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<mods:title id="8CB09E575CEEE12F8E2431CCAAE9BCD8">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVII. Revision of Neobuthus, with description of seven new species from Ethiopia, Kenya and Somaliland (Buthidae)</mods:title>
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<taxonomicName id="BDE24D45FFAE392DFC59BB95FB1AFC18" ID-CoL="46C28" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[968,1229,910,937]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="60" pageNumber="59" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">Neobuthus maidensis</taxonomicName>
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(
<figureCitation id="E2D92A43FFAE392DFCCABBABFC20FC76" box="[859,1015,944,967]" captionStart="Figures 255–267" captionStartId="52.[192,278,1738,1762]" captionTargetBox="[192,1435,197,1709]" captionTargetId="figure-16@52.[192,1439,842,1279]" captionTargetPageId="52" captionText="Figures 255–267: Figures 255–262: Neobuthus gubanensis sp. n. from type locality. Figures 255–257. Female paratype, right chelicera, dorsal (255) and ventral (256) views, and ventral view under UV fluorescence (257). Scale bar: 400 µm. Figures 258– 262. Male paratype (No. 1300), right hemispermatophore capsule region in posterior (258), convex compressed (259), convex (260) and anterior (261) views, and right hemispermatophore, convex view (262). Scale bars: 200 µm, 500 µm. Figures 263– 267: Neobuthus maidensis sp. n. male holotype (No. 1320), left hemispermatophore (mirror image): capsule region in posterior (263), convex compressed (264), convex (265) and anterior (266) views, hemispermatophore, convex view (267). Scale bars: 200 µm, 500 µm." pageId="60" pageNumber="59">Figs. 263–267</figureCitation>
, 
<figureCitation id="E2D92A43FFAE392DFB93BBABFBB5FC76" box="[1026,1122,944,967]" captionStart-0="Figures 311–314" captionStart-1="Figures 315–322" captionStart-2="Figures 323–330" captionStart-3="Figures 331–346" captionStart-4="Figures 347–348" captionStartId-0="61.[192,278,1712,1736]" captionStartId-1="62.[192,278,1471,1495]" captionStartId-2="63.[192,278,1656,1680]" captionStartId-3="64.[192,278,1314,1338]" captionStartId-4="65.[192,278,1688,1712]" captionTargetBox-0="[207,1425,243,1660]" captionTargetBox-1="[191,1437,191,1442]" captionTargetBox-2="[203,1439,178,1608]" captionTargetBox-3="[192,1430,200,1275]" captionTargetBox-4="[191,1440,191,1660]" captionTargetId-0="figure-16@61.[192,1439,239,1680]" captionTargetId-1="figure-293@62.[192,1437,192,1441]" captionTargetId-2="figure-47@63.[192,1439,168,1627]" captionTargetId-3="figure-11@64.[192,1437,192,1285]" captionTargetId-4="figure-36@65.[192,1439,192,1659]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionTargetPageId-2="63" captionTargetPageId-3="64" captionTargetPageId-4="65" captionText-0="Figures 311–314: Neobuthus maidensis sp. n.. Figures 311–312. Male holotype, dorsal (311) and ventral (312) views. Figures 313–314. Female paratype, dorsal (313) and ventral (314) views. Scale bar: 10 mm." captionText-1="Figures 315–322: Neobuthus maidensis sp. n.. Figures 315 and 317. Male holotype, carapace and tergites I–IV (315) and coxosternal area and sternites (317). Figures 316, 318–322. Female paratype, carapace and tergites I–IV (316), coxosternal area and sternites III–VI (318), and right legs I–IV, retrolateral aspect (319–322)." captionText-2="Figures 323–330: Neobuthus maidensis sp. n.. Figures 323, 328–330. Male holotype, metasoma V and telson lateral view (323), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. Figures. 324–327. Female paratype, metasoma V and telson lateral view (324), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. Scale bar: 10 mm (325– 330)." captionText-3="Figures 331–346: Neobuthus maidensis sp. n., pedipalp segments. Figures 331–339. Female paratype, pedipalp chela, dorsal (331), external (332), and ventral (333) views, pedipalp patella, dorsal (334), external (335) and ventral (336) views, pedipalp femur and trochanter ventral (337), internal (338) and dorsal (339) views. Figures 340–346. Male holotype, pedipalp chela, dorsal (340), external (341), and ventral (342) views, pedipalp patella, dorsal (343) and ventral (344) views, pedipalp femur and trochanter dorsal (345) and ventral (346) views. The trichobothrial pattern is indicated in Figures 332–335 and 338–339 (white circles)." captionText-4="Figures 347–348: Neobuthus maidensis sp. n. in vivo habitus. Male holotype (347) and female paratype (348)." pageId="60" pageNumber="59">311–348</figureCitation>
, 
<figureCitation id="E2D92A43FFAE392DFBFCBBABFB4FFC76" box="[1133,1176,944,967]" captionStart="Figures 402–415" captionStartId="75.[192,278,1796,1820]" captionTargetBox="[198,1435,192,1766]" captionTargetId="figure-195@75.[192,1439,192,1766]" captionTargetPageId="75" captionText="Figures 402–415: Pedipalp femur, patella and chela dorsal views of female and male. Figure 402. N. amoudensis sp. n. Figure 403. N. awashensis. Figure 404. N. berberensis (from sandy area). Figure 405. N. berberensis (from stony area). Figure 406. N. cloudsleythompsoni. Figure 407. N. erigavoensis sp. n. Figure 408. N. eritreaensis. Figure 409. N. factorio sp. n. Figure 410. N. ferrugineus. Figure 411. N. gubanensis sp. n. Figure 412. N. kloppersi sp. n. Figure 413. N. kutcheri. Figure 414. N. maidensis sp. n. Figure 415. N. montanus sp. n." pageId="60" pageNumber="59">414</figureCitation>
, 
