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<document id="9F90B73C144BDAF30FB94F1588A68586" ID-CLB-Dataset="36842" ID-DOI="10.11646/zootaxa.3808.1.1" ID-GBIF-Dataset="154a67eb-e80e-451e-8afd-497de87d068a" ID-ISSN="1175-5326" ID-Zenodo-Dep="286171" ID-ZooBank="E4C0E93F-DBAB-4788-A8F7-C03E2C6F0306" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460429779140" checkinUser="plazi" docAuthor="Boumans, Louis &amp; Murányi, Dávid" docDate="2014" docId="B023F04EFFC1FF87D2ADFA5AFB323FC0" docLanguage="en" docName="zt03808p091.pdf" docOrigin="Zootaxa 3808 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Zwicknia westermanni Boumans &amp; Muranyi, sp. n." docType="treatment" docVersion="8" lastPageNumber="10" masterDocId="4C1A8836FFC4FF8DD23AFF88FFBB3F38" masterDocTitle="Two new species of Zwicknia Murányi, with molecular data on the phylogenetic position of the genus (Plecoptera, Capniidae)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="6" updateTime="1698331207130" updateUser="ExternalLinkService">
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<mods:title id="02E59E7F74A1C960FF73B463FD63E01A">Two new species of Zwicknia Murányi, with molecular data on the phylogenetic position of the genus (Plecoptera, Capniidae)</mods:title>
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<mods:namePart id="1FE0CD9DB6F5FA9A2E4BC31EEFD25C65">Boumans, Louis</mods:namePart>
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<mods:namePart id="E14BC7C041646F3A917648C18DFB9BE9">Murányi, Dávid</mods:namePart>
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<mods:title id="1CAAF788719E66CC60067F9A28B28BFD">Zootaxa</mods:title>
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<mods:date id="98C73A64394C068D260295084B5E7A48">2014</mods:date>
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<treatment id="B023F04EFFC1FF87D2ADFA5AFB323FC0" ID-DOI="http://doi.org/10.5281/zenodo.5618746" ID-GBIF-Taxon="127668164" ID-Zenodo-Dep="5618746" LSID="urn:lsid:plazi:treatment:B023F04EFFC1FF87D2ADFA5AFB323FC0" httpUri="http://treatment.plazi.org/id/B023F04EFFC1FF87D2ADFA5AFB323FC0" lastPageId="10" lastPageNumber="10" pageId="5" pageNumber="6">
<subSubSection id="709012D3FFC1FF88D2ADFA5AFE4E3936" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="38354158FFC1FF88D2ADFA5AFCB73AD5" blockId="5.[151,780,1490,1550]" box="[151,780,1490,1517]" pageId="5" pageNumber="6">
<heading id="637DF634FFC1FF88D2ADFA5AFCB73AD5" bold="true" box="[151,780,1490,1517]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="0AFE9D4AFFC1FF88D2ADFA5AFCB73AD5" bold="true" box="[151,780,1490,1517]" pageId="5" pageNumber="6">
<taxonomicName id="FF8A3ADBFFC1FF88D2ADFA5AFD063AD5" ID-CoL="7GGKK" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[151,701,1490,1517]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFC1FF88D2ADFA5AFE123AD4" bold="true" box="[151,425,1490,1516]" italics="true" pageId="5" pageNumber="6">Zwicknia westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFC1FF88D0F0FA5BFCB73AD5" box="[714,780,1491,1517]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="38354158FFC1FF88D2ADFA7DFE4E3936" blockId="5.[151,780,1490,1550]" box="[151,501,1525,1550]" pageId="5" pageNumber="6">
(Figs. 1–5;
<figureCitation id="A0B15DDDFFC1FF88D32DFA7DFEE03936" box="[279,347,1525,1550]" captionStart="FIGURE 6" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@8.[151,1436,193,1049]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Zwicknia westermanni Boumans &amp; Murányi, sp. n., mating pair, France, Burgundy, Saône-et-Loire department, Curgy. Photo A. Ruffoni." httpUri="https://zenodo.org/record/286172/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
;
<figureCitation id="A0B15DDDFFC1FF88D35DFA7DFE573936" box="[359,492,1525,1550]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="5" pageNumber="6">Figs. 14–19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFC1FF88D2ADF9BFFCD238E7" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="38354158FFC1FF88D2ADF9BFFE8439DE" blockId="5.[151,1437,1590,1766]" pageId="5" pageNumber="6">
<taxonomicName id="FF8A3ADBFFC1FF88D2ADF9BFFE653974" ID-CoL="QQPJ" authority="Newman, 1838" authorityName="Newman" authorityYear="1838" box="[151,478,1590,1612]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFC1FF88D2ADF9BFFE953974" box="[151,302,1590,1612]" italics="true" pageId="5" pageNumber="6">Capnia bifrons</emphasis>
(Newman, 1838)
</taxonomicName>
wing measurements and figure of brachypterous male habitus; abiotic habitat characteristics at
<typeStatus id="E731FFFAFFC1FF88D2FDF9DEFF483953" box="[199,243,1622,1643]" pageId="5" pageNumber="6">type</typeStatus>
locality—Westermann 1993 (the micropterous population in this paper refers to
<taxonomicName id="FF8A3ADBFFC1FF88D61FF9DEFB123953" ID-CoL="7GGKF" box="[1061,1193,1621,1643]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="0AFE9D4AFFC1FF88D61FF9DEFB123953" box="[1061,1193,1621,1643]" italics="true" pageId="5" pageNumber="6">Z. rupprechti</emphasis>
</taxonomicName>
)
<taxonomicName id="FF8A3ADBFFC1FF88D2ADF9FDFE5B39B2" ID-CoL="QQPJ" authority="Newman, 1838" authorityName="Newman" authorityYear="1838" box="[151,480,1652,1674]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFC1FF88D2ADF9FDFE9439B2" box="[151,303,1652,1674]" italics="true" pageId="5" pageNumber="6">Capnia bifrons</emphasis>
(Newman, 1838)
</taxonomicName>
‘Cappan’ race sensu
<bibRefCitation id="5C1B3CA9FFC1FF88D080F9FCFCE139B2" author="Rupprecht" box="[698,858,1652,1674]" pageId="5" pageNumber="6" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera: biology, ecology, systematics. MTL, Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
—
<bibRefCitation id="5C1B3CA9FFC1FF88D149F9FCFBA739B2" author="Rupprecht" box="[883,1052,1652,1674]" pageId="5" pageNumber="6" refString="Rupprecht, R. (1997) An attempt to explain different drumming signals within Capnia bifrons. In: Landolt, P. &amp; Sartori, M. (Eds.), Ephemeroptera &amp; Plecoptera: biology, ecology, systematics. MTL, Fribourg, pp. 93 - 98." type="book chapter" year="1997">Rupprecht 1997</bibRefCitation>
: 96 fig. 4 (drumming signals).
<taxonomicName id="FF8A3ADBFFC1FF88D2ADF91BFE653991" ID-CoL="QQPJ" authority="Newman, 1838" authorityName="Newman" authorityYear="1838" box="[151,478,1683,1705]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFC1FF88D2ADF91BFE953991" box="[151,302,1683,1705]" italics="true" pageId="5" pageNumber="6">Capnia bifrons</emphasis>
(Newman, 1838)
</taxonomicName>
brachypterous population—Haybach 2004 (material not studied, but brachypterous population is likely to belong to
<taxonomicName id="FF8A3ADBFFC1FF88D387F93AFDE439FF" ID-CoL="7GGKK" box="[445,607,1714,1735]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC1FF88D387F93AFDE439FF" box="[445,607,1714,1735]" italics="true" pageId="5" pageNumber="6">Z. westermanni</emphasis>
</taxonomicName>
; the micropterous population referred to is probably either
<taxonomicName id="FF8A3ADBFFC1FF88D6CBF93AFAE239FF" ID-CoL="QQPJ" box="[1265,1369,1713,1735]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFC1FF88D6CBF93AFAE239FF" box="[1265,1369,1713,1735]" italics="true" pageId="5" pageNumber="6">Z. bifrons</emphasis>
</taxonomicName>
or
<taxonomicName id="FF8A3ADBFFC1FF88D7BDF93AFE8A39DE" ID-CoL="7GGKF" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="0AFE9D4AFFC1FF88D7BDF93AFE8A39DE" italics="true" pageId="5" pageNumber="6">Z. rupprechti</emphasis>
</taxonomicName>
.)
