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<mods:title id="B79371CF06D6D2F5525D9FF8D441E3EB">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:namePart id="A624E07370C2DB1BA13FBCA33C631C9A">Wu, Jingyu</mods:namePart>
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<mods:affiliation id="811E6C12D6A45543802EB3B372A61B8B">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:affiliation id="245B508959A1E3ED67A5381EC7DBB6C1">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<subSubSection id="C30B650DFFB5FFB79EE7C92F9DED4A74" box="[151,715,1837,1864]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BAE3686FFB5FFB79EE7C92F9DED4A74" blockId="18.[151,715,1837,1899]" box="[151,715,1837,1864]" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EE7C92F9DED4A74" bold="true" box="[151,715,1837,1864]" pageId="18" pageNumber="19">
[5]
<taxonomicName id="4C114D05FFB5FFB79EB1C92F9DED4A74" ID-CoL="66SKQ" authority="Tsukaguchi, 1995" authorityName="Tsukaguchi" authorityYear="1995" box="[193,715,1837,1864]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B965EA94FFB5FFB79EB1C92F9ECC4A74" bold="true" box="[193,490,1837,1864]" italics="true" pageId="18" pageNumber="19">Ankylopteryx ferruginea</emphasis>
<bibRefCitation id="EF804B77FFB5FFB79F81C92F9DED4A74" author="Tsukaguchi, S." box="[497,715,1837,1863]" pageId="18" pageNumber="19" refId="ref91372" refString="Tsukaguchi, S. (1995) Chrysopidae of Japan (Insecta, Neuroptera). Privately printed, Osaka, 224 pp." type="book" year="1995">Tsukaguchi, 1995</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
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<paragraph id="8BAE3686FFB5FFB79EE7C9539E014A58" blockId="18.[151,715,1837,1899]" box="[151,295,1873,1899]" pageId="18" pageNumber="19">
(
<figureCitation id="132A2A03FFB5FFB79EEFC9539FD04A58" box="[159,246,1873,1899]" captionStart="FIGURE 10" captionStartId="20.[151,250,1904,1928]" captionTargetBox="[219,1400,212,1859]" captionTargetId="figure-33@20.[183,1404,181,1880]" captionTargetPageId="20" captionText="FIGURE 10. Ankylopteryx ferruginea Tsukaguchi, 1995. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcus and entoprocessus, dorsal view; G. Gonarcus and entoprocessus, lateral view; H. Pseudopenis, dorsal view; I. Pseudopenis, lateral view; J. Female genitalia, lateral view; K. Subgenitale; L. Spermatheca. Scale bar: 5 mm (A), 1 mm (BD)." figureDoi="http://doi.org/10.5281/zenodo.14240528" httpUri="https://zenodo.org/record/14240528/files/figure.png" pageId="18" pageNumber="19">Figs 10</figureCitation>
,
<figureCitation id="132A2A03FFB5FFB79F72C9539E394A58" box="[258,287,1873,1899]" captionStart="FIGURE 79" captionStartId="142.[151,250,1894,1918]" captionTargetBox="[196,1392,1098,1870]" captionTargetId="figure-46@142.[196,1392,1098,1870]" captionTargetPageId="142" captionText="FIGURE 79. Distribution map of species of Ankylopteryx from China." figureDoi="http://doi.org/10.5281/zenodo.14240704" httpUri="https://zenodo.org/record/14240704/files/figure.png" pageId="18" pageNumber="19">79</figureCitation>
)
</paragraph>
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<paragraph id="8BAE3686FFB5FFB79EE7C9949A164A9D" blockId="18.[151,1328,1942,1966]" box="[151,1328,1942,1966]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB0111A8FFB5FFB79EE7C9949A164A9D" box="[151,1328,1942,1966]" pageId="18" pageNumber="19">
<treatmentCitation id="0AB01097FFB5FFB79EE7C9949D584A9D" author="Tsukaguchi, S." box="[151,638,1942,1966]" page="127" pageId="18" pageNumber="19" year="1995">
<taxonomicName id="4C114D05FFB5FFB79EE7C9949D584A9D" ID-CoL="66SKQ" authority="Tsukaguchi, 1995: 127" authorityName="Tsukaguchi" authorityPageNumber="127" authorityYear="1995" box="[151,638,1942,1966]" class="Insecta" family="Chrysopidae" genus="Ankylopteryx" kingdom="Animalia" order="Neuroptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="B965EA94FFB5FFB79EE7C9949EA84A9D" box="[151,398,1942,1966]" italics="true" pageId="18" pageNumber="19">Ankylopteryx ferruginea</emphasis>
