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<document id="054D843DEDBCF0675D1402D8D7E8FFA0" ID-CLB-Dataset="80832" ID-DOI="10.5281/zenodo.6707142" ID-GBIF-Dataset="ab66b2b7-9544-4411-bf61-5bc3651d7bca" ID-ISBN="978-84-16728-04-6" ID-Zenodo-Dep="6707142" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1656002532852" checkinUser="carolina" docAuthor="Don E. Wilson, Russell A. Mittermeier &amp; Thomas E. Lacher, Jr" docDate="2017" docId="03F06D13FF2720EE0DA31EFF009CFAB4" docLanguage="en" docName="hbmw_7_Cricetidae_0204.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 7 Rodents II, Barcelona: Lynx Edicions" docTitle="Nephelomys caracolus" docType="treatment" docVersion="14" lastPageNumber="416" masterDocId="FFC9156BFFAE20670D37145C0837FFDB" masterDocTitle="Cricetidae" masterLastPageNumber="535" masterPageNumber="204" pageNumber="416" updateTime="1718799336328" updateUser="carolina">
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<mods:title id="A8EF7BB64F39AFD1E2272124922FA603">Cricetidae</mods:title>
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<mods:namePart id="F1DC1D7213E64C8B068890590A515D9B">Don E. Wilson</mods:namePart>
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<mods:namePart id="6E84E1FAB2BD91405A3B25C0F04EAA92">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="6812A33609920320445103843599587E">Thomas E. Lacher, Jr</mods:namePart>
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<mods:title id="B40FFB90D47C74C91E7710DA474EA88C">Handbook of the Mammals of the World Volume 7 Rodents II</mods:title>
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<treatment id="03F06D13FF2720EE0DA31EFF009CFAB4" ID-DOI="http://doi.org/10.5281/zenodo.6726800" ID-GBIF-Taxon="196221059" ID-Zenodo-Dep="6726800" LSID="urn:lsid:plazi:treatment:03F06D13FF2720EE0DA31EFF009CFAB4" httpUri="http://treatment.plazi.org/id/03F06D13FF2720EE0DA31EFF009CFAB4" lastPageNumber="416" pageId="137" pageNumber="416">
<subSubSection id="C3438F8EFF2720EE0DA31EFF08D0F50E" box="[148,231,2723,2773]" pageId="137" pageNumber="416" type="multiple">
<paragraph id="8BE6DC05FF2720EE0DA31EFF08D0F50E" blockId="137.[146,1293,2723,2853]" box="[148,231,2723,2773]" pageId="137" pageNumber="416">
<heading id="D0AE6B69FF2720EE0DA31EFF08D0F50E" box="[148,231,2723,2773]" pageId="137" pageNumber="416">
<figureCitation id="1362C080FF2720EE0DA31EFF08D0F50E" box="[148,231,2723,2773]" captionStart="Plate 20: Cricetidae" captionStartId="141.[156,186,3186,3207]" captionTargetBox="[12,2771,14,3647]" captionTargetPageId="130" captionText="353. Bogota Grass Mouse (Neomicroxus bogotensts), 354. Ecuadorean Grass Mouse (Neomucroxus latebricola), 355. Altiplano Chinchilla Mouse (Chinchillula sahamae), 356. Small Red-nosed Tree Mouse (Juliomys pictipes), 357. Delicate Red-nosed Tree Mouse (Juliomys ossitenuis), 358. Montane Red-rumped Tree Mouse (Juliomys rimofrons), 359. Araucaria Forest Tree Mouse (Juliomys ximenezi), 360. Ruschis Spiny Mouse (Abrawayaomys ruschii), 361. Chebezs Spiny Mouse (Abrawayaomys chebezi), 362. Montane Atlantic Forest Rat (Delomys altimontanus), 363. Pallid Atlantic Forest Rat (Delomys sublineatus), 364. Striped Atlantic Forest Rat (Delomys dorsalis), 365. North-western Conyrat (Reithrodon caurinus), 366. Naked-soled Conyrat (Reithrodon typicus), 367. Hairy-soled Conyrat (Reithrodon awritus), 368. Short-tailed Cane Mouse (Zygodontomys brevicauda), 369. Colombian Cane Mouse (Zygodontomys brunneus), 370. Ucayali South American Spiny Mouse (Scolomys ucayalensts), 371. Gray South American Spiny Mouse (Scolomys melanops), 372. Boquete Rice Rat (Nephelomys devius), 373. Santa Marta Rice Rat (Nephelomys maculiventer), 374. Mount Pirre Rice Rat (Nephelomys pirrensis), 375. Western Colombian Rice Rat (Nephelomys pectoralis), 376. Coastal Cordilleran Rice Rat (Nephelomys caracolus), 377. Merida Rice Rat (Nephelomys meridensis), 378. Childs Rice Rat (Nephelomys childi), 379. White-throated Rice Rat (Nephelomys albigularis), 380. Greater Golden-bellied Rice Rat (Nephelomys aurirventer), 381. Gray-bellied Rice Rat (Nephelomys moerex), 382. Lesser Golden-bellied Rice Rat (Nephelomys nimbosus), 383. Keayss Rice Rat (Nephelomys keaysi), 384. Nimble-footed Rice Rat (Nephelomys levipes), 385. Hammond's Rice Rat (Mindomys hammondi), 386. White-bellied Arboreal Rice Rat (Oecomys bicolor), 387. Yellow Arboreal Rice Rat (Oecomys flavicans), 388. Savanna Arboreal Rice Rat (Oecomys speciosus), 389. Long-furred Arboreal Rice Rat (Oecomys trinitatis), 390. Robert's Arboreal Rice Rat (Oecomys robert), 391. Unicolored Arboreal Rice Rat (Oecomys concolor), 392. Guianan Arboreal Rice Rat (Oecomys awyantepui), 393. King Arboreal Rice Rat (Oecomys rex), 394. Red Arboreal Rice Rat (Oecomys rutilus), 395. Brazilian Arboreal Rice Rat (Oecomys paricola), 396. Foothill Arboreal Rice Rat (Oecomys superans), 397. Dusky Arboreal Rice Rat (Oecomys phaeotis), 398. Mamore Arboreal Rice Rat (Oecomys mamorae), 399. Andersons Arboreal Rice Rat (Oecomys sydandersoni), 400. Franciscos Arboreal Rice Rat (Oecomys franciscorum), 401. Atlantic Forest Arboreal Rice Rat (Oecomys catherinae), 402. Clebers Arboreal Rice Rat (Oecomys cleber), 403. McConnells Rice Rat (Euryoryzomys macconnelli), 404. Emmonss Rice Rat (Euryoryzomys emmonsae), 405. Big-headed Rice Rat (Euryoryzomys legatus), 406. Russet Rice Rat (Euryoryzomys russatus), 407. Bufty-sided Rice Rat (Euryoryzomys lamia), 408. Elegant Rice Rat (Euryoryzomys nitidus), 409. Long-whiskered Rice Rat (Transandinomys bolivaris), 410. Talamancan Rice Rat (Transandinomys talamancae), 411. Long-nosed Rice Rat “ Handleyomys (™ rostratus), 412. Chapmans Rice Rat “ Handleyomys (” chapman), 413. Black-eared Rice Rat “ Handleyomys (™ melanotis), 414. Guerrero Rice Rat “ Handleyomys (™ guerrerensis), 415. Cloud Forest Rice Rat “ Handleyomys (” saturatior), 416. Highland Rice Rat “ Handleyomys (” rhabdops), 417. Alfaros Rice Rat “ Handleyomys (” alfaroi), 418. Colombian Western Andes Cloud Forest Mouse (Handleyomys fuscatus), 419. Colombian Central Andes Cloud Forest Mouse (Handleyomys intectus)" figureDoi="http://doi.org/10.5281/zenodo.6708503" httpUri="https://zenodo.org/record/6708503/files/figure.png" pageId="137" pageNumber="416">376.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE0DCF1EFF0B7FF50E" box="[248,840,2723,2773]" pageId="137" pageNumber="416" type="vernacular_names">
<paragraph id="8BE6DC05FF2720EE0DCF1EFF0B7FF50E" blockId="137.[146,1293,2723,2853]" box="[248,840,2723,2773]" pageId="137" pageNumber="416">
<heading id="D0AE6B69FF2720EE0DCF1EFF0B7FF50E" box="[248,840,2723,2773]" pageId="137" pageNumber="416">
<vernacularName id="055AAC2BFF2720EE0DCF1EFF0B7FF50E" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[248,840,2723,2773]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Coastal Cordilleran Rice Rat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE0EBA1EFF0D3AF50E" box="[909,1293,2723,2773]" pageId="137" pageNumber="416" type="nomenclature">
<paragraph id="8BE6DC05FF2720EE0EBA1EFF0D3AF50E" blockId="137.[146,1293,2723,2853]" box="[909,1293,2723,2773]" pageId="137" pageNumber="416">
<heading id="D0AE6B69FF2720EE0EBA1EFF0D3AF50E" box="[909,1293,2723,2773]" pageId="137" pageNumber="416">
<taxonomicName id="4C59A786FF2720EE0EBA1EFF0D3AF50E" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[909,1293,2723,2773]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">
<emphasis id="B92D0017FF2720EE0EBA1EFF0D3AF50E" box="[909,1293,2723,2773]" italics="true" pageId="137" pageNumber="416">Nephelomys caracolus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE0DA31EBB0C4BF4F8" pageId="137" pageNumber="416" type="vernacular_names">
<paragraph id="8BE6DC05FF2720EE0DA31EBB0CE3F527" blockId="137.[146,1293,2723,2853]" box="[148,1236,2791,2812]" pageId="137" pageNumber="416">
<heading id="D0AE6B69FF2720EE0DA31EBB0CE3F527" box="[148,1236,2791,2812]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE0DA31EBB08E8F527" bold="true" box="[148,223,2791,2812]" pageId="137" pageNumber="416">French:</emphasis>
<vernacularName id="055AAC2BFF2720EE0DDE1EBB09EDF527" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[233,474,2791,2812]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="fra" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Néphélomys de Caracas</vernacularName>
/
<emphasis id="B92D0017FF2720EE0CD81EBB0A7EF527" bold="true" box="[495,585,2791,2812]" pageId="137" pageNumber="416">German:</emphasis>
<vernacularName id="055AAC2BFF2720EE0F631EBB0B57F527" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[596,864,2791,2812]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="deu" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Kiistenkordilleren-Reisratte</vernacularName>
/
<emphasis id="B92D0017FF2720EE0E421EBB0BE7F527" bold="true" box="[885,976,2791,2812]" pageId="137" pageNumber="416">Spanish:</emphasis>
<vernacularName id="055AAC2BFF2720EE0EED1EBB0CE3F527" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[986,1236,2791,2812]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="esp" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Rata arrocera de la Costa</vernacularName>
</heading>
</paragraph>
<paragraph id="8BE6DC05FF2720EE0DA41F520C4BF4F8" blockId="137.[146,1293,2723,2853]" box="[147,1148,2830,2851]" pageId="137" pageNumber="416">
<heading id="D0AE6B69FF2720EE0DA41F520C4BF4F8" box="[147,1148,2830,2851]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE0DA41F5209BDF4F8" bold="true" box="[147,394,2830,2851]" pageId="137" pageNumber="416">Other common names:</emphasis>
<vernacularName id="055AAC2BFF2720EE0CA41F520A0EF4F8" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[403,569,2830,2851]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Caracol Rice Rat</vernacularName>
,
<vernacularName id="055AAC2BFF2720EE0F711F520B4DF4F8" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[582,890,2830,2851]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Coastal Cordillera Nephelomys</vernacularName>
,
<vernacularName id="055AAC2BFF2720EE0EBF1F520C4BF4F8" ID-CoL="73WZ9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[904,1148,2830,2851]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" language="eng" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Costa Central Oryzomys</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE0FC01F080D18F41A" pageId="137" pageNumber="416" type="reference_group">
<paragraph id="8BE6DC05FF2720EE0FC01F080D18F41A" blockId="137.[759,1353,2900,3323]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE0FC01F080BA5F4AA" bold="true" box="[759,914,2900,2929]" pageId="137" pageNumber="416">Taxonomy.</emphasis>
<taxonomicName id="4C59A786FF2720EE0E861F080B0BF442" ID-CoL="74ZH5" authority="Thomas" authorityName="Thomas" authorityYear="1914" class="Mammalia" family="Cricetidae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Oryzomys caracolus Thomas, 1914</taxonomicName>
,
<materialsCitation id="3B31D658FF2720EE0E791FDC0D18F41A" ID-GBIF-Occurrence="3818814881" pageId="137" pageNumber="416">
“Galiparé, Cerro del Avila, near Caracas,
<collectingCountry id="F34E9C95FF2720EE0E7A1FF40BD7F41A" box="[845,992,2984,3009]" name="Venezuela" pageId="137" pageNumber="416">Venezuela</collectingCountry>
. Alt. 6000 [=
<quantity id="4CA171E0FF2720EE099E1FF40D24F41A" box="[1193,1299,2984,3009]" metricMagnitude="3" metricUnit="m" metricValue="1.829" pageId="137" pageNumber="416" unit="m" value="1829.0">1829 m</quantity>
].”
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</paragraph>
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<subSubSection id="C3438F8EFF2720EE0FCF1F960FF3FEB4" pageId="137" pageNumber="416" type="discussion">
<paragraph id="8BE6DC05FF2720EE0FCF1F960FF3FEB4" blockId="137.[759,1353,2900,3323]" lastBlockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<taxonomicName id="4C59A786FF2720EE0FCF1F960C2DF43C" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[760,1050,3018,3047]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">Nephelomys caracolus</taxonomicName>
was originally described in 1914 as a species of
<taxonomicName id="4C59A786FF2720EE09FC1FB30D76F3CB" box="[1227,1345,3055,3088]" class="Mammalia" family="Cricetidae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="genus">Oryzomys</taxonomicName>
; from 1960 on, it was included in
<taxonomicName id="4C59A786FF2720EE09F818490D77F3ED" baseAuthorityName="Olfers" baseAuthorityYear="1818" box="[1231,1344,3093,3126]" class="Mammalia" family="Cricetidae" genus="Oryzomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="capito">O. capito</taxonomicName>
,
<taxonomicName id="4C59A786FF2720EE0FCE18600C0CF386" baseAuthorityName="Fischer" baseAuthorityYear="1814" box="[761,1083,3132,3165]" class="Mammalia" family="Cricetidae" genus="Hylaeamys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="megacephalus">Hylaeamys megacephalus</taxonomicName>
, and subsequently
<taxonomicName id="4C59A786FF2720EE0FCC18380B9EF35E" baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[763,937,3172,3205]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="albigularis">O. albigularis</taxonomicName>
. In the 1990s, studies of Venezuelan populations in the
<taxonomicName id="4C59A786FF2720EE09A118D70D7EF377" baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[1174,1353,3211,3244]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="417" phylum="Chordata" rank="species" species="albigularis">O. albigularis</taxonomicName>
group, based on karyotypes and morphometrics, provided evidence for distinctiveness of
<taxonomicName id="4C59A786FF2720EE0C37195D09ABF2F9" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[256,412,3329,3362]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">O. caracolus</taxonomicName>
and the other Venezuelan species of the group (e.g.
<taxonomicName id="4C59A786FF2720EE09BD195D0D0EF2F9" baseAuthorityName="Thomas" baseAuthorityYear="1894" box="[1162,1337,3329,3362]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="meridensis">O. meridensis</taxonomicName>
). Patterns differed among studies, and some authors chose to retain
<taxonomicName id="4C59A786FF2720EE095F19740D3FF292" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[1128,1288,3368,3401]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">O. caracolus</taxonomicName>
and
<taxonomicName id="4C59A786FF2720EE0DAF190C0974F2AA" baseAuthorityName="Thomas" baseAuthorityYear="1894" box="[152,323,3408,3441]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="meridensis">O. meridensis</taxonomicName>
as subspecies of
<taxonomicName id="4C59A786FF2720EE0F0C190C0ADCF2AA" baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[571,747,3408,3441]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="albigularis">O. albigularis</taxonomicName>
, suggesting need for further study of classification of Venezuelan members of the group. When members of the
<taxonomicName id="4C59A786FF2720EE09AD192B0D7EF243" baseAuthorityName="Tomes" baseAuthorityYear="1860" box="[1178,1353,3447,3480]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="albigularis">O. albigularis</taxonomicName>
group were transferred to the new genus
<taxonomicName id="4C59A786FF2720EE0AE1157A0050FE9C" box="[2006,2151,294,327]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="genus">Nephelomys</taxonomicName>
in 2006,
<taxonomicName id="4C59A786FF2720EE05DC157A0168FE9C" baseAuthorityName="Thomas" baseAuthorityYear="1914" box="[2283,2399,294,327]" class="Mammalia" family="Cricetidae" genus="Nephelomys" kingdom="Animalia" order="Rodentia" pageId="137" pageNumber="416" phylum="Chordata" rank="species" species="caracolus">caracolus</taxonomicName>
was moved and retained as a distinct species. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BE15290008FE4D" box="[1417,2111,373,406]" pageId="137" pageNumber="416" type="distribution">
<caption id="DF268C8DFF2720EE08BE15290008FE4D" ID-DOI="http://doi.org/10.5281/zenodo.6761260" ID-Zenodo-Dep="6761260" box="[1417,2111,373,406]" httpUri="https://zenodo.org/record/6761260/files/figure.png" inLine="true" pageId="137" pageNumber="416" startId="137.[1417,1589,373,406]" targetBox="[145,735,2908,3322]" targetPageId="137">
<paragraph id="8BE6DC05FF2720EE08BE15290008FE4D" blockId="137.[1416,2624,294,1394]" box="[1417,2111,373,406]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BE15290E0EFE4D" bold="true" box="[1417,1593,373,406]" pageId="137" pageNumber="416">Distribution.</emphasis>
<taxonomicName id="4C59A786FF2720EE0B7215290EC8FE4D" authority="de" box="[1605,1791,373,406]" class="Magnoliopsida" family="Chrysobalanaceae" genus="Cordillera" kingdom="Plantae" order="Malpighiales" pageId="137" pageNumber="497" phylum="Tracheophyta" rank="genus">Cordillera de</taxonomicName>
la Costa, N Venezuela.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BE15F80EA4FD71" pageId="137" pageNumber="416" type="description">
<paragraph id="8BE6DC05FF2720EE08BE15F80EA4FD71" blockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BE15F80EB1FE66" bold="true" box="[1417,1670,420,445]" pageId="137" pageNumber="416">Descriptive notes.</emphasis>
Head-body 141 mm, tail 162 mm, ear 24 mm, hindfoot 34 mm; weight 66 g (mean values for a sample from Henri Pittier National Park). The Coastal Cordilleran Rice Rat is medium-sized, with males slightly larger than females. Dorsal pelage is long, dense, and yellowish, intensely grizzled with brown; venter is grayish to whitish, with narrow white pectoral patches variably present (in some specimens, patches are especially long, extending to gular region). Chromosomal complement is 2n = 66, FN = 90.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BD16F30003FC9C" pageId="137" pageNumber="416" type="biology_ecology">
<paragraph id="8BE6DC05FF2720EE08BD16F30003FC9C" blockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BD16F30DCEFD0B" bold="true" box="[1418,1529,687,720]" pageId="137" pageNumber="416">Habitat.</emphasis>
Mature evergreen forest with rich subcanopy of ferns and vines and covered with abundant moss and epiphytes, disturbed forests, and drier evergreen and cloud forests with limited epiphytic cover, mostly associated with Caribbean slope of the
<taxonomicName id="4C59A786FF2720EE073216A20E38FC9C" authority="de" class="Magnoliopsida" family="Chrysobalanaceae" genus="Cordillera" kingdom="Plantae" order="Malpighiales" pageId="137" pageNumber="497" phylum="Tracheophyta" rank="genus">Cordillera de</taxonomicName>
la Costa, at elevations of 1050-2300 m.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BD171101D3FC66" pageId="137" pageNumber="416" type="food_feeding">
<paragraph id="8BE6DC05FF2720EE08BD171101D3FC66" blockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BD17110EAFFCB5" bold="true" box="[1418,1688,845,878]" pageId="137" pageNumber="416">Food and Feeding.</emphasis>
The Coastal Cordilleran Rice Rat eats fruits, seeds, and invertebrates. Stomach contents revealed ¢.50% fruits (
<taxonomicName id="4C59A786FF2720EE050F17200092FC4E" box="[2104,2213,892,917]" class="Magnoliopsida" family="Fabaceae" genus="Inga" kingdom="Plantae" order="Fabales" pageId="137" pageNumber="416" phylum="Tracheophyta" rank="species" species="undetermined">Inga sp.</taxonomicName>
,
<taxonomicName id="4C59A786FF2720EE058017200109FC4E" box="[2231,2366,892,917]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="137" pageNumber="416" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), c.20% insect larvae, ¢.20% palm seeds, c¢.2% fungi, and c.2%
<taxonomicName id="4C59A786FF2720EE053D17F8015BFC66" authorityName="Godart" authorityYear="1819" box="[2058,2412,932,957]" class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Plantae" order="Zingiberales" pageId="137" pageNumber="416" phylum="Tracheophyta" rank="genus">Heliconia (Heliconiaceae)</taxonomicName>
flowers.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BC179F0E91FB80" pageId="137" pageNumber="416" type="breeding">
<paragraph id="8BE6DC05FF2720EE08BC179F0E91FB80" blockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BC179F0E26FC3F" bold="true" box="[1419,1553,963,996]" pageId="137" pageNumber="416">Breeding.</emphasis>
Coastal Cordilleran Rice Rats breed year-round, with peak in wet season. Sexual maturity is reached at 47 days old for males and 52 days old for females. Litters have 2-5 young. Laboratory results strongly suggest that the Coastal Cordilleran Rice Rat is monogamous.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BE103D01EFFB59" box="[1417,2520,1121,1154]" pageId="137" pageNumber="416" type="activity">
<paragraph id="8BE6DC05FF2720EE08BE103D01EFFB59" blockId="137.[1416,2624,294,1394]" box="[1417,2520,1121,1154]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BE103D0E43FB59" bold="true" box="[1417,1652,1121,1154]" pageId="137" pageNumber="416">Activity patterns.</emphasis>
The Coastal Cordilleran Rice Rat is nocturnal and terrestrial.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BC10D00100FB72" box="[1419,2359,1164,1193]" pageId="137" pageNumber="416" type="biology_ecology">
<paragraph id="8BE6DC05FF2720EE08BC10D00100FB72" blockId="137.[1416,2624,294,1394]" box="[1419,2359,1164,1193]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BC10D0007FFB72" bold="true" box="[1419,2120,1164,1193]" pageId="137" pageNumber="416">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BC10E801F5FB0A" box="[1419,2498,1204,1233]" pageId="137" pageNumber="416" type="conservation">
<paragraph id="8BE6DC05FF2720EE08BC10E801F5FB0A" blockId="137.[1416,2624,294,1394]" box="[1419,2498,1204,1233]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BC10E80EDFFB0A" bold="true" box="[1419,1768,1204,1233]" pageId="137" pageNumber="416">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List.
</paragraph>
</subSubSection>
<subSubSection id="C3438F8EFF2720EE08BB10BC009CFAB4" pageId="137" pageNumber="416" type="bibRefCitation_list">
<paragraph id="8BE6DC05FF2720EE08BB10BC009CFAB4" blockId="137.[1416,2624,294,1394]" pageId="137" pageNumber="416">
<emphasis id="B92D0017FF2720EE08BB10BC0E12FB22" bold="true" box="[1420,1573,1248,1273]" pageId="137" pageNumber="416">Bibliography.</emphasis>
Aguilera (2016¢), Aguilera et al. (1995), Anderson (2003), Anderson &amp; Raza (2010), Cardenas (2017), Diaz de Pascual (1988),
<taxonomicName id="4C59A786FF2720EE0BD0115B00BFFAFB" authority=", Delgado-Jaramillo et al. (2012)" authorityName="Delgado-Jaramillo" authorityYear="2012" box="[1767,2184,1287,1312]" class="Magnoliopsida" family="Euphorbiaceae" genus="Garcia" kingdom="Plantae" order="Malpighiales" pageId="137" pageNumber="416" phylum="Tracheophyta" rank="genus">Garcia, Delgado-Jaramillo et al. (2012)</taxonomicName>
, Handley (1976), Marquez et al. (2000), Moscarella (1997), Moscarella &amp; Aguilera (1999), Moscarella et al. (2001), Musser &amp; Carleton (2005), Percequillo (2015g), Rivas (1993, 1997), Rivas &amp; Péfaur (1999), Thomas (1914a).
</paragraph>
</subSubSection>
</treatment>
</document>