<figureCitation id="E2D92A43FFAE392DFB33BBABFB1AFC76" box="[1186,1229,944,967]" captionStart="Figures 416–431" captionStartId="76.[192,278,1722,1746]" captionTargetBox="[192,1439,165,1704]" captionTargetId="figure-119@76.[192,1439,163,1704]" captionTargetPageId="76" captionText="Figures 416–431: Neobuthus, main characters used in the key to species. Figures 416–419. Pedipalp patella, dorsal view, female of N. berberensis (416), female of N. maidensis sp. n. (417), female of N. awashensis (418), and female of N. gubanensis sp. n. (419). Figures 420–421. Telotarsus III, ventral aspect, female of N. kutcheri (420), and female of N. awashensis (421). Figures 422–424. Metasoma IV–I, dorsal view, female of N. erigavoensis sp. n. (422), female of N. gubanensis sp. n. (423), and female of N. kloppersi sp. n. (424). Figures 425–427. Metasoma IV–I, ventral view, male of N. kloppersi sp. n. (425), male of N. erigavoensis sp. n. (426) and male of N. berberensis (427). Figures 428–429. Metasoma III–II, lateral view, female of N. gubanensis sp. n. (428), and female of N. kloppersi sp. n. (429). Figures 430–431. Sternites III–IV, male of N. montanus sp. n. (430), and male of N. gubanensis sp. n. (431)." pageId="60" pageNumber="59">417</figureCitation>
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<subSubSection id="32F8654DFFAE392DFCAFBD25FAB2FAE4" box="[830,1381,1342,1365]" pageId="60" pageNumber="59" type="etymology">
<paragraph id="7A5D36C6FFAE392DFCAFBD25FAB2FAE4" blockId="60.[830,1381,1342,1365]" box="[830,1381,1342,1365]" pageId="60" pageNumber="59">ETYMOLOGY. Named after the village of collection.</paragraph>
</subSubSection>
<subSubSection id="32F8654DFFAE392FFCAFBD67FD3EF920" lastPageId="62" lastPageNumber="61" pageId="60" pageNumber="59" type="diagnosis">
<paragraph id="7A5D36C6FFAE392FFCAFBD67FD3EF920" blockId="60.[830,1441,1404,1887]" lastBlockId="62.[192,802,1597,1681]" lastPageId="62" lastPageNumber="61" pageId="60" pageNumber="59">
DIAGNOSIS. Total length 
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(male), 
<quantity id="BD1A9B23FFAE392DFABEBD67FA77FA22" box="[1327,1440,1404,1427]" metricMagnitude="-2" metricUnit="m" metricValue="2.246" pageId="60" pageNumber="59" unit="mm" value="22.46">22.46 mm</quantity>
(female); carapace with area between anterior median carinae yellow to orange; tergites with 3 incomplete dark stripes; pedipalp relatively very slender, male with femur L/ 
<geoCoordinate id="1FD65001FFAE392DFC1BBDEDFC0BF9BC" box="[906,988,1526,1549]" degrees="2.56" direction="west" orientation="longitude" pageId="60" pageNumber="59" precision="555" value="-2.56">W 2.56</geoCoordinate>
, patella L/ 
<geoCoordinate id="1FD65001FFAE392DFBC1BDEDFB74F9BC" box="[1104,1187,1526,1549]" degrees="2.81" direction="west" orientation="longitude" pageId="60" pageNumber="59" precision="555" value="-2.81">W 2.81</geoCoordinate>
, chela L/ 
<geoCoordinate id="1FD65001FFAE392DFA99BDEDFA8CF9BC" box="[1288,1371,1526,1549]" degrees="5.33" direction="west" orientation="longitude" pageId="60" pageNumber="59" precision="555" value="-5.33">W 5.33</geoCoordinate>
; chela movable finger with 5 subrows of primary denticles, 4 external accessory denticles flanking proximal end of each subrow; trichobothria 
<emphasis id="4896EAD4FFAE392DFBF7BE49FBA4F9D8" box="[1126,1139,1618,1641]" italics="true" pageId="60" pageNumber="59">d</emphasis>
<subScript id="E6663483FFAE392DFBE5BE46FBAAF9DD" attach="left" box="[1140,1149,1629,1644]" fontSize="6" pageId="60" pageNumber="59">2</subScript>
usually absent from femur and patella; dorsoexternal and ventroexternal carinae on pedipalp patella in female present, smooth; posterior margins of tergites sparsely setose (male) or with 1–2 pairs of macrosetae (female); pedipalps, legs, metasoma and telson with moderately short, non- spiniform macrosetae in male, and long, fine setae in female; male with sternites III–VI shagreened to smooth medially, sternite VII finely granulated with 4 granulated carinae; female with sternites III–VI smooth, sternite VII with 4 weak granulated carinae; metasoma I–III with median lateral carinae present in both sexes; lateral surface of metasoma V in male densely granulated, with granules separated; soles of telotarsi with relatively sparse setation, leg III of adults with 12–14 ventral macrosetae on telotarsus; pectine teeth: 19–21 (male), 15–16 (female).
</paragraph>
</subSubSection>
<caption id="2E9D664EFFAF392CFF51BEABFBEFF951" pageId="61" pageNumber="60" startId="61.[192,278,1712,1736]" targetBox="[207,1425,243,1660]" targetPageId="61">
<paragraph id="7A5D36C6FFAF392CFF51BEABFBEFF951" blockId="61.[192,1422,1712,1760]" pageId="61" pageNumber="60">
<emphasis id="4896EAD4FFAF392CFF51BEABFE50F979" bold="true" box="[192,391,1712,1736]" pageId="61" pageNumber="60">Figures 311–314:</emphasis>
<taxonomicName id="BDE24D45FFAF392CFE03BEA9FDB5F976" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[402,610,1714,1735]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="61" pageNumber="60" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFAF392CFE03BEA9FDB5F976" box="[402,610,1714,1735]" italics="true" pageId="61" pageNumber="60">Neobuthus maidensis</emphasis>
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<emphasis id="4896EAD4FFAF392CFDFCBEA9FD53F976" bold="true" box="[621,644,1714,1735]" pageId="61" pageNumber="60">sp</emphasis>
. 
<emphasis id="4896EAD4FFAF392CFD05BEA9FD76F976" bold="true" box="[660,673,1714,1735]" pageId="61" pageNumber="60">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="4896EAD4FFAF392CFD29BEAAFCB4F977" bold="true" box="[696,867,1713,1735]" pageId="61" pageNumber="60">Figures 311–312</emphasis>
. Male holotype, dorsal (311) and ventral (312) views. 