<materialsCitation id="3B793CDBFFB5FFB79FE4C9949D584A9D" box="[404,638,1942,1966]" collectionCode="UOP" country="Japan" location="Iriomote Island" pageId="18" pageNumber="19" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<bibRefCitation id="EF804B77FFB5FFB79FE4C9949D584A9D" author="Tsukaguchi, S." box="[404,638,1942,1966]" pageId="18" pageNumber="19" refId="ref91372" refString="Tsukaguchi, S. (1995) Chrysopidae of Japan (Insecta, Neuroptera). Privately printed, Osaka, 224 pp." type="book" year="1995">Tsukaguchi, 1995: 127</bibRefCitation>
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. Type locality:
<collectingCountry id="F3067616FFB5FFB79D69C9949C754A9E" box="[793,851,1942,1965]" name="Japan" pageId="18" pageNumber="19">Japan</collectingCountry>
(
<collectingRegion id="49D5F864FFB5FFB79D10C9949C994A9D" box="[864,959,1942,1966]" country="Japan" name="Okinawa" pageId="18" pageNumber="19">Okinawa</collectingRegion>
,
<location id="8ECE605DFFB5FFB79DBAC9949B4F4A9D" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB5FFB79DBAC9949B4F4A9D" box="[970,1129,1942,1966]" country="Japan" name="Iriomote Island" pageId="18" pageNumber="19" stateProvince="Okinawa">Iriomote Island</location>
).
<typeStatus id="54AA8824FFB5FFB79A0AC9949BFE4A9D" box="[1146,1240,1942,1966]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
in UOP.
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</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB5FFB69EE7C9DE9DD84DCA" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BAE3686FFB5FFB69EE7C9DE9DD84DCA" blockId="18.[151,1437,2012,2039]" lastBlockId="19.[151,1437,151,1977]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B965EA94FFB5FFB79EE7C9DE9E344AC5" bold="true" box="[151,274,2012,2038]" pageId="18" pageNumber="19">Diagnosis.</emphasis>
This species is characterized by the presence of the scape and pedicel without stripes, the small pale brown spots present on the anterolateral angle of pronotum, the pro-, meso- and metatibia with median black spot, the triangular cell
<emphasis id="B965EA94FFB4FFB69F2FCEB99E5D4DE7" box="[351,379,187,212]" italics="true" pageId="19" pageNumber="20">im</emphasis>
, the arched inner gradate series, the male entoprocessus attached at about basal 1/3 of gonarcus, and the narrowed and projected apex of entoprocessus.
</paragraph>
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<subSubSection id="C30B650DFFB4FFB69EB7CF009D964922" pageId="19" pageNumber="20" type="description">
<paragraph id="8BAE3686FFB4FFB69EB7CF009AA94C2E" blockId="19.[151,1437,151,1977]" box="[199,1423,258,285]" pageId="19" pageNumber="20">
<emphasis id="B965EA94FFB4FFB69EB7CF009E7F4C2F" bold="true" box="[199,345,258,284]" pageId="19" pageNumber="20">Description.</emphasis>
Male. Body length
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, forewing length 10.0
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, hindwing length
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.
</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CF259B604C56" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Head yellow, vertex raised. Compound eyes brown. Antennal scape and pedicel without stripes. Clypeal margin with brown stripes. Gena without spots. Maxillary and labial palpi brownish yellow.</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CF6D9D844C9E" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Pronotum anterior margin shorter than posterior margin, with small pale brown spots on anterolateral angle; mesonotum and metanotum without markings.</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CFB59FD24CC6" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Legs with dense setae. Pro-, meso- and metatibia with median black spot. Pretarsal claws brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CFFD9A4D4FB6" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">
Forewing broad, with brown shadings on base of Rs, psm-psc crossveins and CuP. Pterostigma brown. Costal area broad basally, with 2024 crossveins; 910 radial crossveins and 7 psm-psc crossveins present.