</paragraph>
<paragraph id="38354158FFC1FF88D2ADF89AFCD238E7" blockId="5.[151,1436,1810,2015]" pageId="5" pageNumber="6">
<emphasis id="0AFE9D4AFFC1FF88D2ADF89AFE7F3813" bold="true" box="[151,452,1810,1835]" pageId="5" pageNumber="6">Morphological diagnosis.</emphasis>
Male brachypterous, ventral vesicle half as wide as the subgenital plate (Fig. 4). Main epiproct sclerite (Ep-scl): narrow and pointed in dorsal view (Fig. 2), tip straight, moderately acute in lateral view (Figs. 3, 14, 16, 18); ventral membranous section ends before the base in lateral view, apical spines stout, distributed not only on membranous parts but a few also on the sclerite (
<figureCitation id="A0B15DDDFFC1FF88D186F8F6FBD538AF" box="[956,1134,1918,1943]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="5" pageNumber="6">Figs. 14, 16, 18</figureCitation>
). Process of male tergite 9: high, perpendicularly raised, less than twice as wide as main epiproct sclerite (Fig. 2, 5), trapezoid and with sides not or just slightly indenting in caudal view (Figs. 5, 15, 17, 19).
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFC2FF8BD2FDFF1FFDE63C2C" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="38354158FFC2FF8BD2FDFF1FFC813FC0" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
<emphasis id="0AFE9D4AFFC2FF8BD2FDFF1FFECF3F88" bold="true" box="[199,372,151,176]" pageId="6" pageNumber="7">
<typeStatus id="E731FFFAFFC2FF8BD2FDFF1FFEB93F88" box="[199,258,151,176]" pageId="6" pageNumber="7">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="E731FFFAFFC2FF8BD3B3FF1FFE4A3F88" box="[393,497,151,176]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="409D01C8FFC2FF8BD071FF10FD613F88" box="[587,730,152,176]" name="Germany" pageId="6" pageNumber="7">GERMANY</collectingCountry>
: Lower Saxony, Hils Massif, Coppengrave, Duinger Wald, Hohenförsiek stream, N 5158.813' E 0941.732',
<quantity id="FF72ECBDFFC2FF8BD0EFFF34FC9F3FEC" box="[725,804,188,212]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="7" unit="m" value="210.0">210 m</quantity>
, 0 5.03.2011, leg. L. Boumans &amp; P.J. Nellestijn:
<specimenCount id="2E8C8AD1FFC2FF8BD756FF34FA273FEC" box="[1388,1436,188,212]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD2A5FF68FF403FC0" box="[159,251,224,248]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="6" pageNumber="7">NHMO</collectionCode>
: 10073107; used for molecular studies as No. 681).
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFE8DFBA73E25" blockId="6.[151,1437,151,1869]" box="[199,1052,260,285]" pageId="6" pageNumber="7">
Note: all
<typeStatus id="E731FFFAFFC2FF8BD315FE8DFEE43E25" box="[303,351,261,285]" pageId="6" pageNumber="7">type</typeStatus>
specimens used for molecular studies have the head detached.
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFEA0FCFA3D05" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
<typeStatus id="E731FFFAFFC2FF8BD2FDFEA0FE8F3E78" box="[199,308,296,320]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
<specimenCount id="2E8C8AD1FFC2FF8BD37AFEAFFE1A3E78" box="[320,417,295,320]" pageId="6" pageNumber="7" type="male">5 males</specimenCount>
:
<collectingCountry id="409D01C8FFC2FF8BD38BFEA0FDFB3E78" box="[433,576,296,320]" name="Germany" pageId="6" pageNumber="7">GERMANY</collectingCountry>
: Lower Saxony, Hils massif, Coppengrave, Duinger Wald, Hohenförsiek stream, N 5158.813’ E 0941.732’,
<quantity id="FF72ECBDFFC2FF8BD01AFEC4FDD13E5C" box="[544,618,332,356]" metricMagnitude="2" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="7" unit="m" value="210.0">210 m</quantity>
, 0 5.03.2011, leg. L. Boumans &amp; P.J. Nellestijn:
<specimenCount id="2E8C8AD1FFC2FF8BD6B1FEC4FB0D3E5C" box="[1163,1206,332,356]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD6FCFEC4FA993E5C" LSID="urn:lsid:biocol.org:col:34991" box="[1222,1314,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
: PLP4271; used for molecular studies as No.680, used for drawings Figs. 1–5,
<figureCitation id="A0B15DDDFFC2FF8BD1B7FEE7FBAD3EB0" box="[909,1046,367,392]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="6" pageNumber="7">Figs. 14–15</figureCitation>
);
<specimenCount id="2E8C8AD1FFC2FF8BD610FEF8FBEF3EBF" box="[1066,1108,368,392]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD65EFEF8FB043EB0" LSID="urn:lsid:biocol.org:col:34991" box="[1124,1215,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
: PLP4272; used for molecular studies as No.677, used for drawings
<figureCitation id="A0B15DDDFFC2FF8BD08FFE1CFC873E95" box="[693,828,404,429]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="6" pageNumber="7">Figs. 16–17</figureCitation>
); Idem, Hilsbach Stream, N 5158.769'
<geoCoordinate id="5DBE279FFFC2FF8BD6D7FE1DFADE3E95" box="[1261,1381,404,429]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="55" value="942.642">E 942.642</geoCoordinate>
',
<quantity id="FF72ECBDFFC2FF8BD748FE1CFF143EF7" metricMagnitude="2" metricUnit="m" metricValue="1.88" pageId="6" pageNumber="7" unit="m" value="188.0">188 m</quantity>
,
<date id="4C346798FFC2FF8BD287FE3FFE853EE8" box="[189,318,439,464]" pageId="6" pageNumber="7" value="2013-02-26">26.02.2013</date>
, leg. L. Boumans &amp; F. Westermann:
<specimenCount id="2E8C8AD1FFC2FF8BD0CAFE30FCA63EF7" box="[752,797,440,464]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD114FE30FC323EE8" box="[814,905,440,464]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="6" pageNumber="7">NHMO</collectionCode>
: 10073047; used for molecular studies as No. 1876);
<collectingCountry id="409D01C8FFC2FF8BD2D2FE54FEE03ECC" box="[232,347,476,500]" name="France" pageId="6" pageNumber="7">FRANCE</collectingCountry>
: Burgundy, Saône-et-Loire department, Curgy, Boisserand stream,
<geoCoordinate id="5DBE279FFFC2FF8BD674FE55FB123ECD" box="[1102,1193,476,501]" direction="north" orientation="latitude" pageId="6" pageNumber="7" precision="555" value="46.99">N 46.99</geoCoordinate>
<geoCoordinate id="5DBE279FFFC2FF8BD68BFE55FAB03ECC" box="[1201,1291,476,500]" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="9" value="0.073">E 0 4.38</geoCoordinate>
,
<date id="4C346798FFC2FF8BD72DFE54FA233ECC" box="[1303,1432,476,500]" pageId="6" pageNumber="7" value="2010-02-18">18.02.2010</date>
, leg. A. Ruffoni:
<specimenCount id="2E8C8AD1FFC2FF8BD368FD88FEC63D2F" box="[338,381,512,536]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD3B7FD88FE523D20" LSID="urn:lsid:biocol.org:col:34991" box="[397,489,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
: 4437; used for molecular studies as No.1868, used for drawings
<figureCitation id="A0B15DDDFFC2FF8BD6E8FE77FAE73D20" box="[1234,1372,511,536]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="6" pageNumber="7">Figs. 18–19</figureCitation>
);
<specimenCount id="2E8C8AD1FFC2FF8BD74BFD88FA273D2F" box="[1393,1436,512,536]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD2A5FDACFF403D04" box="[159,251,548,572]" httpUri="http://grbio.org/cool/2knt-7f1r" name="Natural History Museum, University of Oslo" pageId="6" pageNumber="7">NHMO</collectionCode>
: 10073059; used for molecular studies as No.1869).