<emphasis id="4896EAD4FFAF392CFF51BED1FEBFF96E" bold="true" box="[192,360,1738,1760]" pageId="61" pageNumber="60">Figures 313–314</emphasis>
. Female paratype, dorsal (313) and ventral (314) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="2E9D664EFFAC392FFF51BDA4FC76F9B9" pageId="62" pageNumber="61" startId="62.[192,278,1471,1495]" targetBox="[191,1437,191,1442]" targetPageId="62">
<paragraph id="7A5D36C6FFAC392FFF51BDA4FC76F9B9" blockId="62.[192,1441,1471,1544]" pageId="62" pageNumber="61">
<emphasis id="4896EAD4FFAC392FFF51BDA4FE51FA66" bold="true" box="[192,390,1471,1495]" pageId="62" pageNumber="61">Figures 315–322:</emphasis>
<taxonomicName id="BDE24D45FFAC392FFE1EBDD9FD88FA66" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[399,607,1474,1495]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="62" pageNumber="61" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFAC392FFE1EBDD9FD88FA66" box="[399,607,1474,1495]" italics="true" pageId="62" pageNumber="61">Neobuthus maidensis</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFAC392FFDF9BDDAFDA8FA67" bold="true" box="[616,639,1473,1494]" pageId="62" pageNumber="61">sp</emphasis>
. 
<emphasis id="4896EAD4FFAC392FFD1FBDDAFD4CFA67" bold="true" box="[654,667,1473,1494]" pageId="62" pageNumber="61">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="4896EAD4FFAC392FFD20BDDAFCFCFA67" bold="true" box="[689,811,1473,1495]" pageId="62" pageNumber="61">Figures 315</emphasis>
and 
<emphasis id="4896EAD4FFAC392FFCF1BDDAFC53FA67" bold="true" box="[864,900,1473,1494]" pageId="62" pageNumber="61">317</emphasis>
. Male holotype, carapace and tergites I–IV (315) and coxosternal area and sternites (317). 
<emphasis id="4896EAD4FFAC392FFDB9BDC2FD48FA5E" bold="true" box="[552,671,1497,1519]" pageId="62" pageNumber="61">Figures 316</emphasis>
, 
<emphasis id="4896EAD4FFAC392FFD3DBDC1FCD7FA5E" bold="true" box="[684,768,1498,1519]" pageId="62" pageNumber="61">318–322</emphasis>
. Female paratype, carapace and tergites I–IV (316), coxosternal area and sternites III–VI (318), and right legs I–IV, retrolateral aspect (319–322).
</paragraph>
</caption>
<subSubSection id="32F8654DFFAC3952FF51BEA3FC17F92D" lastPageId="67" lastPageNumber="66" pageId="62" pageNumber="61" type="description">
<paragraph id="7A5D36C6FFAC392EFF51BEA3FE17F887" blockId="62.[192,802,1720,1896]" lastBlockId="63.[192,802,1792,1907]" lastPageId="63" lastPageNumber="62" pageId="62" pageNumber="61">
DESCRIPTION. Total length of adult male 
<quantity id="BD1A9B23FFAC392FFD1ABEA3FD2DF97E" box="[651,762,1720,1743]" metricMagnitude="-2" metricUnit="m" metricValue="1.985" pageId="62" pageNumber="61" unit="mm" value="19.85">19.85 mm</quantity>
, of adult female 
<quantity id="BD1A9B23FFAC392FFEC1BECDFE14F95C" box="[336,451,1750,1773]" metricMagnitude="-2" metricUnit="m" metricValue="2.246" pageId="62" pageNumber="61" unit="mm" value="22.46">22.46 mm</quantity>
; measurements of carapace, telson, segments of metasoma and pedipalps given in 
<tableCitation id="3760037DFFAC392FFD74BEEEFF1AF89A" captionStart="Table 1" captionStartId="38.[192,257,795,819]" captionText="Table 1: Comparative measurements of adults of Neobuthus amoudensis sp. n. and N. factorio sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="62" pageNumber="61">Table 1</tableCitation>
; positions and distribution of trichobothria of pedipalps shown in 
<figureCitation id="E2D92A43FFAC392FFEB8BF29FD82F8F8" box="[297,597,1842,1865]" captionStart="Figures 331–346" captionStartId="64.[192,278,1314,1338]" captionTargetBox="[192,1430,200,1275]" captionTargetId="figure-11@64.[192,1437,192,1285]" captionTargetPageId="64" captionText="Figures 331–346: Neobuthus maidensis sp. n., pedipalp segments. Figures 331–339. Female paratype, pedipalp chela, dorsal (331), external (332), and ventral (333) views, pedipalp patella, dorsal (334), external (335) and ventral (336) views, pedipalp femur and trochanter ventral (337), internal (338) and dorsal (339) views. Figures 340–346. Male holotype, pedipalp chela, dorsal (340), external (341), and ventral (342) views, pedipalp patella, dorsal (343) and ventral (344) views, pedipalp femur and trochanter dorsal (345) and ventral (346) views. The trichobothrial pattern is indicated in Figures 332–335 and 338–339 (white circles)." pageId="62" pageNumber="61">Figs. 332–335 and 338–339</figureCitation>
; trichobothrium 
<emphasis id="4896EAD4FFAC392FFC9DBF29FCCEF8F8" box="[780,793,1842,1865]" italics="true" pageId="62" pageNumber="61">d</emphasis>
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usually absent from femur and patella; base color pale yellow to white with variable fuscous pigmentation and patterns of dark maculation on metasoma, and very weakly on pedipalps and legs; chelicerae yellow with dark reticulation on anterior manus, dentition reddish. 
<emphasis id="4896EAD4FFAC392FFCAFBEA3FBC2F97E" box="[830,1045,1720,1743]" italics="true" pageId="62" pageNumber="61">Sexual dimorphism</emphasis>
: strong, adult males substantially smaller, but without differences in shapes of pedipalps, metasoma and telson; pedipalp patella and femur granulate and matte in male, smooth and glossy in female; sternites smooth in females and partly granulated in male; macrosetae on pedipalps, legs, metasoma and telson much longer and finer in female than male; other sex differences cited below.
</paragraph>
<caption id="2E9D664EFFAD392EFF51BE63FF24F96B" pageId="63" pageNumber="62" startId="63.[192,278,1656,1680]" targetBox="[203,1439,178,1608]" targetPageId="63">
<paragraph id="7A5D36C6FFAD392EFF51BE63FF24F96B" blockId="63.[192,1440,1656,1754]" pageId="63" pageNumber="62">
<emphasis id="4896EAD4FFAD392EFF51BE63FE51F921" bold="true" box="[192,390,1656,1680]" pageId="63" pageNumber="62">Figures 323–330:</emphasis>
<taxonomicName id="BDE24D45FFAD392EFE01BE60FD88F921" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[400,607,1659,1680]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="63" pageNumber="62" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFAD392EFE01BE60FD88F921" box="[400,607,1659,1680]" italics="true" pageId="63" pageNumber="62">Neobuthus maidensis</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFAD392EFDF9BE61FDA8F93E" bold="true" box="[616,639,1658,1679]" pageId="63" pageNumber="62">sp</emphasis>
. 