<emphasis id="B965EA94FFB4FFB69B6ECC219A674F0E" box="[1310,1345,547,573]" italics="true" pageId="19" pageNumber="20">m1</emphasis>
smaller than
<emphasis id="B965EA94FFB4FFB69EBECC459FD64F52" box="[206,240,583,609]" italics="true" pageId="19" pageNumber="20">m2</emphasis>
,
<emphasis id="B965EA94FFB4FFB69E8DCC459E3F4F53" box="[253,281,583,608]" italics="true" pageId="19" pageNumber="20">im</emphasis>
triangular.
<emphasis id="B965EA94FFB4FFB69FE8CC459E954F52" box="[408,435,583,609]" italics="true" pageId="19" pageNumber="20">c1</emphasis>
smaller than
<emphasis id="B965EA94FFB4FFB69C38CC459D454F52" box="[584,611,583,609]" italics="true" pageId="19" pageNumber="20">c2</emphasis>
,
<emphasis id="B965EA94FFB4FFB69C00CC459DB14F52" box="[624,663,583,609]" italics="true" pageId="19" pageNumber="20">dcc</emphasis>
opened. A1 forked, A2 and A3 simple. Gradate series with small pale brown shadings, inner gradate series arched and with 47 crossveins, outer gradate series with 68 crossveins.
</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CC8D9BFA4FC2" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Hindwing narrow, with brown shadings on base of Rs and middle posterior margin crossveins. Pterostigma pale brown. Costal area with 1516 crossveins; 89 radial crossveins and 6 psm-psc crossveins present. A1 forked, A2 and A3 simple. Inner gradate series with 35 crossveins, outer gradate series with 57 crossveins.</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CCF99C184EB2" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Abdomen with dense short setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; sternum VIII fused with sternum IX; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, lateral arms narrow at base and wide at tip; entoprocessus attached at about basal 1/3 of gonarcus, narrowed and projected at tip; pseudopenis widest near tip and curved.</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CD899A0C4E96" blockId="19.[151,1437,151,1977]" box="[199,1322,907,933]" pageId="19" pageNumber="20">
Female. Body length 7.210.0 mm, forewing length
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, hindwing length
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.
</paragraph>
<paragraph id="8BAE3686FFB4FFB69EB7CDAD9D964922" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">Sternum VII longer than tergum VII; tergum VII subquadrate in lateral view; tergum VIII shorter than tergum VII; tergum IX fused with ectoproct. Callus cerci ovoid. Subgenitale bilobed apically; spermatheca thick; vela and ventral impression absent; duct long and coiled.</paragraph>
</subSubSection>
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<emphasis id="B965EA94FFB4FFB69EB7CA189E104907" bold="true" box="[199,310,1050,1076]" pageId="19" pageNumber="20">Material.</emphasis>
<materialsCitation id="3B793CDBFFB4FFB69F4ECA199A624906" box="[318,1348,1050,1077]" collectingDate="2020-05-20" collectionCode="CAU" collectorName="Yuchen Zheng" country="China" county="Zhangzhou" elevation="250" location="Xiangxi" municipality="Nanjing" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Fujian">
<specimenCount id="9D17FD0FFFB4FFB69F4ECA199E444906" box="[318,354,1051,1077]" count="2" pageId="19" pageNumber="20" type="female">2♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69F1FCA199E934906" box="[367,437,1051,1077]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69FB0CA199D2D4906" box="[448,523,1051,1077]" country="China" name="Fujian" pageId="19" pageNumber="20">Fujian</collectingRegion>
,
<collectingCounty id="62CF4E0AFFB4FFB69C66CA199DB14906" box="[534,663,1051,1077]" pageId="19" pageNumber="20">Zhangzhou</collectingCounty>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69CD2CA199DD94906" box="[674,767,1051,1077]" pageId="19" pageNumber="20">Nanjing</collectingMunicipality>
,