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFDCFFDE63C2C" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
Other material - Records based on morphology:
<collectingCountry id="409D01C8FFC2FF8BD13FFDC0FCC33D58" box="[773,888,584,608]" name="France" pageId="6" pageNumber="7">FRANCE</collectingCountry>
: Provence-Alpes-Côte d’Azur, Var department, Pourrières,
<date id="4C346798FFC2FF8BD32DFDE4FE233DBC" box="[279,408,620,644]" pageId="6" pageNumber="7" value="1994-12-02">12.02.1994</date>
, leg. G. Vinçon:
<specimenCount id="2E8C8AD1FFC2FF8BD075FDE4FDC23DBC" box="[591,633,620,644]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD0BDFDE4FD5C3DBC" box="[647,743,620,644]" pageId="6" pageNumber="7">HMHM</collectionCode>
: PLP4284); Provence-Alpes-Côte d’Azur, Bouches-du-Rhône Department, Aix-en-Provence, Sainte Victoire, Infernet,
<date id="4C346798FFC2FF8BD113FD18FC113D90" box="[809,938,656,680]" pageId="6" pageNumber="7" value="1994-12-02">12.02.1994</date>
, leg. G. Vinçon:
<specimenCount id="2E8C8AD1FFC2FF8BD64CFD18FB1E3D9F" box="[1142,1189,656,680]" pageId="6" pageNumber="7" type="generic">2 ♂</specimenCount>
(
<collectionCode id="5E9BD99DFFC2FF8BD682FD18FAAF3D90" LSID="urn:lsid:biocol.org:col:34991" box="[1208,1300,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
: PLP4285); Provence-Alpes-Côte d’Azur, Var Department, La Garde-Freinet, Nible, N074 bridge, 0 2.01.2009, leg. M. Brulin:
<specimenCount id="2E8C8AD1FFC2FF8BD2ADFD50FF7C3DD7" box="[151,199,728,752]" pageId="6" pageNumber="7" type="generic">4 ♂</specimenCount>
, 1 ♀ nymph (CGV); Provence-Alpes-Côte d’Azur, Var Department, Plan-de-la-Tour, Largoustaou,
<quantity id="FF72ECBDFFC2FF8BD770FD5FFA2C3DD7" box="[1354,1431,727,752]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="6" pageNumber="7" unit="m" value="205.0">205 m</quantity>
,
<date id="4C346798FFC2FF8BD2ADFD74FEA33C2C" box="[151,280,764,788]" pageId="6" pageNumber="7" value="2009-01-31">31.01.2009</date>
, leg. J. Le Doar:
<specimenCount id="2E8C8AD1FFC2FF8BD3EDFD74FDBA3C2C" box="[471,513,764,788]" pageId="6" pageNumber="7" type="generic">1 ♂</specimenCount>
(CGV).
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFC2FF8BD2FDFC97FEFD3875" pageId="6" pageNumber="7" type="description">
<paragraph id="38354158FFC2FF8BD2FDFC97FB643C47" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
<emphasis id="0AFE9D4AFFC2FF8BD2FDFC97FEEA3C00" bold="true" box="[199,337,799,824]" pageId="6" pageNumber="7">Description</emphasis>
. Head, thorax, appendages and basal segments of the abdomen as is typical for the genus. Males brachypterous, females macropterous (
<figureCitation id="A0B15DDDFFC2FF8BD06CFCCCFD253C65" box="[598,670,836,861]" captionStart="FIGURE 6" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1049]" captionTargetId="figure@8.[151,1436,193,1049]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Zwicknia westermanni Boumans &amp; Murányi, sp. n., mating pair, France, Burgundy, Saône-et-Loire department, Curgy. Photo A. Ruffoni." httpUri="https://zenodo.org/record/286172/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
). Body length:
<typeStatus id="E731FFFAFFC2FF8BD162FCCCFC013C65" box="[856,954,836,861]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
<quantity id="FF72ECBDFFC2FF8BD1FCFCCCFB993C64" box="[966,1058,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="7" unit="mm" value="5.2">5.2 mm</quantity>
, male
<typeStatus id="E731FFFAFFC2FF8BD649FCCDFB653C65" box="[1139,1246,837,861]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
5.0–6.0, female 7.5–8.0 mm; forewing length:
<typeStatus id="E731FFFAFFC2FF8BD3D7FCEFFDF43CB8" box="[493,591,871,896]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
<quantity id="FF72ECBDFFC2FF8BD06CFCE0FD153C47" box="[598,686,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="7" unit="mm" value="2.3">2.3 mm</quantity>
, male
<typeStatus id="E731FFFAFFC2FF8BD0CDFCE0FCD93CB8" box="[759,866,872,896]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
<quantity id="FF72ECBDFFC2FF8BD150FCE0FC493C47" box="[874,1010,872,896]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="6" pageNumber="7" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1–2.9 mm</quantity>
, female 8.0–9.0 mm.
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFC04FAC03A35" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
Male terminalia (Figs. 1–5,
<figureCitation id="A0B15DDDFFC2FF8BD3C5FC04FD333C9C" box="[511,648,908,933]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="6" pageNumber="7">Figs. 14–19</figureCitation>
): Process of tergite 9 high, perpendicularly elevated, its apex is &lt;2× as wide as the medial section of the main epiproct sclerite (Ep-scl, sensu
<bibRefCitation id="5C1B3CA9FFC2FF8BD18BFC27FB2E3CF0" author="Muranyi" box="[945,1173,943,968]" pageId="6" pageNumber="7" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC2FF8BD623FC39FBE93CF0" box="[1049,1106,944,968]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2014
</bibRefCitation>
); its shape is trapezoid, the apex rounded or bearings two inconspicuous, hump-like tips; sides not or slightly indented in caudal view, the membranous portion narrowest apically. Tergite 10, basal and laterobasal epiproct sclerites as is typical for the genus. Main epiproct sclerite narrow and pointed in dorsal view, medially not swollen, its medial width is half of basal width; apex straight and moderately acute in lateral view, divided section long. Ventral membranous part between the left and right divisions of the main epiproct sclerite ends before the base in lateral view; apical spines stout, distributed on the membranous part with a few spines occurring on the sclerite. Inner sclerite (I-scl, sensu
<bibRefCitation id="5C1B3CA9FFC2FF8BD2ADFB24FEC63BFC" author="Muranyi" box="[151,381,1196,1221]" pageId="6" pageNumber="7" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC2FF8BD33AFB25FE813BFC" box="[256,314,1196,1220]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2014
</bibRefCitation>
) as is typical for the genus, eversible crest (Ec) long (
<figureCitation id="A0B15DDDFFC2FF8BD1D6FB24FBFA3BFC" box="[1004,1089,1196,1221]" captionStart="FIGURES 14 – 21" captionStartId="12.[151,264,1638,1661]" captionTargetBox="[312,1266,182,1638]" captionTargetId="figure@12.[293,1293,147,1648]" captionTargetPageId="12" captionText="FIGURES 14 – 21. Male terminalia of Zwicknia westermanni Boumans &amp; Murányi, sp. n. and Z. komica Murányi &amp; Boumans, sp. n. — 14, 16, 18, 20: main epiproct sclerite (Ep-scl), lateral view; 15, 17, 19, 21: hump of tergite 9, caudal view— 14 – 17: Z. westermanni paratypes, Germany, Lower Saxony, Hohenförsiek; 18 – 19: Z. westermanni paratypes France, Burgundy, Curgy; 20 – 21: Z. komica paratype, Russia, Komi Republic, Tyla-yu stream—scale 0.5 mm." httpUri="https://zenodo.org/record/286176/files/figure.png" pageId="6" pageNumber="7">Fig. 18</figureCitation>
). Sternite 9 slightly projecting medially, vesicle rather large, half as wide as the subgenital plate (Fig. 4). Subgenital plate rounded with triangular shape, tip slightly incised (Fig. 5). Paraprocts, fusion plate, retractoral plate and cerci as is typical for the genus.