<emphasis id="4896EAD4FFAD392EFD1FBE61FD4CF93E" bold="true" box="[654,667,1658,1679]" pageId="63" pageNumber="62">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="4896EAD4FFAD392EFD20BE61FCFBF93E" bold="true" box="[689,812,1658,1680]" pageId="63" pageNumber="62">Figures 323</emphasis>
, 
<emphasis id="4896EAD4FFAD392EFCAABE61FC58F93E" bold="true" box="[827,911,1658,1679]" pageId="63" pageNumber="62">328–330</emphasis>
. Male holotype, metasoma V and telson lateral view (323), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. 
<emphasis id="4896EAD4FFAD392EFC44BE88FBF6F918" bold="true" box="[981,1057,1683,1705]" pageId="63" pageNumber="62">Figures</emphasis>
. 
<emphasis id="4896EAD4FFAD392EFBBEBE88FB54F919" bold="true" box="[1071,1155,1683,1704]" pageId="63" pageNumber="62">324–327</emphasis>
. Female paratype, metasoma V and telson lateral view (324), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. Scale bar: 10 mm (325– 330).
</paragraph>
</caption>
<paragraph id="7A5D36C6FFAD3951FF51BF27FE48F981" blockId="63.[192,802,1792,1907]" lastBlockId="64.[192,802,1500,1891]" lastPageId="64" lastPageNumber="63" pageId="63" pageNumber="62">
<emphasis id="4896EAD4FFAD392EFF51BF27FEF2F8E5" bold="true" box="[192,293,1852,1876]" pageId="63" pageNumber="62">Pedipalp</emphasis>
(
<figureCitation id="E2D92A43FFAD392EFEA7BF26FE07F8E5" box="[310,464,1853,1876]" captionStart="Figures 331–346" captionStartId="64.[192,278,1314,1338]" captionTargetBox="[192,1430,200,1275]" captionTargetId="figure-11@64.[192,1437,192,1285]" captionTargetPageId="64" captionText="Figures 331–346: Neobuthus maidensis sp. n., pedipalp segments. Figures 331–339. Female paratype, pedipalp chela, dorsal (331), external (332), and ventral (333) views, pedipalp patella, dorsal (334), external (335) and ventral (336) views, pedipalp femur and trochanter ventral (337), internal (338) and dorsal (339) views. Figures 340–346. Male holotype, pedipalp chela, dorsal (340), external (341), and ventral (342) views, pedipalp patella, dorsal (343) and ventral (344) views, pedipalp femur and trochanter dorsal (345) and ventral (346) views. The trichobothrial pattern is indicated in Figures 332–335 and 338–339 (white circles)." pageId="63" pageNumber="62">Figs. 331–346</figureCitation>
). Pedipalp mostly sparsely hirsute; finely granulated in male and smooth in female; femur with five conspicuously granulose carinae, more strongly developed in male; patella with seven granulose carinae developed in male, and five smooth carinae in female; ventroexternal and dorsoexternal carinae on pedipalp patella in female present, smooth; chela with smooth carinae indicated in male, and more clearly developed in female.
</paragraph>
<caption id="2E9D664EFFD23951FF51BD39FEDAFA07" pageId="64" pageNumber="63" startId="64.[192,278,1314,1338]" targetBox="[192,1430,200,1275]" targetPageId="64">
<paragraph id="7A5D36C6FFD23951FF51BD39FEDAFA07" blockId="64.[192,1441,1314,1462]" pageId="64" pageNumber="63">
<emphasis id="4896EAD4FFD23951FF51BD39FE53FA8B" bold="true" box="[192,388,1314,1338]" pageId="64" pageNumber="63">Figures 331–346:</emphasis>
<taxonomicName id="BDE24D45FFD23951FE1DBD3EFD8DFA8B" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[396,602,1317,1338]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="64" pageNumber="63" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFD23951FE1DBD3EFD8DFA8B" box="[396,602,1317,1338]" italics="true" pageId="64" pageNumber="63">Neobuthus maidensis</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFD23951FDF0BD3FFDAFFA88" bold="true" box="[609,632,1316,1337]" pageId="64" pageNumber="63">sp</emphasis>
. 
<emphasis id="4896EAD4FFD23951FD14BD3FFD45FA88" bold="true" box="[645,658,1316,1337]" pageId="64" pageNumber="63">n</emphasis>
.
</taxonomicNameLabel>
, pedipalp segments. 
<emphasis id="4896EAD4FFD23951FCF9BD3FFBC7FA88" bold="true" box="[872,1040,1316,1338]" pageId="64" pageNumber="63">Figures 331–339</emphasis>
. Female paratype, pedipalp chela, dorsal (331), external (332), and ventral (333) views, pedipalp patella, dorsal (334), external (335) and ventral (336) views, pedipalp femur and trochanter ventral (337), internal (338) and dorsal (339) views. 
<emphasis id="4896EAD4FFD23951FC23BD4DFB8AFADA" bold="true" box="[946,1117,1366,1388]" pageId="64" pageNumber="63">Figures 340–346</emphasis>
. Male holotype, pedipalp chela, dorsal (340), external (341), and ventral (342) views, pedipalp patella, dorsal (343) and ventral (344) views, pedipalp femur and trochanter dorsal (345) and ventral (346) views. The trichobothrial pattern is indicated in Figures 332–335 and 338–339 (white c ircles).