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)
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<materialsCitation id="3B793CDBFFB4FFB69C0ACB5D9D9F48AE" collectingDate="1954-05-17" collectionCode="CAU" collectorName="Fasehng Li" country="China" county="Chongzuo" elevation="180" location="Longrui National Nature Reserve" municipality="Ningming" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi">
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<collectingCounty id="62CF4E0AFFB4FFB69D07CB5D9CCB484A" box="[887,1005,1375,1401]" pageId="19" pageNumber="20">Chongzuo</collectingCounty>
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)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69CB6CB819D7C48F2" collectingDate="2013-04-15" collectionCode="CAU" collectorName="Jianyun Wang &amp; Pei Jiang" country="China" location="Yinggeling National Nature Reserve" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Hainan">
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<location id="8ECE605DFFB4FFB69DDDCB819A7648AE" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69DDDCB819A7648AE" box="[941,1360,1411,1437]" country="China" name="Yinggeling National Nature Reserve" pageId="19" pageNumber="20" stateProvince="Hainan">Yinggeling National Nature Reserve</location>
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<collectingDate id="EFEBE9AEFFB4FFB69B2DCB819FF148F2" pageId="19" pageNumber="20" value="2013-04-15">2013. IV.15</collectingDate>
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<collectorName id="26E45350FFB4FFB69FEFCBA59D2548F2" box="[415,515,1447,1473]" pageId="19" pageNumber="20">Pei Jiang</collectorName>
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)
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;
<materialsCitation id="3B793CDBFFB4FFB69C14CBA59EBD48D6" collectingDate="2014-07-26" collectionCode="CAU" collectorName="Lu Yue" country="China" elevation="512" location="Maolan National Nature Reserve" municipality="Libo" pageId="19" pageNumber="20" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
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<location id="8ECE605DFFB4FFB69DE1CBA59A2748F2" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69DE1CBA59A2748F2" box="[913,1281,1447,1473]" country="China" municipality="Libo" name="Maolan National Nature Reserve" pageId="19" pageNumber="20" stateProvince="Guizhou">Maolan National Nature Reserve</location>
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<collectingDate id="EFEBE9AEFFB4FFB69B2DCBA59FC548D6" pageId="19" pageNumber="20" value="2014-07-26">2014. VII.26</collectingDate>
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<materialsCitation id="3B793CDBFFB4FFB69FD8CBC99EEE4B3A" collectingDate="2019-04-28" collectingDateMax="2019-04-29" collectingDateMin="2019-04-28" collectorName="Hongyu Li" country="China" county="Xishuangbanna" elevation="780" location="No." municipality="Menglun" pageId="19" pageNumber="20" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Yunnan">
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<location id="8ECE605DFFB4FFB69D95CBC99B2848D6" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69D95CBC99B2848D6" box="[997,1038,1483,1509]" country="China" county="Xishuangbanna" municipality="Menglun" name="No." pageId="19" pageNumber="20" stateProvince="Yunnan">No.</location>
55 experimental site, light,
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<collectingDate id="EFEBE9AEFFB4FFB69EE7CBED9E664B3A" box="[151,320,1519,1545]" pageId="19" pageNumber="20" value="2019-04-28" valueMax="2019-04-29" valueMin="2019-04-28">2019.IV.2829</collectingDate>
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<materialsCitation id="3B793CDBFFB4FFB69FA3CBED9E0A4B1E" collectionCode="CAU" collectorName="Yunlan Jiang" country="China" county="Xishuangbanna" elevation="680" location="Yexianggu" municipality="Mengyang" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