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFA9FFB293A08" blockId="6.[151,1437,151,1869]" box="[199,1170,1303,1329]" pageId="6" pageNumber="7">
Female subgenital plate: As is typical for the genus (
<bibRefCitation id="5C1B3CA9FFC2FF8BD129FA9FFC433A08" author="Muranyi" box="[787,1016,1303,1328]" pageId="6" pageNumber="7" refString="Muranyi, D., Gamboa, M. &amp; Orci, K. M. (2014) Zwicknia gen. n. for the Capnia bifrons species group, with descriptions of three new species based on morphology, drumming signals and molecular genetics, and a synopsis of the West Palaearctic and Nearctic genera of Capniidae (Plecoptera). Zootaxa, in press." type="book" year="2014">
Murányi
<emphasis id="0AFE9D4AFFC2FF8BD146FA91FC0E3A08" box="[892,949,1304,1328]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2014
</bibRefCitation>
: Figs. 62-65).
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDFAB4FEF63931" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
<emphasis id="0AFE9D4AFFC2FF8BD2FDFAB4FE4E3A6D" bold="true" box="[199,501,1340,1365]" pageId="6" pageNumber="7">Morphological affinities.</emphasis>
Males of the species can be easily distinguished from congeners by being brachypterous and by the relatively large ventral vesicle. In other morphological features,
<taxonomicName id="FF8A3ADBFFC2FF8BD6B8FAE9FA953A4F" box="[1154,1326,1376,1400]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC2FF8BD6B8FAE9FA953A4F" box="[1154,1326,1376,1400]" italics="true" pageId="6" pageNumber="7">Z. westermanni</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="FF8A3ADBFFC2FF8BD28EFA0DFEAA3AA4" box="[180,273,1413,1436]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="0AFE9D4AFFC2FF8BD28EFA0DFEAA3AA4" box="[180,273,1413,1436]" italics="true" pageId="6" pageNumber="7">Z. acuta</emphasis>
</taxonomicName>
, but the following differences appear to be constant and distinctive: epiproct is more robust, less acute and the ventral membranous part ends closer to the base in the new species; process of tergite 9 lower, sides not or only slightly indented while the apex bears no conspicuous humps. Females are morphologically indistinguishable from congeners.
</paragraph>
<paragraph id="38354158FFC2FF8BD2FDF99CFEFD3875" blockId="6.[151,1437,151,1869]" pageId="6" pageNumber="7">
<emphasis id="0AFE9D4AFFC2FF8BD2FDF99CFEE93915" bold="true" box="[199,338,1556,1581]" pageId="6" pageNumber="7">Drumming.</emphasis>
<taxonomicName id="FF8A3ADBFFC2FF8BD35AF99CFDD83914" box="[352,611,1556,1580]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFC2FF8BD35AF99CFDD83914" box="[352,611,1556,1580]" italics="true" pageId="6" pageNumber="7">Zwicknia westermanni</emphasis>
</taxonomicName>
intersexual communication consists of three-way, monophasic signal exchanges (
<figureCitation id="A0B15DDDFFC2FF8BD327F9BFFEDF3968" box="[285,356,1591,1616]" captionStart="FIGURE 7" captionStartId="9.[151,250,712,734]" captionTargetBox="[151,1436,373,691]" captionTargetId="figure@9.[151,1436,373,691]" captionTargetPageId="9" captionText="FIGURE 7. Oscillogram showing an excerpt of a vibration duet of a pair of Z. westermanni Boumans &amp; Murányi, sp. n. from the Hohenförsiek stream at 20 ° C." httpUri="https://zenodo.org/record/286173/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
).
<tableCitation id="750874E3FFC2FF8BD346F9BFFE6A3968" box="[380,465,1591,1616]" captionStart="TABLE 2" captionStartId="8.[151,239,1163,1186]" captionTargetBox="[159,1288,1277,1795]" captionTargetPageId="8" captionText="TABLE 2. Descriptive statistics of duets of Z. westermanni Boumans &amp; Murányi, sp. n. based on three pairs from the Hohenförsiek stream recorded at different temperatures. 11 to 12 signals of each type were measured at 20 ° C, 11 to 13 at 22 ° C and 16 to 17 at 24 ° C. The average beat interval is calculated as (average signal duration) / (average beat nr. - 1)." httpUri="http://table.plazi.org/id/6CF511D0FFCCFF85D2ADFB03FAD13BDA" pageId="6" pageNumber="7" tableUuid="6CF511D0FFCCFF85D2ADFB03FAD13BDA">Table 2</tableCitation>
gives statistical values for the characteristics of duetting of three pairs (six different individuals) from the Hohenförsiek stream recorded at different temperatures. The average interval between beats of around 60 ms at 20°C is a coarse description. The second and/or third intervals are often the shortest; thereafter intervals gradually lengthen in a nonlinear way as shown in
<figureCitation id="A0B15DDDFFC2FF8BD170F92CFC343984" box="[842,911,1700,1725]" captionStart="FIGURE 8" captionStartId="9.[151,250,1802,1824]" captionTargetBox="[276,1313,815,1773]" captionTargetId="figure@9.[273,1313,815,1781]" captionTargetPageId="9" captionText="FIGURE 8. Beat interval length as a function of interval number in a duet of Z. westermanni Boumans &amp; Murányi, sp. n. from the Hohenförsiek stream at 20 ° C. Data points show the median value of seven measured signals." httpUri="https://zenodo.org/record/286174/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
. Note that the Call has the highest initial beat frequency, while the Reply has the largest number of beats. After each Call-Answer-Reply sequence, the male starts searching. A new signal exchange sequence typically follows after 3–6 seconds but large differences were found between pairs (
<tableCitation id="750874E3FFC2FF8BD3B7F887FE5D3810" box="[397,486,1807,1832]" captionStart="TABLE 2" captionStartId="8.[151,239,1163,1186]" captionTargetBox="[159,1288,1277,1795]" captionTargetPageId="8" captionText="TABLE 2. Descriptive statistics of duets of Z. westermanni Boumans &amp; Murányi, sp. n. based on three pairs from the Hohenförsiek stream recorded at different temperatures. 11 to 12 signals of each type were measured at 20 ° C, 11 to 13 at 22 ° C and 16 to 17 at 24 ° C. The average beat interval is calculated as (average signal duration) / (average beat nr. - 1)." httpUri="http://table.plazi.org/id/6CF511D0FFCCFF85D2ADFB03FAD13BDA" pageId="6" pageNumber="7" tableUuid="6CF511D0FFCCFF85D2ADFB03FAD13BDA">Table 2</tableCitation>
). This value was not calculated for the duet recorded at 22°C because not all Calls were answered.
