AMNH" county="Bexar Co." location="Examined" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
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||
<emphasis id="B9183A33FFE5FFCCFF50F943D08AF8C1" bold="true" box="[151,421,1767,1793]" pageId="51" pageNumber="52">
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||
<typeStatus id="54D75883FFE5FFCCFF50F943D010F8C1" box="[151,319,1767,1793]" pageId="51" pageNumber="52" type="holotype">HOLOTYPE</typeStatus>
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female:
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</emphasis>
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“
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<collectingRegion id="49A828C3FFE5FFCCFE7EF943D32BF8C1" box="[441,516,1767,1793]" country="United States of America" name="Texas" pageId="51" pageNumber="52">Texas</collectingRegion>
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; Braken Bat Cave,
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<collectingCounty id="62B29EADFFE5FFCCFD29F943D245F8C1" box="[750,874,1767,1793]" pageId="51" pageNumber="52">Bexar Co.</collectingCounty>
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,
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||
<date id="FFD2C0E1FFE5FFCCFCBFF943D54CF8C1" box="[888,1123,1767,1793]" pageId="51" pageNumber="52" value="1980-11-22">
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<collectingDate id="EF963909FFE5FFCCFCBFF943D54CF8C1" box="[888,1123,1767,1793]" pageId="51" pageNumber="52" value="1980-11-22">22 November 1980</collectingDate>
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||
</date>
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. George Veni”.
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<location id="8EB3B0FAFFE5FFCCFAE4F943D4B4F8C1" LSID="urn:lsid:plazi:treatment:03C55737FFE5FFCBFF50F9B4D091FD9B:8EB3B0FAFFE5FFCCFAE4F943D4B4F8C1" box="[1315,1435,1767,1793]" county="Bexar Co." name="Examined" pageId="51" pageNumber="52" stateProvince="Texas">Examined</location>
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||
(
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<collectionCode id="ED7D7EE4FFE5FFCCFF58F8AAD1D0F8E8" box="[159,255,1806,1832]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="51" pageNumber="52">AMNH</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C376B5AAFFE5FFCBFF00F890D275FE5A" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
|
||
<paragraph id="8BD3E621FFE5FFCCFF00F890D305F88E" blockId="51.[151,1436,1767,1948]" box="[199,554,1844,1870]" pageId="51" pageNumber="52">
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||
<emphasis id="B9183A33FFE5FFCCFF00F890D070F88E" bold="true" box="[199,351,1844,1870]" pageId="51" pageNumber="52">Description.</emphasis>
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||
<emphasis id="B9183A33FFE5FFCCFEA1F891D305F88E" box="[358,554,1845,1870]" italics="true" pageId="51" pageNumber="52">
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||
Female
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||
<typeStatus id="54D75883FFE5FFCCFE02F891D305F88E" box="[453,554,1845,1870]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
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||
<paragraph id="8BD3E621FFE5FFCBFF00F8FFD275FE5A" blockId="51.[151,1436,1767,1948]" lastBlockId="52.[151,1436,152,603]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">Carapace length: 1.62; carapace width: 1.17. Eyes absent. Cheliceral promargin right side with 3 teeth (1 large, 2 small), left side likewise; retromargin right side with 6 denticles (3 large, 3 small), left side likewise. Epigynum: 0.28 (half size). Epigynal ventral plate with short transverse slit, bearing medial, shallow, inverted U-shaped indentation; atrium squared, two-thirds intruded by rounded dorsal plate, atrial aperture small; bursa not enlarged nor fused; copulatory ducts touching at midpoint, rounded, reaching over primary lobe apex; copulatory ducts constriction located basally; primary pores not visible; stalk short, straight, horizontally directed, joining the primary lobe at the aperture of the dictynoid pore; dictynoid pore elongated; primary lobe bean-shaped; secondary lobe absent; fertilization canal short, internally positioned from the dictynoid pore; fertilization duct medially attached to the primary lobe base.</paragraph>