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<collectorName id="26E45350FFB4FFB69B3DCBED9FF44B1E" pageId="19" pageNumber="20">Yunlan Jiang</collectorName>
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<collectingMunicipality id="6BCAACFCFFB4FFB69FADC8599D084B46" box="[477,558,1627,1653]" pageId="19" pageNumber="20">Qujing</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69C4AC8599DA64B46" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69C4AC8599DA64B46" box="[570,640,1627,1653]" country="China" municipality="Qujing" name="Huize" pageId="19" pageNumber="20" stateProvince="Yunnan">Huize</location>
,
<quantity id="4CE99B63FFB4FFB69CFCC8599DF04B46" box="[652,726,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="m" value="800.0">
<elevation id="003CD1B5FFB4FFB69CFCC8599DF04B46" box="[652,726,1627,1653]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="m" value="800.0">800 m</elevation>
</quantity>
,
<date id="FFAF1046FFB4FFB69C93C8599C784B46" box="[739,862,1627,1653]" pageId="19" pageNumber="20" value="1979-04-18">
<collectingDate id="EFEBE9AEFFB4FFB69C93C8599C784B46" box="[739,862,1627,1653]" pageId="19" pageNumber="20" value="1979-04-18">1979.IV.18</collectingDate>
</date>
(
<collectionCode id="ED00AE43FFB4FFB69D1FC8599C964B46" box="[879,944,1627,1653]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69DB4C8599E1F4BAA" collectingDate="2017-07-20" collectionCode="CAU" country="China" location="Lancang" municipality="Puer" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D17FD0FFFB4FFB69DB4C8599CC14B46" box="[964,999,1627,1653]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69D87C8599B1B4B46" box="[1015,1085,1627,1653]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69A38C8599B874B46" box="[1096,1185,1627,1653]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69ADCC8599BC74B46" box="[1196,1249,1627,1653]" pageId="19" pageNumber="20">Puer</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69A9EC8599A774B46" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69A9EC8599A774B46" box="[1262,1361,1627,1653]" country="China" municipality="Puer" name="Lancang" pageId="19" pageNumber="20" stateProvince="Yunnan">Lancang</location>
,
<date id="FFAF1046FFB4FFB69B2DC8599FC74BAA" pageId="19" pageNumber="20" value="2017-07-20">
<collectingDate id="EFEBE9AEFFB4FFB69B2DC8599FC74BAA" pageId="19" pageNumber="20" value="2017-07-20">2017. VII.20</collectingDate>
</date>
(
<collectionCode id="ED00AE43FFB4FFB69E80C87D9E174BAA" box="[240,305,1663,1689]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69F34C87D9E334B8E" collectingDate="2019-10-05" collectingDateMax="2019-10-06" collectingDateMin="2019-10-05" collectionCode="CAU" collectorName="Hongyu Li" country="China" elevation="600" location="Nangunhe National Nature Reserve" municipality="Cangyuan" pageId="19" pageNumber="20" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan">
<specimenCount id="9D17FD0FFFB4FFB69F34C87D9E4E4BAA" box="[324,360,1663,1689]" count="2" pageId="19" pageNumber="20" type="female">2♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69F04C87D9E9C4BAA" box="[372,442,1663,1689]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69FB3C87D9D3A4BAA" box="[451,540,1663,1689]" country="China" name="Yunnan" pageId="19" pageNumber="20">Yunnan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69C57C87D9DBD4BAA" box="[551,667,1663,1689]" pageId="19" pageNumber="20">Cangyuan</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69CD6C87D9B104BAA" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69CD6C87D9B104BAA" box="[678,1078,1663,1689]" country="China" municipality="Cangyuan" name="Nangunhe National Nature Reserve" pageId="19" pageNumber="20" stateProvince="Yunnan">Nangunhe National Nature Reserve</location>
,
<quantity id="4CE99B63FFB4FFB69A30C87D9BAF4BAA" box="[1088,1161,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="m" value="600.0">
<elevation id="003CD1B5FFB4FFB69A30C87D9BAF4BAA" box="[1088,1161,1663,1689]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="19" pageNumber="20" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FFAF1046FFB4FFB69AE4C87C9A1E4BAA" box="[1172,1336,1662,1689]" pageId="19" pageNumber="20" value="2019-10-05" valueMax="2019-10-06" valueMin="2019-10-05">
<collectingDate id="EFEBE9AEFFB4FFB69AE4C87C9A1E4BAA" box="[1172,1336,1662,1689]" pageId="19" pageNumber="20" value="2019-10-05" valueMax="2019-10-06" valueMin="2019-10-05">2019.X.0506</collectingDate>
</date>
,
<collectorName id="26E45350FFB4FFB69B32C87D9F964B8E" pageId="19" pageNumber="20">Hongyu Li</collectorName>
(
<collectionCode id="ED00AE43FFB4FFB69EB2C8A19E2A4B8E" box="[194,268,1699,1725]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69F6AC8A19A434B8E" box="[282,1381,1699,1725]" collectingDate="2019-06-13" collectionCode="CAU" collectorName="Xingyue Liu" country="China" elevation="400" location="Guanyin Waterfall" municipality="Ren'ai" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Nantou">
<specimenCount id="9D17FD0FFFB4FFB69F6AC8A19E154B8E" box="[282,307,1699,1725]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69F31C8A19EA14B8E" box="[321,391,1699,1725]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
,
<collectingCountry id="F3067616FFB4FFB69FE4C8A19ECF4B8E" box="[404,489,1699,1725]" name="Taiwan" pageId="19" pageNumber="20">Taiwan</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69F86C8A19D6D4B8E" box="[502,587,1699,1725]" country="Taiwan" name="Nantou" pageId="19" pageNumber="20">Nantou</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69C28C8A19D834B8E" box="[600,677,1699,1725]" pageId="19" pageNumber="20">Renai</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69CC2C8A19CA04B8E" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69CC2C8A19CA04B8E" box="[690,902,1699,1725]" country="China" municipality="Ren'ai" name="Guanyin Waterfall" pageId="19" pageNumber="20" stateProvince="Nantou">Guanyin Waterfall</location>
,
<quantity id="4CE99B63FFB4FFB69DE3C8A19CF84B8E" box="[915,990,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="400.0">
<elevation id="003CD1B5FFB4FFB69DE3C8A19CF84B8E" box="[915,990,1699,1725]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<date id="FFAF1046FFB4FFB69D9BC8A19B4B4B8E" box="[1003,1133,1699,1725]" pageId="19" pageNumber="20" value="2019-06-13">
<collectingDate id="EFEBE9AEFFB4FFB69D9BC8A19B4B4B8E" box="[1003,1133,1699,1725]" pageId="19" pageNumber="20" value="2019-06-13">2019.VI.13</collectingDate>
</date>
,
<collectorName id="26E45350FFB4FFB69A0BC8A19A2D4B8E" box="[1147,1291,1699,1725]" pageId="19" pageNumber="20">Xingyue Liu</collectorName>
(
<collectionCode id="ED00AE43FFB4FFB69B6CC8A19A7B4B8E" box="[1308,1373,1699,1725]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69B02C8A19CD24BD2" collectingDate="2016-03-26" collectionCode="CAU" collectorName="Xingyue Liu" country="Laos" elevation="250" location="Hatdai" municipality="Xaignabouri" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D17FD0FFFB4FFB69B02C8A19AB34B8E" box="[1394,1429,1699,1725]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69EE7C8C59FF64BD2" box="[151,208,1735,1761]" name="Laos" pageId="19" pageNumber="20">Laos</collectingCountry>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69E90C8C59E494BD2" box="[224,367,1735,1761]" pageId="19" pageNumber="20">Xaignabouri</collectingMunicipality>
, near
<location id="8ECE605DFFB4FFB69FCAC8C59D2E4BD2" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69FCAC8C59D2E4BD2" box="[442,520,1735,1761]" country="Laos" municipality="Xaignabouri" name="Hatdai" pageId="19" pageNumber="20">Hatdai</location>