</paragraph>
</subSubSection>
<paragraph id="38354158FFC3FF8AD2ADF801FF7D38E5" pageId="7" pageNumber="8">
<table id="4A8AB3F8FFC30072D2ADF801FA2738E5" box="[151,1436,1929,2013]" gridcols="3" gridrows="3" pageId="7" pageNumber="8">
<tr id="86BA431AFFC30072D2ADF801FA273898" box="[151,1436,1929,1952]" gridrow="0" pageId="7" pageNumber="8">
<th id="C56B2A66FFC30072D2ADF801FDD03898" box="[151,619,1929,1952]" gridcol="0" gridrow="0" pageId="7" pageNumber="8">
<emphasis id="0AFE9D4AFFC3FF8AD2ADF801FE8738A7" bold="true" box="[151,316,1929,1952]" pageId="7" pageNumber="8">FIGURES 1–5.</emphasis>
Male terminalia of
<taxonomicName id="FF8A3ADBFFC3FF8AD037F802FDD13898" box="[525,618,1930,1952]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="0AFE9D4AFFC3FF8AD037F802FDD13898" box="[525,618,1930,1952]" italics="true" pageId="7" pageNumber="8">Zwicknia</emphasis>
</taxonomicName>
</th>
<th id="C56B2A66FFC30072D048F801FB353898" box="[626,1166,1929,1952]" gridcol="1" gridrow="0" pageId="7" pageNumber="8">
<taxonomicName id="FF8A3ADBFFC3FF8AD048F803FC613898" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[626,986,1930,1952]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFC3FF8AD048F803FD483898" box="[626,755,1931,1952]" italics="true" pageId="7" pageNumber="8">westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFC3FF8AD1DFF802FB9B3898" box="[997,1056,1930,1952]" pageId="7" pageNumber="8" rank="species">
<emphasis id="0AFE9D4AFFC3FF8AD1DFF802FBA138A7" bold="true" box="[997,1050,1930,1951]" pageId="7" pageNumber="8">sp. n</emphasis>
.
</taxonomicNameLabel>
; paratype,
</th>
<th id="C56B2A66FFC30072D6AFF801FA273898" box="[1173,1436,1929,1952]" gridcol="2" gridrow="0" pageId="7" pageNumber="8">Germany, Lower Saxony,</th>
</tr>
<tr id="86BA431AFFC30072D2ADF821FA273887" box="[151,1436,1961,1983]" gridrow="1" pageId="7" pageNumber="8">
<th id="C56B2A66FFC30072D2ADF821FDD03887" box="[151,619,1961,1983]" gridcol="0" gridrow="1" pageId="7" pageNumber="8">Coppengrave, Hohenförsiek—1: dorso-caudal</th>
<td id="C56B2A66FFC30072D048F821FB353887" box="[626,1166,1961,1983]" gridcol="1" gridrow="1" pageId="7" pageNumber="8">view; 2: dorsal view; 3: lateral view; 4: ventral view;</td>
<td id="C56B2A66FFC30072D6AFF821FA273887" box="[1173,1436,1961,1983]" gridcol="2" gridrow="1" pageId="7" pageNumber="8">5: caudal view—scale 0.5</td>
</tr>
<tr id="86BA431AFFC30072D2ADF840FA2738E5" box="[151,1436,1992,2013]" gridrow="2" pageId="7" pageNumber="8" rowspan-1="1" rowspan-2="1">
<th id="C56B2A66FFC30072D2ADF840FDD038E5" box="[151,619,1992,2013]" gridcol="0" gridrow="2" pageId="7" pageNumber="8">mm.</th>
</tr>
</table>
</paragraph>
<caption id="6CF511D0FFCCFF85D2ADFBA7FDAE3B5B" httpUri="https://zenodo.org/record/286172/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1436,193,1049]" targetPageId="8">
<paragraph id="38354158FFCCFF85D2ADFBA7FDAE3B5B" blockId="8.[151,1436,1071,1123]" pageId="8" pageNumber="9">
<emphasis id="0AFE9D4AFFCCFF85D2ADFBA7FEAE3B7C" bold="true" box="[151,277,1071,1093]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="FF8A3ADBFFCCFF85D3A9FBA7FCD93B7D" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[403,866,1071,1093]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFCCFF85D3A9FBA7FDC13B7D" box="[403,634,1071,1093]" italics="true" pageId="8" pageNumber="9">Zwicknia westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFCCFF85D154FBA7FC113B7C" box="[878,938,1071,1092]" pageId="8" pageNumber="9" rank="species">
<emphasis id="0AFE9D4AFFCCFF85D154FBA7FC113B7C" bold="true" box="[878,938,1071,1092]" pageId="8" pageNumber="9">sp. n.</emphasis>
</taxonomicNameLabel>
, mating pair, France, Burgundy, Saône-et-Loire department, Curgy. Photo A. Ruffoni.
</paragraph>
</caption>
<caption id="6CF511D0FFCCFF85D2ADFB03FAD13BDA" ID-Table-UUID="6CF511D0FFCCFF85D2ADFB03FAD13BDA" httpUri="http://table.plazi.org/id/6CF511D0FFCCFF85D2ADFB03FAD13BDA" pageId="8" pageNumber="9" targetBox="[159,1288,1277,1795]" targetIsTable="true" targetPageId="8">
<paragraph id="38354158FFCCFF85D2ADFB03FAD13BDA" blockId="8.[151,1436,1163,1250]" pageId="8" pageNumber="9">
<emphasis id="0AFE9D4AFFCCFF85D2ADFB03FEB03B9A" bold="true" box="[151,267,1163,1186]" pageId="8" pageNumber="9">TABLE 2.</emphasis>
Descriptive statistics of duets of
<taxonomicName id="FF8A3ADBFFCCFF85D04DFB03FBB63B9A" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[631,1037,1163,1186]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFCCFF85D04DFB03FCA63B9A" box="[631,797,1163,1186]" italics="true" pageId="8" pageNumber="9">Z. westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFCCFF85D623FB03FBED3B9A" box="[1049,1110,1163,1186]" pageId="8" pageNumber="9" rank="species">
<emphasis id="0AFE9D4AFFCCFF85D623FB03FBEB3B9A" bold="true" box="[1049,1104,1163,1186]" pageId="8" pageNumber="9">sp. n</emphasis>
.
</taxonomicNameLabel>
based on three pairs from the Hohenförsiek stream recorded at different temperatures. 11 to 12 signals of each type were measured at 20°C, 11 to 13 at 22°C and 16 to 17 at 24°C. The average beat interval is calculated as (average signal duration)/(average beat nr. -1).