</paragraph>
</caption>
<paragraph id="7A5D36C6FFD23951FF51BE2CFDB7F8D2" blockId="64.[192,802,1500,1891]" pageId="64" pageNumber="63">
<emphasis id="4896EAD4FFD23951FF51BE2CFEF9F9FE" bold="true" box="[192,302,1591,1615]" pageId="64" pageNumber="63">Carapace</emphasis>
(
<figureCitation id="E2D92A43FFD23951FED5BE23FE32F9FE" box="[324,485,1592,1615]" captionStart="Figures 315–322" captionStartId="62.[192,278,1471,1495]" captionTargetBox="[191,1437,191,1442]" captionTargetId="figure-293@62.[192,1437,192,1441]" captionTargetPageId="62" captionText="Figures 315–322: Neobuthus maidensis sp. n.. Figures 315 and 317. Male holotype, carapace and tergites I–IV (315) and coxosternal area and sternites (317). Figures 316, 318–322. Female paratype, carapace and tergites I–IV (316), coxosternal area and sternites III–VI (318), and right legs I–IV, retrolateral aspect (319–322)." pageId="64" pageNumber="63">Figs. 315–316</figureCitation>
). Strongly trapezoidal (narrower anteriorly), wider than long (L/ 
<geoCoordinate id="1FD65001FFD23951FD11BE4DFD01F9DC" box="[640,726,1622,1645]" degrees="0.82" direction="west" orientation="longitude" pageId="64" pageNumber="63" precision="555" value="-0.82">W 0.82</geoCoordinate>
–0.83); posterior median postocular area flat, anterior median preocular area gently sloped downwards towards anterior margin; lateral flanks steeply sloped; ocular tubercle broad, prominent, located slightly anterior to middle of carapace; anterior margin straight, finely microdenticulate, with coarser granules overlapping edge, bearing 6–8 macrosetae; all carinae indistinct; dense granulation covering most of carapace.
</paragraph>
<paragraph id="7A5D36C6FFD23951FCAFBDC0FAEBF978" blockId="64.[830,1441,1499,1891]" pageId="64" pageNumber="63">
<emphasis id="4896EAD4FFD23951FCAFBDC0FC7BFA42" bold="true" box="[830,940,1499,1523]" pageId="64" pageNumber="63">Chelicera</emphasis>
. Fingers with typical buthid dentition (Vachon, 196 3, Lowe &amp; Kovařík, 2016); fixed finger with large distal denticle, 1 subdistal denticle and 2 basal denticles fused into bicusp, single denticle on ventral surface at level of bicusp; dorsal margin of movable finger with 5 denticles: 1 large distal denticle, mediumsized subdistal and medial, and 2 small, partially fused basal denticles; ventral margin with 2 denticles.
</paragraph>
<paragraph id="7A5D36C6FFD23953FCAFBECBFEB5F94A" blockId="64.[830,1441,1499,1891]" lastBlockId="66.[192,802,1672,1848]" lastPageId="66" lastPageNumber="65" pageId="64" pageNumber="63">
<emphasis id="4896EAD4FFD23951FCAFBECBFC61F959" bold="true" box="[830,950,1744,1768]" pageId="64" pageNumber="63">Mesosoma</emphasis>
(
<figureCitation id="E2D92A43FFD23951FC5DBECAFBBBF959" box="[972,1132,1745,1768]" captionStart-0="Figures 311–314" captionStart-1="Figures 315–322" captionStartId-0="61.[192,278,1712,1736]" captionStartId-1="62.[192,278,1471,1495]" captionTargetBox-0="[207,1425,243,1660]" captionTargetBox-1="[191,1437,191,1442]" captionTargetId-0="figure-16@61.[192,1439,239,1680]" captionTargetId-1="figure-293@62.[192,1437,192,1441]" captionTargetPageId-0="61" captionTargetPageId-1="62" captionText-0="Figures 311–314: Neobuthus maidensis sp. n.. Figures 311–312. Male holotype, dorsal (311) and ventral (312) views. Figures 313–314. Female paratype, dorsal (313) and ventral (314) views. Scale bar: 10 mm." captionText-1="Figures 315–322: Neobuthus maidensis sp. n.. Figures 315 and 317. Male holotype, carapace and tergites I–IV (315) and coxosternal area and sternites (317). Figures 316, 318–322. Female paratype, carapace and tergites I–IV (316), coxosternal area and sternites III–VI (318), and right legs I–IV, retrolateral aspect (319–322)." pageId="64" pageNumber="63">Figs. 311–318</figureCitation>
). Tergites I–VI bear three carinae of which the lateral pair may be less conspicuous mainly on tergites I–IV; tergite VII bears five welldefined carinae (median, submedians and laterals); tergites I–VI granular, with coarser granules on posterior lateral areas; tergite VII densely granular; sternites III– VI smooth medially and granulate laterally in male and smooth in female; sternite VII with four well-defined carinae, sternite VII densely, finely granulated in male and weakly granulated in female; sternum 
<typeStatus id="A5598864FFD33950FD36BF44FD02F8C7" box="[679,725,1887,1910]" pageId="65" pageNumber="64">type</typeStatus>
1, triangular in shape, smooth, with deep posteromedian invagination; genital opercula smooth; genital papillae present; pectines extending to around a quarter of sternite V in male and around end of sternite IV in female; pectine teeth 19 and 
<quantity id="BD1A9B23FFD33950FBBEBF44FBB1F8C7" box="[1071,1126,1887,1910]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="65" pageNumber="64" unit="in" value="21.0">21 in</quantity>
male, 15 and 
<quantity id="BD1A9B23FFD33950FA98BF44FA97F8C7" box="[1289,1344,1887,1910]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="65" pageNumber="64" unit="in" value="16.0">16 in</quantity>
female; combs with 3 marginal lamellae and 8–9 middle lamellae; marginal lamellae, middle lamellae and fulcra with dense cover of short dark reddish macrosetae; fulcra with 2–5 setae.
</paragraph>
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<paragraph id="7A5D36C6FFD33950FF51BE83FAD7F901" blockId="65.[192,1280,1688,1712]" box="[192,1280,1688,1712]" pageId="65" pageNumber="64">
<emphasis id="4896EAD4FFD33950FF51BE83FE54F901" bold="true" box="[192,387,1688,1712]" pageId="65" pageNumber="64">Figures 347–348:</emphasis>
<taxonomicName id="BDE24D45FFD33950FE18BE80FD82F901" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[393,597,1691,1712]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="65" pageNumber="64" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFD33950FE18BE80FD82F901" box="[393,597,1691,1712]" italics="true" pageId="65" pageNumber="64">Neobuthus maidensis</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFD33950FDCDBE81FDA4F91E" bold="true" box="[604,627,1690,1711]" pageId="65" pageNumber="64">sp</emphasis>
. 
<emphasis id="4896EAD4FFD33950FDEFBE81FD5CF91E" bold="true" box="[638,651,1690,1711]" pageId="65" pageNumber="64">n</emphasis>
.
</taxonomicNameLabel>
in vivo habitus. Male holotype (347) and female paratype (348).
</paragraph>
</caption>
<caption id="2E9D664EFFD03953FF51BE2AFB95F9D3" pageId="66" pageNumber="65" startId="66.[192,278,1585,1609]" targetBox="[198,1413,201,1550]" targetPageId="66">
<paragraph id="7A5D36C6FFD03953FF51BE2AFB95F9D3" blockId="66.[192,1422,1585,1634]" pageId="66" pageNumber="65">
<emphasis id="4896EAD4FFD03953FF51BE2AFE50F9FB" bold="true" box="[192,391,1585,1610]" pageId="66" pageNumber="65">Figures 349–352:</emphasis>
<taxonomicName id="BDE24D45FFD03953FE03BE2FFDB7F9F8" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[402,608,1588,1609]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="66" pageNumber="65" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
<emphasis id="4896EAD4FFD03953FE03BE2FFDB7F9F8" box="[402,608,1588,1609]" italics="true" pageId="66" pageNumber="65">Neobuthus montanus</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFD03953FDFABE2FFD55F9F8" bold="true" box="[619,642,1588,1609]" pageId="66" pageNumber="65">sp</emphasis>
. 
<emphasis id="4896EAD4FFD03953FD03BE2FFD48F9F8" bold="true" box="[658,671,1588,1609]" pageId="66" pageNumber="65">n</emphasis>
</taxonomicNameLabel>
.. 
<emphasis id="4896EAD4FFD03953FD27BE28FCB5F9F9" bold="true" box="[694,866,1587,1609]" pageId="66" pageNumber="65">Figures 34 9–350</emphasis>
. Male holotype, dorsal (349) and ventral (350) views. 
<emphasis id="4896EAD4FFD03953FF51BE57FEBFF9D0" bold="true" box="[192,360,1612,1634]" pageId="66" pageNumber="65">Figures 351–352</emphasis>
. Female paratype, dorsal (351) and ventral (352) views. Scale bars: 10 mm.
</paragraph>
</caption>
<paragraph id="7A5D36C6FFD03953FF51BF1AFED2F889" blockId="66.[192,802,1672,1848]" pageId="66" pageNumber="65">
<emphasis id="4896EAD4FFD03953FF51BF1AFE7BF8A8" bold="true" box="[192,428,1793,1817]" pageId="66" pageNumber="65">Hemispermatophore</emphasis>
(
<figureCitation id="E2D92A43FFD03953FE50BF19FDB6F8A8" box="[449,609,1794,1817]" captionStart="Figures 255–267" captionStartId="52.[192,278,1738,1762]" captionTargetBox="[192,1435,197,1709]" captionTargetId="figure-16@52.[192,1439,842,1279]" captionTargetPageId="52" captionText="Figures 255–267: Figures 255–262: Neobuthus gubanensis sp. n. from type locality. Figures 255–257. Female paratype, right chelicera, dorsal (255) and ventral (256) views, and ventral view under UV fluorescence (257). Scale bar: 400 µm. Figures 258– 262. Male paratype (No. 1300), right hemispermatophore capsule region in posterior (258), convex compressed (259), convex (260) and anterior (261) views, and right hemispermatophore, convex view (262). Scale bars: 200 µm, 500 µm. Figures 263– 267: Neobuthus maidensis sp. n. male holotype (No. 1320), left hemispermatophore (mirror image): capsule region in posterior (263), convex compressed (264), convex (265) and anterior (266) views, hemispermatophore, convex view (267). Scale bars: 200 µm, 500 µm." pageId="66" pageNumber="65">Figs. 263–267</figureCitation>
). Typical of the genus.
</paragraph>
<paragraph id="7A5D36C6FFD03952FCAFBE9CFE29FA74" blockId="66.[830,1440,1671,1848]" lastBlockId="67.[192,802,1332,1906]" lastPageId="67" lastPageNumber="66" pageId="66" pageNumber="65">
<emphasis id="4896EAD4FFD03953FCAFBE9CFCA4F92E" bold="true" box="[830,883,1671,1695]" pageId="66" pageNumber="65">Legs</emphasis>
(
<figureCitation id="E2D92A43FFD03953FC13BE93FBF5F92E" box="[898,1058,1672,1695]" captionStart="Figures 315–322" captionStartId="62.[192,278,1471,1495]" captionTargetBox="[191,1437,191,1442]" captionTargetId="figure-293@62.[192,1437,192,1441]" captionTargetPageId="62" captionText="Figures 315–322: Neobuthus maidensis sp. n.. Figures 315 and 317. Male holotype, carapace and tergites I–IV (315) and coxosternal area and sternites (317). Figures 316, 318–322. Female paratype, carapace and tergites I–IV (316), coxosternal area and sternites III–VI (318), and right legs I–IV, retrolateral aspect (319–322)." pageId="66" pageNumber="65">Figs. 319–322</figureCitation>
). Coxa, femora, patella and tibia of all legs bearing variable numbers of short to medium length, straight, dark-reddish macrosetae; tarsi with mix of short and longer, dark-reddish macrosetae; basitarsi I– III slightly compressed with flat retrolateral surfaces, with bristle combs consisting of retrosuperior series of longer macrosetae, plus retroinferior and proinferior series of shorter macrosetae; telotarsi with two rows of short setae on ventral aspect, 12–14 macrosetae on telotarsus III; tibial spurs moderate on leg IV and reduced, very short on leg III.
</paragraph>
<caption id="2E9D664EFFD13952FF51BCD5FC52FB4E" pageId="67" pageNumber="66" startId="67.[192,278,1230,1254]" targetBox="[191,1437,191,1201]" targetPageId="67">
<paragraph id="7A5D36C6FFD13952FF51BCD5FC52FB4E" blockId="67.[192,1440,1230,1279]" pageId="67" pageNumber="66">
<emphasis id="4896EAD4FFD13952FF51BCD5FE50FB57" bold="true" box="[192,391,1230,1254]" pageId="67" pageNumber="66">Figures 353–354:</emphasis>
<taxonomicName id="BDE24D45FFD13952FE03BCCAFDB7FB57" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[402,608,1233,1254]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
<emphasis id="4896EAD4FFD13952FE03BCCAFDB7FB57" box="[402,608,1233,1254]" italics="true" pageId="67" pageNumber="66">Neobuthus montanus</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFD13952FDFABCCBFD55FB54" bold="true" box="[619,642,1232,1253]" pageId="67" pageNumber="66">sp</emphasis>
. 
<emphasis id="4896EAD4FFD13952FD03BCCBFD48FB54" bold="true" box="[658,671,1232,1253]" pageId="67" pageNumber="66">n</emphasis>
.
</taxonomicNameLabel>
, male paratype (No. 1541) from type locality, carapace and tergites (353), coxosternal area and sternites (354). UV fluorescence. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="7A5D36C6FFD13952FF51BDD7FA95FAD8" blockId="67.[192,802,1332,1906]" lastBlockId="67.[830,1440,1332,1385]" pageId="67" pageNumber="66">
<emphasis id="4896EAD4FFD13952FF51BDD7FE6EFA55" bold="true" box="[192,441,1484,1508]" pageId="67" pageNumber="66">Metasoma and telson</emphasis>
(
<figureCitation id="E2D92A43FFD13952FE5BBDD6FDB0FA55" box="[458,615,1485,1508]" captionStart="Figures 323–330" captionStartId="63.[192,278,1656,1680]" captionTargetBox="[203,1439,178,1608]" captionTargetId="figure-47@63.[192,1439,168,1627]" captionTargetPageId="63" captionText="Figures 323–330: Neobuthus maidensis sp. n.. Figures 323, 328–330. Male holotype, metasoma V and telson lateral view (323), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. Figures. 324–327. Female paratype, metasoma V and telson lateral view (324), metasoma and telson lateral (328), dorsal (329), and ventral (330) views. Scale bar: 10 mm (325– 330)." pageId="67" pageNumber="66">Figs. 323–330</figureCitation>
). Metasoma and telson sparsely hirsute, setae moderately short in male and longer in female, straight and reddish; metasomal segments I–III with 10 carinae, IV with 6–8 carinae, V with 2 carinae; segments I–III with moderate, granulate dorsolateral carinae, other carinae relatively well developed; segment IV with weakly indicated dorsolateral carinae; segment V with strong, granulate to dentatelobate ventrolateral carinae; segments I–IV with dense granulation on all intercarinal surfaces including dorsal surfaces; segment V densely granular on lateral and ventral surfaces, more coarsely so on ventral surface, granules not arranged along any traces of carinae; telson smooth, ventral surface sparsely, weakly granular; vesicle slightly elongated; aculeus stout, shorter than vesicle, tip of aculeus almost vertically directed.
</paragraph>
<paragraph id="7A5D36C6FFD13952FCAFBD8BFC17F92D" blockId="67.[830,1441,1424,1692]" pageId="67" pageNumber="66">
AFFINITIES. The described features distinguish 
<taxonomicName id="BDE24D45FFD13952FAD0BD8BFCA4FA74" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFD13952FAD0BD8BFCA4FA74" italics="true" pageId="67" pageNumber="66">N. maidensis</emphasis>
</taxonomicName>
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<emphasis id="4896EAD4FFD13952FC13BDB5FC4CFA77" bold="true" box="[898,923,1454,1478]" pageId="67" pageNumber="66">sp</emphasis>
. 
<emphasis id="4896EAD4FFD13952FC20BDB5FC17FA77" bold="true" box="[945,960,1454,1478]" pageId="67" pageNumber="66">n</emphasis>
.
</taxonomicNameLabel>
from all other species of the genus. 
<taxonomicName id="BDE24D45FFD13952FA19BDB5FC7EFA55" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFD13952FA19BDB5FC7EFA55" italics="true" pageId="67" pageNumber="66">N. maidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53A557AFFFD13952FC28BDD7FC28FA55" box="[953,1023,1484,1508]" pageId="67" pageNumber="66" rank="species">
<emphasis id="4896EAD4FFD13952FC28BDD7FC05FA55" bold="true" box="[953,978,1484,1508]" pageId="67" pageNumber="66">sp</emphasis>
. 
<emphasis id="4896EAD4FFD13952FC78BDD7FC2FFA55" bold="true" box="[1001,1016,1484,1508]" pageId="67" pageNumber="66">n</emphasis>
.
</taxonomicNameLabel>
has the narrowest pedipalp patella among all 
<taxonomicName id="BDE24D45FFD13952FC51BDF7FB59F9B2" box="[960,1166,1516,1539]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4896EAD4FFD13952FC51BDF7FBE5F9B2" box="[960,1074,1516,1539]" italics="true" pageId="67" pageNumber="66">Neobuthus</emphasis>
species
</taxonomicName>
(
<figureCitation id="E2D92A43FFD13952FB34BDF7FADEF9B2" box="[1189,1289,1516,1539]" captionStart="Figures 416–431" captionStartId="76.[192,278,1722,1746]" captionTargetBox="[192,1439,165,1704]" captionTargetId="figure-119@76.[192,1439,163,1704]" captionTargetPageId="76" captionText="Figures 416–431: Neobuthus, main characters used in the key to species. Figures 416–419. Pedipalp patella, dorsal view, female of N. berberensis (416), female of N. maidensis sp. n. (417), female of N. awashensis (418), and female of N. gubanensis sp. n. (419). Figures 420–421. Telotarsus III, ventral aspect, female of N. kutcheri (420), and female of N. awashensis (421). Figures 422–424. Metasoma IV–I, dorsal view, female of N. erigavoensis sp. n. (422), female of N. gubanensis sp. n. (423), and female of N. kloppersi sp. n. (424). Figures 425–427. Metasoma IV–I, ventral view, male of N. kloppersi sp. n. (425), male of N. erigavoensis sp. n. (426) and male of N. berberensis (427). Figures 428–429. Metasoma III–II, lateral view, female of N. gubanensis sp. n. (428), and female of N. kloppersi sp. n. (429). Figures 430–431. Sternites III–IV, male of N. montanus sp. n. (430), and male of N. gubanensis sp. n. (431)." pageId="67" pageNumber="66">Fig. 417</figureCitation>
) and several other unique characters, mainly almost white color (
<figureCitation id="E2D92A43FFD13952FAFABE11FC4BF9F1" captionStart="Figures 347–348" captionStartId="65.[192,278,1688,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-36@65.[192,1439,192,1659]" captionTargetPageId="65" captionText="Figures 347–348: Neobuthus maidensis sp. n. in vivo habitus. Male holotype (347) and female paratype (348)." pageId="67" pageNumber="66">Figs. 347–348</figureCitation>
), and indistinct anterior median carapacial carinae. The limited available sample of specimens precludes us from characterizing intraspecific variability of this species.