,
<quantity id="4CE99B63FFB4FFB69C67C8C49D424BD2" box="[535,612,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="m" value="250.0">
<elevation id="003CD1B5FFB4FFB69C67C8C49D424BD2" box="[535,612,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="19" pageNumber="20" unit="m" value="250.0">250 m</elevation>
</quantity>
,
<date id="FFAF1046FFB4FFB69C05C8C59DD34BD2" box="[629,757,1735,1761]" pageId="19" pageNumber="20" value="2016-03-26">
<collectingDate id="EFEBE9AEFFB4FFB69C05C8C59DD34BD2" box="[629,757,1735,1761]" pageId="19" pageNumber="20" value="2016-03-26">2016.III.26</collectingDate>
</date>
,
<collectorName id="26E45350FFB4FFB69D75C8C59CBE4BD2" box="[773,920,1735,1761]" pageId="19" pageNumber="20">Xingyue Liu</collectorName>
(
<collectionCode id="ED00AE43FFB4FFB69DDBC8C59CCA4BD2" box="[939,1004,1735,1761]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69A74C8C59DF54A36" collectingDate="2012-07-02" collectionCode="CAU" collectorName="Xingyue Liu" country="Japan" location="Arrogawa Kon" municipality="Iriomotejima" pageId="19" pageNumber="20" specimenCount="2" specimenCount-male="2" stateProvince="Okinawa">
<specimenCount id="9D17FD0FFFB4FFB69A74C8C59B0C4BD2" box="[1028,1066,1735,1761]" count="2" pageId="19" pageNumber="20" type="male">2♂</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69A49C8C59B5A4BD2" box="[1081,1148,1735,1761]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69AFCC8C59BD24BD2" box="[1164,1268,1735,1761]" country="Japan" name="Okinawa" pageId="19" pageNumber="20">Okinawa</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69B73C8C59ABE4BD2" box="[1283,1432,1735,1761]" pageId="19" pageNumber="20">Iriomotejima</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69EE7C8E99E654A36" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69EE7C8E99E654A36" box="[151,323,1771,1797]" country="Japan" municipality="Iriomotejima" name="Arrogawa Kon" pageId="19" pageNumber="20" stateProvince="Okinawa">Arrogawa Kon</location>
,
<date id="FFAF1046FFB4FFB69F20C8E99EFA4A36" box="[336,476,1771,1797]" pageId="19" pageNumber="20" value="2012-07-02">
<collectingDate id="EFEBE9AEFFB4FFB69F20C8E99EFA4A36" box="[336,476,1771,1797]" pageId="19" pageNumber="20" value="2012-07-02">2012.VII.02</collectingDate>
</date>
,
<collectorName id="26E45350FFB4FFB69F99C8E99D5F4A36" box="[489,633,1771,1797]" pageId="19" pageNumber="20">Xingyue Liu</collectorName>
(
<collectionCode id="ED00AE43FFB4FFB69CFAC8E99DED4A36" box="[650,715,1771,1797]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B793CDBFFB4FFB69C90C8E99DD54A1A" collectingDate="2018-07-17" collectingDateMax="2018-07-22" collectingDateMin="2018-07-17" collectionCode="CAU" collectorName="Yunlan Jiang" country="Malaysia" elevation="1230" location="Trus Madi Entomology Camp" municipality="Kota Kinabalu" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Sabah">
<specimenCount id="9D17FD0FFFB4FFB69C90C8E99C254A36" box="[736,771,1771,1797]" count="1" pageId="19" pageNumber="20" type="female">1♀</specimenCount>
,
<collectingCountry id="F3067616FFB4FFB69D63C8E99C5A4A36" box="[787,892,1771,1797]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
,
<collectingRegion id="49D5F864FFB4FFB69DF9C8E99CF64A36" box="[905,976,1771,1797]" country="Malaysia" name="Sabah" pageId="19" pageNumber="20">Sabah</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFB4FFB69DAEC8E99BA14A36" box="[990,1159,1771,1797]" pageId="19" pageNumber="20">Kota Kinabalu</collectingMunicipality>
,
<location id="8ECE605DFFB4FFB69AE3C8E99FFB4A1A" LSID="urn:lsid:plazi:treatment:03B88790FFB5FFB69EE7C92F9E694A8A:8ECE605DFFB4FFB69AE3C8E99FFB4A1A" country="Malaysia" municipality="Kota Kinabalu" name="Trus Madi Entomology Camp" pageId="19" pageNumber="20" stateProvince="Sabah">Trus Madi Entomology Camp</location>
,