</paragraph>
</caption>
<paragraph id="38354158FFCCFF85D0BFFB75FB2E383B" pageId="8" pageNumber="9">
<table id="4A8AB3F8FFCC0072D2A5FB75FAB3383B" box="[159,1288,1277,1795]" gridcols="4" gridrows="13" pageId="8" pageNumber="9">
<tr id="86BA431AFFCC0072D2A5FB75FAB33A2B" box="[159,1288,1277,1299]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
<th id="C56B2A66FFCC0072D0BFFB75FCBE3A2B" box="[645,773,1277,1299]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">20°C</th>
<th id="C56B2A66FFCC0072D148FB75FC773A2B" box="[882,972,1277,1299]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">22°C</th>
<th id="C56B2A66FFCC0072D641FB75FAB33A2B" box="[1147,1288,1277,1299]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">24°C</th>
</tr>
<tr id="86BA431AFFCC0072D2A5FAAFFAB33A05" box="[159,1288,1319,1341]" gridrow="1" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FAAFFD833A05" box="[159,568,1319,1341]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">interval between 3-way sequences (ms)</th>
<td id="C56B2A66FFCC0072D0BFFAAFFCBE3A05" box="[645,773,1319,1341]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">4.450±1.181</td>
<td id="C56B2A66FFCC0072D148FAAFFC773A05" box="[882,972,1319,1341]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">n/a</td>
<td id="C56B2A66FFCC0072D641FAAFFAB33A05" box="[1147,1288,1319,1341]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">23.523±9.929</td>
</tr>
<tr id="86BA431AFFCC0072D2A5FAD8FAB33A5E" box="[159,1288,1360,1382]" gridrow="2" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FAD8FD833A5E" box="[159,568,1360,1382]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">♂ CALL nr. of beats</th>
<td id="C56B2A66FFCC0072D0BFFAD8FCBE3A5E" box="[645,773,1360,1382]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">8.6±2.8</td>
<td id="C56B2A66FFCC0072D148FAD8FC773A5E" box="[882,972,1360,1382]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">9.0±0.5</td>
<td id="C56B2A66FFCC0072D641FAD8FAB33A5E" box="[1147,1288,1360,1382]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">8.3±0.7</td>
</tr>
<tr id="86BA431AFFCC0072D2A5FAF1FAB33AB7" box="[159,1288,1401,1423]" gridrow="3" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FAF1FD833AB7" box="[159,568,1401,1423]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">duration (ms)</th>
<td id="C56B2A66FFCC0072D0BFFAF1FCBE3AB7" box="[645,773,1401,1423]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">441±50</td>
<td id="C56B2A66FFCC0072D148FAF1FC773AB7" box="[882,972,1401,1423]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">392±39</td>
<td id="C56B2A66FFCC0072D641FAF1FAB33AB7" box="[1147,1288,1401,1423]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">289±36</td>
</tr>
<tr id="86BA431AFFCC0072D2A5FA2BFAB33A81" box="[159,1288,1443,1465]" gridrow="4" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FA2BFD833A81" box="[159,568,1443,1465]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">avg beat interval</th>
<td id="C56B2A66FFCC0072D0BFFA2BFCBE3A81" box="[645,773,1443,1465]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">58</td>
<td id="C56B2A66FFCC0072D148FA2BFC773A81" box="[882,972,1443,1465]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">49</td>
<td id="C56B2A66FFCC0072D641FA2BFAB33A81" box="[1147,1288,1443,1465]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">35</td>
</tr>
<tr id="86BA431AFFCC0072D2A5FA44FAB33ADA" box="[159,1288,1484,1506]" gridrow="5" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FA44FD833ADA" box="[159,568,1484,1506]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">♀ response time</th>
<td id="C56B2A66FFCC0072D0BFFA44FCBE3ADA" box="[645,773,1484,1506]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">734±215</td>
<td id="C56B2A66FFCC0072D148FA44FC773ADA" box="[882,972,1484,1506]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">504±100</td>
<td id="C56B2A66FFCC0072D641FA44FAB33ADA" box="[1147,1288,1484,1506]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">376±94</td>
</tr>
<tr id="86BA431AFFCC0072D2A5FA7DFAB33933" box="[159,1288,1525,1547]" gridrow="6" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5FA7DFD833933" box="[159,568,1525,1547]" gridcol="0" gridrow="6" pageId="8" pageNumber="9">♀ ANSWER nr. of beats</th>
<td id="C56B2A66FFCC0072D0BFFA7DFCBE3933" box="[645,773,1525,1547]" gridcol="1" gridrow="6" pageId="8" pageNumber="9">8.9±1.8</td>
<td id="C56B2A66FFCC0072D148FA7DFC773933" box="[882,972,1525,1547]" gridcol="2" gridrow="6" pageId="8" pageNumber="9">8.1±0.3</td>
<td id="C56B2A66FFCC0072D641FA7DFAB33933" box="[1147,1288,1525,1547]" gridcol="3" gridrow="6" pageId="8" pageNumber="9">7.7±0.9</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F997FAB3390D" box="[159,1288,1567,1589]" gridrow="7" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F997FD83390D" box="[159,568,1567,1589]" gridcol="0" gridrow="7" pageId="8" pageNumber="9">duration (ms)</th>
<td id="C56B2A66FFCC0072D0BFF997FCBE390D" box="[645,773,1567,1589]" gridcol="1" gridrow="7" pageId="8" pageNumber="9">471±105</td>
<td id="C56B2A66FFCC0072D148F997FC77390D" box="[882,972,1567,1589]" gridcol="2" gridrow="7" pageId="8" pageNumber="9">408±29</td>
<td id="C56B2A66FFCC0072D641F997FAB3390D" box="[1147,1288,1567,1589]" gridcol="3" gridrow="7" pageId="8" pageNumber="9">365±33</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F9C0FAB33966" box="[159,1288,1608,1630]" gridrow="8" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F9C0FD833966" box="[159,568,1608,1630]" gridcol="0" gridrow="8" pageId="8" pageNumber="9">avg beat interval</th>
<td id="C56B2A66FFCC0072D0BFF9C0FCBE3966" box="[645,773,1608,1630]" gridcol="1" gridrow="8" pageId="8" pageNumber="9">60</td>
<td id="C56B2A66FFCC0072D148F9C0FC773966" box="[882,972,1608,1630]" gridcol="2" gridrow="8" pageId="8" pageNumber="9">57</td>
<td id="C56B2A66FFCC0072D641F9C0FAB33966" box="[1147,1288,1608,1630]" gridcol="3" gridrow="8" pageId="8" pageNumber="9">47</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F9F9FAB339BF" box="[159,1288,1649,1671]" gridrow="9" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F9F9FD8339BF" box="[159,568,1649,1671]" gridcol="0" gridrow="9" pageId="8" pageNumber="9">♂ response time</th>
<td id="C56B2A66FFCC0072D0BFF9F9FCBE39BF" box="[645,773,1649,1671]" gridcol="1" gridrow="9" pageId="8" pageNumber="9">157±92</td>