</paragraph>
</subSubSection>
<subSubSection id="32F8654DFFD13955FCAFBED9FE76F8FB" lastPageId="68" lastPageNumber="67" pageId="67" pageNumber="66" type="discussion">
<paragraph id="7A5D36C6FFD13955FCAFBED9FE76F8FB" blockId="67.[830,1441,1730,1906]" lastBlockId="68.[192,802,1645,1866]" lastPageId="68" lastPageNumber="67" pageId="67" pageNumber="66">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. The 
<typeStatus id="A5598864FFD13952FCAFBEFAFCBBF949" box="[830,876,1761,1784]" pageId="67" pageNumber="66">type</typeStatus>
locality, 
<emphasis id="4896EAD4FFD13952FC49BEFBFBC2F949" bold="true" box="[984,1045,1760,1784]" pageId="67" pageNumber="66">17SN</emphasis>
is sandy semi-desert to desert (fig. 
<quantity id="BD1A9B23FFD13952FCAFBF1BFC52F8A6" box="[830,901,1792,1815]" metricMagnitude="0" metricUnit="m" metricValue="3.048" pageId="67" pageNumber="66" unit="in" value="120.0">120 in</quantity>
<bibRefCitation id="1E734B37FFD13952FC1FBF1BFB4FF8A6" author="KOVARIK" box="[910,1176,1792,1815]" pageId="67" pageNumber="66" pagination="1 - 41" refId="ref28340" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík et al., 2018: 26</bibRefCitation>
and fig. 
<quantity id="BD1A9B23FFD13952FB6EBF1BFAEEF8A6" box="[1279,1337,1792,1815]" metricMagnitude="-1" metricUnit="m" metricValue="9.652" pageId="67" pageNumber="66" unit="in" value="38.0">38 in</quantity>
<bibRefCitation id="1E734B37FFD13952FAD3BF1BFC58F884" author="KOVARIK" pageId="67" pageNumber="66" pagination="1 - 41" refId="ref28340" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík, 2018: 8</bibRefCitation>
). The 
<typeStatus id="A5598864FFD13952FC46BF05FBD8F884" box="[983,1039,1822,1845]" pageId="67" pageNumber="66">types</typeStatus>
of 
<taxonomicName id="BDE24D45FFD13952FBA4BF05FB69F884" authority="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi, 2018" authorityName="Kovařík &amp; Lowe &amp; Awale &amp; Elmi &amp; Abdi" authorityYear="2018" box="[1077,1214,1822,1845]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="4896EAD4FFD13952FBA4BF05FB69F884" box="[1077,1214,1822,1845]" italics="true" pageId="67" pageNumber="66">N. maidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53A557AFFFD13952FB54BF05FAD5F884" box="[1221,1282,1822,1846]" pageId="67" pageNumber="66" rank="species">
<emphasis id="4896EAD4FFD13952FB54BF05FB09F887" bold="true" box="[1221,1246,1822,1846]" pageId="67" pageNumber="66">sp</emphasis>
. 
<emphasis id="4896EAD4FFD13952FB7DBF05FB2CF887" bold="true" box="[1260,1275,1822,1846]" pageId="67" pageNumber="66">n</emphasis>
.
</taxonomicNameLabel>
were obtained at night during UV collecting together with 
<taxonomicName id="BDE24D45FFD13952FAD1BF26FB7CF8C3" authority="Kovarik, 2018" authorityName="Kovarik" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="maidensis">
<emphasis id="4896EAD4FFD13952FAD1BF26FBD2F8C3" italics="true" pageId="67" pageNumber="66">Compsobuthus maidensis</emphasis>
<bibRefCitation id="1E734B37FFD13952FB83BF40FB7CF8C3" author="KOVARIK" box="[1042,1195,1883,1906]" pageId="67" pageNumber="66" pagination="1 - 41" refId="ref28340" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík, 2018</bibRefCitation>
</taxonomicName>
(
<typeStatus id="A5598864FFD13952FB51BF40FB27F8C3" box="[1216,1264,1883,1906]" pageId="67" pageNumber="66">type</typeStatus>
locality), 
<taxonomicName id="BDE24D45FFD13955FAFBBF40FDCFF935" authority="Kovarik et al., 2018" authorityName="Kovarik" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" lastPageId="68" lastPageNumber="67" order="Scorpiones" pageId="67" pageNumber="66" phylum="Arthropoda" rank="species" species="maidensis">
<emphasis id="4896EAD4FFD13955FAFBBF40FEFCF935" italics="true" lastPageId="68" lastPageNumber="67" pageId="67" pageNumber="66">Gint maidensis</emphasis>
<bibRefCitation id="1E734B37FFD63955FEA7BE76FDCFF935" author="KOVARIK" box="[310,536,1645,1668]" pageId="68" pageNumber="67" pagination="1 - 41" refId="ref28340" refString="KOVARIK, F., G. LOWE &amp; F. STAHLAVSKY. 2018. Review of the genus Babycurus Karsch, 1886 (Arachnida, Scorpiones, Buthidae), with description of Barbaracurus gen. n. and two new species from Oman and Yemen. Euscorpius, 267: 1 - 41." type="journal article" year="2018">Kovařík et al., 2018</bibRefCitation>
</taxonomicName>
(
<typeStatus id="A5598864FFD63955FDBABE76FD8CF935" box="[555,603,1645,1668]" pageId="68" pageNumber="67">type</typeStatus>
locality), 
<taxonomicName id="BDE24D45FFD63955FD41BE76FEC9F914" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="68" pageNumber="67" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4896EAD4FFD63955FD41BE76FF26F914" italics="true" pageId="68" pageNumber="67">Hottentotta</emphasis>
sp.
</taxonomicName>
, and 
<taxonomicName id="BDE24D45FFD63955FEF3BE95FD8AF914" box="[354,605,1678,1701]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="68" pageNumber="67" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="4896EAD4FFD63955FEF3BE95FE63F914" box="[354,436,1678,1701]" italics="true" pageId="68" pageNumber="67">Leiurus</emphasis>
sp. (Buthidae)
</taxonomicName>
. The first author arrived at the locality at night on 3th 
<date id="0E5C1006FFD63955FDC6BEB4FCD1F977" box="[599,774,1711,1734]" pageId="68" pageNumber="67" value="2017-09">September 2017</date>
at 21.00. At this time the temperature was 38.6 ºC and humidity 52%. Minimum temperature of 31.9 ºC and humidity of 46% were recorded on 
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4th September 
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</date>
in the early morning.
</paragraph>
</subSubSection>
</treatment>
</document>