<quantity id="4CE99B63FFB4FFB69E98C90D9E194A1A" box="[232,319,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="19" pageNumber="20" unit="m" value="1230.0">
<elevation id="003CD1B5FFB4FFB69E98C90D9E194A1A" box="[232,319,1807,1833]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="19" pageNumber="20" unit="m" value="1230.0">1230 m</elevation>
</quantity>
,
<date id="FFAF1046FFB4FFB69F3BC90D9D264A1A" box="[331,512,1807,1833]" pageId="19" pageNumber="20" value="2018-07-17" valueMax="2018-07-22" valueMin="2018-07-17">
<collectingDate id="EFEBE9AEFFB4FFB69F3BC90D9D264A1A" box="[331,512,1807,1833]" pageId="19" pageNumber="20" value="2018-07-17" valueMax="2018-07-22" valueMin="2018-07-17">2018.VII.1722</collectingDate>
</date>
,
<collectorName id="26E45350FFB4FFB69C7BC90D9DBA4A1A" box="[523,668,1807,1833]" pageId="19" pageNumber="20">Yunlan Jiang</collectorName>
(
<collectionCode id="ED00AE43FFB4FFB69CDBC90D9DCE4A1A" box="[683,744,1807,1833]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="19" pageNumber="20">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB4FFB69EB7C9309D684A42" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BAE3686FFB4FFB69EB7C9309D684A42" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">
<emphasis id="B965EA94FFB4FFB69EB7C9309E474A7F" bold="true" box="[199,353,1842,1868]" pageId="19" pageNumber="20">Distribution.</emphasis>
<collectingCountry id="F3067616FFB4FFB69F1CC9319E894A7E" box="[364,431,1843,1869]" name="China" pageId="19" pageNumber="20">China</collectingCountry>
(
<collectingRegion id="49D5F864FFB4FFB69FB2C9319D284A7E" box="[450,526,1843,1869]" country="China" name="Fujian" pageId="19" pageNumber="20">Fujian</collectingRegion>
,
<collectingRegion id="49D5F864FFB4FFB69C6EC9319D824A7E" box="[542,676,1843,1869]" country="China" name="Guangdong" pageId="19" pageNumber="20">Guangdong</collectingRegion>
,
<collectingRegion id="49D5F864FFB4FFB69CC3C9319C304A7E" box="[691,790,1843,1869]" country="China" name="Guangxi" pageId="19" pageNumber="20">Guangxi</collectingRegion>
,
<collectingRegion id="49D5F864FFB4FFB69D56C9319CAF4A7E" box="[806,905,1843,1869]" country="China" name="Guizhou" pageId="19" pageNumber="20">Guizhou</collectingRegion>
,
<collectingRegion id="49D5F864FFB4FFB69DE9C9319CCB4A7E" box="[921,1005,1843,1869]" country="China" name="Hainan" pageId="19" pageNumber="20">Hainan</collectingRegion>
,
<collectingCountry id="F3067616FFB4FFB69D8CC9319B774A7E" box="[1020,1105,1843,1869]" name="Taiwan" pageId="19" pageNumber="20">Taiwan</collectingCountry>
, Yunnan);
<collectingCountry id="F3067616FFB4FFB69AA1C9319A374A7E" box="[1233,1297,1843,1869]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
(
<collectingRegion id="49D5F864FFB4FFB69B55C9319AB74A7E" box="[1317,1425,1843,1869]" country="Japan" name="Okinawa" pageId="19" pageNumber="20">Okinawa</collectingRegion>
);
<collectingCountry id="F3067616FFB4FFB69EE7C9559FEB4A42" box="[151,205,1879,1905]" name="Laos" pageId="19" pageNumber="20">Laos</collectingCountry>
(Xaignabouri);
<collectingCountry id="F3067616FFB4FFB69FF2C9559ECF4A42" box="[386,489,1879,1905]" name="Malaysia" pageId="19" pageNumber="20">Malaysia</collectingCountry>
(
<collectingRegion id="49D5F864FFB4FFB69F88C9559D644A42" box="[504,578,1879,1905]" country="Malaysia" name="Sabah" pageId="19" pageNumber="20">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFB4FFB69EB7C9799E694A8A" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BAE3686FFB4FFB69EB7C9799E694A8A" blockId="19.[151,1437,151,1977]" pageId="19" pageNumber="20">
<emphasis id="B965EA94FFB4FFB69EB7C9799E1D4AA6" bold="true" box="[199,315,1915,1941]" pageId="19" pageNumber="20">Remarks.</emphasis>
This species can be distinguished easily by the simple markings on body and the arched inner gradate crossvein series.
</paragraph>
</subSubSection>
</treatment>
</document>