<td id="C56B2A66FFCC0072D148F9F9FC7739BF" box="[882,972,1649,1671]" gridcol="2" gridrow="9" pageId="8" pageNumber="9">168±101</td>
<td id="C56B2A66FFCC0072D641F9F9FAB339BF" box="[1147,1288,1649,1671]" gridcol="3" gridrow="9" pageId="8" pageNumber="9">129±68</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F913FAB33989" box="[159,1288,1691,1713]" gridrow="10" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F913FD833989" box="[159,568,1691,1713]" gridcol="0" gridrow="10" pageId="8" pageNumber="9">♂ REPLY nr. of beats</th>
<td id="C56B2A66FFCC0072D0BFF913FCBE3989" box="[645,773,1691,1713]" gridcol="1" gridrow="10" pageId="8" pageNumber="9">14.5±2.5</td>
<td id="C56B2A66FFCC0072D148F913FC773989" box="[882,972,1691,1713]" gridcol="2" gridrow="10" pageId="8" pageNumber="9">10.1±1.5</td>
<td id="C56B2A66FFCC0072D641F913FAB33989" box="[1147,1288,1691,1713]" gridcol="3" gridrow="10" pageId="8" pageNumber="9">11.7±3.0</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F94CFAB339E2" box="[159,1288,1732,1754]" gridrow="11" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F94CFD8339E2" box="[159,568,1732,1754]" gridcol="0" gridrow="11" pageId="8" pageNumber="9">duration (ms)</th>
<td id="C56B2A66FFCC0072D0BFF94CFCBE39E2" box="[645,773,1732,1754]" gridcol="1" gridrow="11" pageId="8" pageNumber="9">767±188</td>
<td id="C56B2A66FFCC0072D148F94CFC7739E2" box="[882,972,1732,1754]" gridcol="2" gridrow="11" pageId="8" pageNumber="9">462±84</td>
<td id="C56B2A66FFCC0072D641F94CFAB339E2" box="[1147,1288,1732,1754]" gridcol="3" gridrow="11" pageId="8" pageNumber="9">493±166</td>
</tr>
<tr id="86BA431AFFCC0072D2A5F965FAB3383B" box="[159,1288,1773,1795]" gridrow="12" pageId="8" pageNumber="9">
<th id="C56B2A66FFCC0072D2A5F965FD83383B" box="[159,568,1773,1795]" gridcol="0" gridrow="12" pageId="8" pageNumber="9">avg beat interval</th>
<td id="C56B2A66FFCC0072D0BFF965FCBE383B" box="[645,773,1773,1795]" gridcol="1" gridrow="12" pageId="8" pageNumber="9">57</td>
<td id="C56B2A66FFCC0072D148F965FC77383B" box="[882,972,1773,1795]" gridcol="2" gridrow="12" pageId="8" pageNumber="9">46</td>
<td id="C56B2A66FFCC0072D641F965FAB3383B" box="[1147,1288,1773,1795]" gridcol="3" gridrow="12" pageId="8" pageNumber="9">42</td>
</tr>
</table>
</paragraph>
<subSubSection id="709012D3FFCCFF84D2FDF8C0FE4538F4" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="distribution">
<paragraph id="38354158FFCCFF84D2FDF8C0FAF03E07" blockId="8.[151,1437,1864,2033]" lastBlockId="9.[151,1437,151,320]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="0AFE9D4AFFCCFF85D2FDF8C0FE473859" bold="true" box="[199,508,1864,1889]" pageId="8" pageNumber="9">Distribution and ecology.</emphasis>
<taxonomicName id="FF8A3ADBFFCCFF85D03DF8C0FCB33858" box="[519,776,1864,1888]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFCCFF85D03DF8C0FCB33858" box="[519,776,1864,1888]" italics="true" pageId="8" pageNumber="9">Zwicknia westermanni</emphasis>
</taxonomicName>
is known from
<collectingCountry id="409D01C8FFCCFF85D1F7F8C0FB8E3858" box="[973,1077,1864,1888]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
(southern Lower Saxony) and
<collectingCountry id="409D01C8FFCCFF85D2ADF8E5FF5838BC" box="[151,227,1901,1924]" name="France" pageId="8" pageNumber="9">France</collectingCountry>
(Burgundy; Provence-Alpes-Côte d’Azur) (
<figureCitation id="A0B15DDDFFCCFF85D0CDF8E4FCEF38BD" box="[759,852,1900,1925]" captionStart="FIGURE 22" captionStartId="13.[151,250,1126,1148]" captionTargetBox="[312,1274,451,1090]" captionTargetId="figure@13.[312,1275,451,1091]" captionTargetPageId="13" captionText="FIGURE 22. Known distribution of Zwicknia westermanni Boumans &amp; Murányi, sp. n. (full circles, type locality indicated with a central dot) and Z. komica Murányi &amp; Boumans, sp. n. (open circle) — known area of the genus Zwicknia Murányi, 2014 indicated in red, grey areas are above 1000 m sea level." httpUri="https://zenodo.org/record/286177/files/figure.png" pageId="8" pageNumber="9">Fig. 22</figureCitation>
).
<bibRefCitation id="5C1B3CA9FFCCFF85D14AF8E5FBEB38BD" author="Westermann" box="[880,1104,1901,1925]" pageId="8" pageNumber="9" refString="Westermann, F. (1990) Limnologische Untersuchungen der Makrobenthosfauna unversauerter und versauerter Bache im Hils (Weser-Leine-Bergland, Sudniedersachsen) [unpublished thesis]. II. Zoologisches Institut. Georg-August-Universitat zu Gottingen, Gottingen." type="book" year="1990">Westermann (1990</bibRefCitation>
,
<bibRefCitation id="5C1B3CA9FFCCFF85D65FF8E5FB2538BD" author="Westermann" box="[1125,1182,1901,1925]" pageId="8" pageNumber="9" refString="Westermann, F. (1993) Wing polymorphism in Capnia bifrons (Plecoptera: Capniidae). Aquatic Insects, 15, 135 - 140. http: // dx. doi. org / 10.1080 / 01650429309361510" type="journal article" year="1993">1993</bibRefCitation>
) provides a detailed description of the
<typeStatus id="E731FFFAFFCCFF85D351F819FE203891" box="[363,411,1937,1961]" pageId="8" pageNumber="9">type</typeStatus>
locality stream, the summer-dry Hohenförsiek stream in the Hils Massif,
<collectingCountry id="409D01C8FFCCFF85D6C3F818FAD83890" box="[1273,1379,1936,1960]" name="Germany" pageId="8" pageNumber="9">Germany</collectingCountry>
. The species was found in sympatry with
<taxonomicName id="FF8A3ADBFFCCFF85D00AF83DFD2738F4" box="[560,668,1972,1996]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFCCFF85D00AF83DFD2738F4" box="[560,668,1972,1996]" italics="true" pageId="8" pageNumber="9">Z. bifrons</emphasis>
</taxonomicName>
at a lower site in the Hohenförsiek stream (N 5158.813'
<geoCoordinate id="5DBE279FFFCCFF85D720F83DFA2938F5" box="[1306,1426,1973,1997]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="55" value="941.732">E 941.732</geoCoordinate>
') in the Hils Massif. In the Boisserand stream in Curgy, Burgundy,
<collectingCountry id="409D01C8FFCCFF85D19CF851FC4938C8" box="[934,1010,2009,2032]" name="France" pageId="8" pageNumber="9">France</collectingCountry>
(N 4659.411'
<geoCoordinate id="5DBE279FFFCCFF85D6A5F851FAA538C9" box="[1183,1310,2008,2033]" direction="east" orientation="longitude" pageId="8" pageNumber="9" precision="55" value="422.852">E 422.852</geoCoordinate>
'), it occurs together with
<taxonomicName id="FF8A3ADBFFCDFF84D30EFF11FE733F88" box="[308,456,152,176]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="0AFE9D4AFFCDFF84D30EFF11FE733F88" box="[308,456,152,176]" italics="true" pageId="9" pageNumber="10">Z. rupprechti</emphasis>
</taxonomicName>
.
<bibRefCitation id="5C1B3CA9FFCDFF84D3EDFF1FFD363F88" author="Haybach" box="[471,653,151,176]" pageId="9" pageNumber="10" refString="Haybach, A. (2004) Zur Kenntnis der Steinfliegenfauna (Insecta: Plecoptera) des Oberen Naheberglandes. Mainzer Naturwissenschaftliches Archiv, 42, 89 - 98." type="journal article" year="2004">Haybach (2004)</bibRefCitation>
reports the occurrence micropterous males of ‘
<taxonomicName id="FF8A3ADBFFCDFF84D6A5FF1FFB483F88" box="[1183,1267,151,176]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">Capnia</taxonomicName>
bifrons’ in the lower part, and brachypterous ones in the upper part of a summer-dry stream in Rhineland-Palatinate,
<collectingCountry id="409D01C8FFCDFF84D715FF34FA223FEC" box="[1327,1433,188,212]" name="Germany" pageId="9" pageNumber="10">Germany</collectingCountry>
. This mirrors the situation we found in the
<typeStatus id="E731FFFAFFCDFF84D09FFF68FD6E3FC0" box="[677,725,224,248]" pageId="9" pageNumber="10">type</typeStatus>
locality stream of
<taxonomicName id="FF8A3ADBFFCDFF84D1F9FF68FBD33FCF" box="[963,1128,224,247]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFCDFF84D1F9FF68FBD33FCF" box="[963,1128,224,247]" italics="true" pageId="9" pageNumber="10">Z.westermanni</emphasis>
</taxonomicName>
, and points to ecological differences between species. We did not study Haybach’s material, yet it is plausible that his brachypterous specimens belong to
<taxonomicName id="FF8A3ADBFFCDFF84D3B8FEA1FD943E07" box="[386,559,296,320]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="westermanni">
<emphasis id="0AFE9D4AFFCDFF84D3B8FEA1FD943E07" box="[386,559,296,320]" italics="true" pageId="9" pageNumber="10">Z. westermanni</emphasis>
</taxonomicName>
, while the micropterous ones may be either
<taxonomicName id="FF8A3ADBFFCDFF84D61AFEA1FB303E78" box="[1056,1163,296,320]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="bifrons">
<emphasis id="0AFE9D4AFFCDFF84D61AFEA1FB303E78" box="[1056,1163,296,320]" italics="true" pageId="9" pageNumber="10">Z. bifrons</emphasis>
</taxonomicName>
or
<taxonomicName id="FF8A3ADBFFCDFF84D68BFEA1FAFF3E78" box="[1201,1348,296,320]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rupprechti">
<emphasis id="0AFE9D4AFFCDFF84D68BFEA1FAFF3E78" box="[1201,1348,296,320]" italics="true" pageId="9" pageNumber="10">Z. rupprechti</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="6CF511D0FFCDFF84D2ADFD40FE513DC5" httpUri="https://zenodo.org/record/286173/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1436,373,691]" targetPageId="9">
<paragraph id="38354158FFCDFF84D2ADFD40FE513DC5" blockId="9.[151,1436,712,765]" pageId="9" pageNumber="10">
<emphasis id="0AFE9D4AFFCDFF84D2ADFD40FEA83DE6" bold="true" box="[151,275,712,734]" pageId="9" pageNumber="10">FIGURE 7.</emphasis>
Oscillogram showing an excerpt of a vibration duet of a pair of
<taxonomicName id="FF8A3ADBFFCDFF84D199FD42FA9B3DE6" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[931,1312,712,735]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFCDFF84D199FD42FBFB3DE6" box="[931,1088,713,735]" italics="true" pageId="9" pageNumber="10">Z. westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFCDFF84D711FD41FADF3DE6" box="[1323,1380,713,734]" pageId="9" pageNumber="10" rank="species">
<emphasis id="0AFE9D4AFFCDFF84D711FD41FADF3DE6" bold="true" box="[1323,1380,713,734]" pageId="9" pageNumber="10">sp. n.</emphasis>
</taxonomicNameLabel>
from the Hohenförsiek stream at 20°C.
</paragraph>
</caption>
<caption id="6CF511D0FFCDFF84D2ADF882FBD23807" httpUri="https://zenodo.org/record/286174/files/figure.png" pageId="9" pageNumber="10" targetBox="[276,1313,815,1773]" targetPageId="9">
<paragraph id="38354158FFCDFF84D2ADF882FBD23807" blockId="9.[151,1436,1802,1855]" pageId="9" pageNumber="10">
<emphasis id="0AFE9D4AFFCDFF84D2ADF882FEA93827" bold="true" box="[151,274,1802,1824]" pageId="9" pageNumber="10">FIGURE 8.</emphasis>
Beat interval length as a function of interval number in a duet of
<taxonomicName id="FF8A3ADBFFCDFF84D19DF883FA993818" authority="Boumans &amp; Muranyi" authorityName="Boumans &amp; Muranyi" box="[935,1314,1802,1824]" class="Insecta" family="Capniidae" genus="Zwicknia" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="westermanni" status="sp. nov.">
<emphasis id="0AFE9D4AFFCDFF84D19DF883FBF93818" box="[935,1090,1803,1824]" italics="true" pageId="9" pageNumber="10">Z. westermanni</emphasis>
Boumans &amp; Murányi
</taxonomicName>
,
<taxonomicNameLabel id="11CD2031FFCDFF84D716F883FADE3818" box="[1324,1381,1803,1824]" pageId="9" pageNumber="10" rank="species">
<emphasis id="0AFE9D4AFFCDFF84D716F883FADE3818" bold="true" box="[1324,1381,1803,1824]" pageId="9" pageNumber="10">sp. n.</emphasis>
</taxonomicNameLabel>
from the Hohenförsiek stream at 20°C. Data points show the median value of seven measured signals.
</paragraph>
</caption>
<paragraph id="38354158FFCDFF84D2FDF8E3FE4538F4" blockId="9.[151,1437,1899,2032]" pageId="9" pageNumber="10">
<emphasis id="0AFE9D4AFFCDFF84D2FDF8E3FEF238BC" bold="true" box="[199,329,1899,1924]" pageId="9" pageNumber="10">Phenology.</emphasis>
In the Hils Massif, adults have been collected from late January to mid-March (
<bibRefCitation id="5C1B3CA9FFCDFF84D732F8E4FF683890" author="Westermann" pageId="9" pageNumber="10" refString="Westermann, F. (1993) Wing polymorphism in Capnia bifrons (Plecoptera: Capniidae). Aquatic Insects, 15, 135 - 140. http: // dx. doi. org / 10.1080 / 01650429309361510" type="journal article" year="1993">Westermann, 1993</bibRefCitation>
). A few specimens reared in the laboratory emerged as late as
<date id="4C346798FFCDFF84D199F818FBAE3890" box="[931,1045,1935,1960]" pageId="9" pageNumber="10" value="1930-03">30 March</date>
. One female lived for two months after emergence in early March.
</paragraph>
</subSubSection>
<subSubSection id="709012D3FFCDFF87D2FDF85FFB323FC0" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="etymology">
<paragraph id="38354158FFCDFF87D2FDF85FFB323FC0" blockId="9.[151,1437,1899,2032]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="0AFE9D4AFFCDFF84D2FDF85FFEF538C8" bold="true" box="[199,334,2007,2032]" pageId="9" pageNumber="10">Etymology.</emphasis>
The species is named after Fulgor Westermann, Mainz,
<collectingCountry id="409D01C8FFCDFF84D1E2F850FBFA38C8" box="[984,1089,2008,2032]" name="Germany" pageId="9" pageNumber="10">Germany</collectingCountry>
, who described the distinctive wing length of the males of this species and the habitat characteristics of the
<typeStatus id="E731FFFAFFCEFF87D1CCFF10FB9D3F88" box="[1014,1062,152,176]" pageId="10" pageNumber="11">type</typeStatus>
locality (
<bibRefCitation id="5C1B3CA9FFCEFF87D6ABFF10FAD33F88" author="Westermann" box="[1169,1384,152,176]" pageId="10" pageNumber="11" refString="Westermann, F. (1993) Wing polymorphism in Capnia bifrons (Plecoptera: Capniidae). Aquatic Insects, 15, 135 - 140. http: // dx. doi. org / 10.1080 / 01650429309361510" type="journal article" year="1993">Westermann, 1993</bibRefCitation>
). He also collected some of the
<typeStatus id="E731FFFAFFCEFF87D3F8FF35FE493FED" box="[450,498,189,213]" pageId="10" pageNumber="11">type</typeStatus>
specimens. His advice and assistance with the fieldwork in the Hils area have been a major contribution. The specific epithet is used as the genitive of a noun of male gender.
</paragraph>
</subSubSection>
</treatment>
</document>
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