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</subSubSection>
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||
<subSubSection id="C376B5AAFFE2FFCBFF00FE02D41FFE00" box="[199,1328,422,448]" pageId="52" pageNumber="53" type="distribution">
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||
<paragraph id="8BD3E621FFE2FFCBFF00FE02D41FFE00" blockId="52.[151,1436,152,603]" box="[199,1328,422,448]" pageId="52" pageNumber="53">
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||
<emphasis id="B9183A33FFE2FFCBFF00FE02D04FFE00" bold="true" box="[199,352,422,448]" pageId="52" pageNumber="53">Distribution</emphasis>
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||
. Only reported from the
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||
<typeStatus id="54D75883FFE2FFCBFD42FE02D398FE00" box="[645,695,422,448]" pageId="52" pageNumber="53">type</typeStatus>
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||
locality: Braken Bat Cave (Bexar County) (
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||
<figureCitation id="1357FAA4FFE2FFCBFB7BFE02D40CFE00" box="[1212,1315,422,448]" captionStart="FIGURE 134" captionStartId="57.[151,257,1593,1617]" captionTargetBox="[201,1369,204,1566]" captionTargetId="figure-238@57.[201,1388,194,1566]" captionTargetPageId="57" captionText="FIGURE 134. Distribution of troglobitic Cicurina (subgenus Cicurella) species in Bexar and Comal Counties. The range of the species is indicated by an area that includes all known locations. Each cave is indicated by a dot. The species ranges are accompanied by the illustration of the holotype epigynum (in ventral view). The type locality of C. platypus, Platypus Pit, apparently also harbors C. bullis, as indicated by the overlap of the ranges. The distribution range of C. madla includes a locality in Uvalde County (see Fig. 135), as suggested by the arrow pointing to the South West. Cicurina vespera is known from a single specimen apparently collected from Government Canyon Bat Cave. The DNA samples included in the analysis of Paquin & Hedin (2004) from this locality suggested that the immature Cicurina collected from Government Canyon Bat Cave are conspecific with C. madla, questioning the identity of the cave at the origin of the specimen used to describe C. vespera. The two localities harboring C. neovespera are rather obscure and could not be established with certainty, other than by an approximate location, resulting in a conflicting range of at least" pageId="52" pageNumber="53">Fig. 134</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C376B5AAFFE2FFCBFF00FE69D091FD9B" pageId="52" pageNumber="53" type="discussion">
|
||
<paragraph id="8BD3E621FFE2FFCBFF00FE69D091FD9B" blockId="52.[151,1436,152,603]" pageId="52" pageNumber="53">
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||
<emphasis id="B9183A33FFE2FFCBFF00FE69D06FFE27" bold="true" box="[199,320,461,487]" pageId="52" pageNumber="53">Remarks.</emphasis>
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Male unknown. Even though the epigynum is damaged, the remnants were enough to produce a complete description. The opisthosoma however, is missing. The entrance of Bracken Bat Cave has unfortunately been sealed with concrete. This species is included on the
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||
<collectingCountry id="F37BA6B1FFE2FFCBFBDBFDBED57FFDF4" box="[1052,1104,538,564]" name="United States of America" pageId="52" pageNumber="53">U.S.</collectingCountry>
|
||
Federal list of endangered species (
|
||
<bibRefCitation id="EFFD9BD0FFE2FFCBFF3CFDE5D09DFD9B" author="Longacre, C." box="[251,434,577,603]" pageId="52" pageNumber="53" pagination="81419 - 81433" refId="ref40235" refString="Longacre, C. (2000) Department of the Interior, Fish and Wildlife Service, 50 CFR part 17, RIN 1018 - AF 33. Endangered and threatened wildlife and plants; final rule to list nine Bexar County, Texas Invertebrate species as endangered. Federal Register, 65 (248), 81419 - 81433." type="journal article" year="2000">Longacre 2000</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |