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<document id="870846263830D64CD5D8BF9826C42111" ID-DOI="10.11646/zootaxa.5360.2.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="10145774" ID-ZooBank="5A6202BB-121B-45E4-8BA7-FFA20B6E9ABC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700174684381" checkinUser="plazi" docAuthor="Mofrad, Farhad Eshraghi" docDate="2023" docId="03E967369376FFEDFF346AFEFAA4779C" docLanguage="en" docName="zootaxa.5360.2.8.pdf" docOrigin="Zootaxa 5360 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5360.2.8/52119" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euboeus (Pelorinus) kandovanensis Mofrad 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="299" masterDocId="FFD01F4E9377FFEEFFA36B24FFCD764D" masterDocTitle="A new species of the genus Euboeus Boieldieu, 1865 from Northern Iran (Coleoptera: Tenebrionidae: Helopini)" masterLastPageNumber="300" masterPageNumber="296" pageNumber="297" updateTime="1700175042480" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EBDD2C6619D87D8E7A88E447E2AB67DA">A new species of the genus Euboeus Boieldieu, 1865 from Northern Iran (Coleoptera: Tenebrionidae: Helopini)</mods:title>
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<treatment id="03E967369376FFEDFF346AFEFAA4779C" LSID="urn:lsid:plazi:treatment:03E967369376FFEDFF346AFEFAA4779C" httpUri="http://treatment.plazi.org/id/03E967369376FFEDFF346AFEFAA4779C" lastPageId="3" lastPageNumber="299" pageId="1" pageNumber="297">
<subSubSection id="C35A85AB9376FFEFFF346AFEFD5B77B8" box="[151,662,474,501]" pageId="1" pageNumber="297" type="nomenclature">
<paragraph id="8BFFD6209376FFEFFF346AFEFD5B77B8" blockId="1.[151,662,474,537]" box="[151,662,474,501]" pageId="1" pageNumber="297">
<heading id="D0B7614C9376FFEFFF346AFEFD5B77B8" bold="true" box="[151,662,474,501]" fontSize="11" level="1" pageId="1" pageNumber="297" reason="1">
<emphasis id="B9340A329376FFEFFF346AFEFD5B77B8" bold="true" box="[151,662,474,501]" pageId="1" pageNumber="297">
<taxonomicName id="4C40ADA39376FFEFFF346AFEFD8177B8" authority="Mofrad, 2023" authorityName="Mofrad" authorityYear="2023" box="[151,588,474,501]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="kandovanensis" status="sp. nov." subGenus="Pelorinus">
<emphasis id="B9340A329376FFEFFF346AFEFECC77B8" bold="true" box="[151,257,474,501]" italics="true" pageId="1" pageNumber="297">Euboeus</emphasis>
(
<emphasis id="B9340A329376FFEFFEB16AFEFE4A77B8" bold="true" box="[274,391,474,501]" italics="true" pageId="1" pageNumber="297">Pelorinus</emphasis>
)
<emphasis id="B9340A329376FFEFFE3A6AFEFD8177B8" bold="true" box="[409,588,474,501]" italics="true" pageId="1" pageNumber="297">kandovanensis</emphasis>
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<taxonomicNameLabel id="A207B7499376FFEFFDF06AFFFD5B77B8" box="[595,662,475,501]" pageId="1" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="C35A85AB9376FFEFFF346ADBFEF77454" box="[151,314,511,537]" pageId="1" pageNumber="297" type="description">
<paragraph id="8BFFD6209376FFEFFF346ADBFEF77454" blockId="1.[151,662,474,537]" box="[151,314,511,537]" pageId="1" pageNumber="297">
(
<figureCitation id="137BCAA59376FFEFFF3C6ADBFEFE7454" box="[159,307,511,537]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="1" pageNumber="297">Figs. 1A—K</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A85AB9376FFEFFF346963FAF774C8" pageId="1" pageNumber="297" type="materials_examined">
<paragraph id="8BFFD6209376FFEFFF346963FAF774C8" blockId="1.[151,1436,582,647]" pageId="1" pageNumber="297">
<emphasis id="B9340A329376FFEFFF346963FE84742D" bold="true" box="[151,329,582,609]" pageId="1" pageNumber="297">Type material.</emphasis>
<typeStatus id="54FB68829376FFEFFEF66963FE0D742C" box="[341,448,583,609]" pageId="1" pageNumber="297" type="holotype">Holotype</typeStatus>
, ♁ and
<materialsCitation id="3B28DC7D9376FFEFFD876963FAF774C8" collectingDate="2023-06-17" collectorName="F. Eshraghi Mofrad." country="Iran" elevation="2950" location="Gachsar" municipality="Central" pageId="1" pageNumber="297" specimenCount="1" specimenCount-female="1" stateProvince="Alborz" typeStatus="paratype">
<typeStatus id="54FB68829376FFEFFD876963FD44742C" box="[548,649,583,609]" pageId="1" pageNumber="297" type="paratype">Paratype</typeStatus>
,
<specimenCount id="9D461DA99376FFEFFD3B6963FD60742C" box="[664,685,583,609]" count="1" pageId="1" pageNumber="297" type="female"></specimenCount>
(HMIM):
<collectingCountry id="F35796B09376FFEFFC8D6963FC93742C" box="[814,862,583,609]" name="Iran" pageId="1" pageNumber="297">Iran</collectingCountry>
,
<collectingRegion id="498418C29376FFEFFCCF6963FBCE742C" box="[876,1027,583,609]" country="Iran" name="Alborz" pageId="1" pageNumber="297">Alborz Prov.</collectingRegion>
,
<collectingMunicipality id="6B9B4C5A9376FFEFFBB16963FBA9742C" box="[1042,1124,583,609]" pageId="1" pageNumber="297">Central</collectingMunicipality>
<collectingRegion id="498418C29376FFEFFBCD6963FB71742C" box="[1134,1212,583,609]" country="Iran" name="Alborz" pageId="1" pageNumber="297">Alborz</collectingRegion>
Mts.,
<location id="8E9F80FB9376FFEFFAAE6963FAA4742C" LSID="urn:lsid:plazi:treatment:03E967369376FFEDFF346AFEFAA4779C:8E9F80FB9376FFEFFAAE6963FAA4742C" box="[1293,1385,583,609]" country="Iran" municipality="Central" name="Gachsar" pageId="1" pageNumber="297" stateProvince="Alborz">Gachsar</location>
env (Kandovan tunnel)., H=
<quantity id="4CB87BC59376FFEFFE02694FFD8D74C8" box="[417,576,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="1" pageNumber="297" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">
<elevation id="006D31139376FFEFFE02694FFD8D74C8" box="[417,576,619,645]" metricMagnitude="3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="1" pageNumber="297" unit="m" value="2950.0" valueMax="3000.0" valueMin="2900.0">29003000 m</elevation>
</quantity>
, 36°09ʹ06.4ʹʹN, 51°18ʹ45.3ʹʹE,
<date id="FFFEF0E09376FFEFFC39694FFBD074C8" box="[922,1053,619,645]" pageId="1" pageNumber="297" value="2023-06-17">
<collectingDate id="EFBA09089376FFEFFC39694FFBD074C8" box="[922,1053,619,645]" pageId="1" pageNumber="297" value="2023-06-17">17.VI.2023</collectingDate>
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; leg.
<collectorName id="26B5B3F69376FFEFFBFA694FFAF774C8" box="[1113,1338,619,645]" pageId="1" pageNumber="297">F. Eshraghi Mofrad.</collectorName>
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</paragraph>
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<caption id="DF3F86A89376FFEFFF346C32FE9571D7" ID-DOI="http://doi.org/10.5281/zenodo.10145776" ID-Zenodo-Dep="10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="1" pageNumber="297" startId="1.[151,250,1814,1838]" targetBox="[350,1233,671,1775]" targetPageId="1" targetType="figure">
<paragraph id="8BFFD6209376FFEFFF346C32FE9571D7" blockId="1.[151,1437,1814,1946]" pageId="1" pageNumber="297">
<emphasis id="B9340A329376FFEFFF346C32FED97163" bold="true" box="[151,276,1814,1838]" pageId="1" pageNumber="297">FIGURE 1.</emphasis>
<taxonomicName id="4C40ADA39376FFEFFEBF6C32FDC27163" authority="Mofrad, 2023" authorityName="Mofrad" authorityYear="2023" box="[284,527,1814,1838]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="297" phylum="Arthropoda" rank="species" species="kandovanensis" status="sp. nov.">
<emphasis id="B9340A329376FFEFFEBF6C32FDC27163" box="[284,527,1814,1838]" italics="true" pageId="1" pageNumber="297">Euboeus kandovanensis</emphasis>
</taxonomicName>
<emphasis id="B9340A329376FFEFFDB46C32FDA47163" bold="true" box="[535,617,1814,1838]" pageId="1" pageNumber="297">
<taxonomicNameLabel id="A207B7499376FFEFFDB46C32FDA47163" box="[535,617,1814,1838]" pageId="1" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally.
</paragraph>
</caption>
<subSubSection id="C35A85AB9375FFECFF646BB2FB1F7038" pageId="2" pageNumber="298" type="description">
<paragraph id="8BFFD6209375FFECFF646BB2FEEB76B4" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
<emphasis id="B9340A329375FFECFF646BB2FE9476FD" bold="true" box="[199,345,150,176]" pageId="2" pageNumber="298">Description.</emphasis>
Male. Body slender, black, pronotum and elytra dorsally shiny, head lesser shiny (
<figureCitation id="137BCAA59375FFECFA926BB3FA4376FC" box="[1329,1422,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1A</figureCitation>
). Measurements: Y= 1.6;
<collectingCountry id="F35796B09375FFECFE016B9FFE0B7698" box="[418,454,187,213]" name="Philippines" pageId="2" pageNumber="298">PH</collectingCountry>
<subScript id="17C4D4659375FFECFE656BE9FE1F7696" attach="left" box="[454,466,205,219]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.66; P
<subScript id="17C4D4659375FFECFD956BE9FD8F7696" attach="both" box="[566,578,205,219]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
P
<subScript id="17C4D4659375FFECFDF26BE9FD9B7696" attach="left" box="[593,598,205,219]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
= 1.29; E
<subScript id="17C4D4659375FFECFD1F6BE9FD0C7696" attach="both" box="[700,705,205,219]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
E
<subScript id="17C4D4659375FFECFD726BE9FD107696" attach="left" box="[721,733,205,219]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.68; EH
<subScript id="17C4D4659375FFECFCF46BE9FCAE7696" attach="left" box="[855,867,205,219]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 2.08; EP
<subScript id="17C4D4659375FFECFC7B6BE9FC297696" attach="left" box="[984,996,205,219]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.25; EP
<subScript id="17C4D4659375FFECFBFA6BE9FB937696" attach="left" box="[1113,1118,205,219]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
= 2.7. Body length
<quantity id="4CB87BC59375FFECFA906B9FFA557698" box="[1331,1432,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="2" pageNumber="298" unit="mm" value="13.2">13.2 mm</quantity>
, width
<quantity id="4CB87BC59375FFECFF7D6BFAFEEF76B4" box="[222,290,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="298" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BFFD6209375FFECFF646A27FF28779C" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Head (
<figureCitation id="137BCAA59375FFECFEB06A27FE937750" box="[275,350,259,285]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1</figureCitation>
E-F). Anterior margin of epistoma slightly rounded. Head widest at eye level. Eyes moderately convex with circumocular impression narrow. Lateral margins of genae evenly rounded. Puncturation of head moderately fine and sparse, interpuncture distance 1.5 times as long as puncture diameter in middle; punctures coarser and denser on genae. Ventral side of epicranium coarsely and densely punctured. Antennae comparatively short, with only two apical antennomeres, extending beyond base of pronotum, reaching less than 1/4 of elytral length.
</paragraph>
<paragraph id="8BFFD6209375FFECFF646AFFFC1575CC" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Prothorax (
<figureCitation id="137BCAA59375FFECFEE76AFFFE6877B8" box="[324,421,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1B</figureCitation>
, E-G). Pronotum widest at middle, transverse, 1.29 times as wide as long (
<figureCitation id="137BCAA59375FFECFB406AFFFAF577B8" box="[1251,1336,475,501]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1E</figureCitation>
). Lateral margins of pronotum weakly evenly rounded, slightly emarginated near base; anterior margin weakly rounded at middle and slightly emarginated near corners; base weakly rounded, almost straight, with emargination at middle. Antero-lateral corners not projected, obtuse, narrowly rounded at apex; postero-lateral corners right, pointed at apex. Lateral margins slightly beaded in anterior half; wider beaded in the basal half (
<figureCitation id="137BCAA59375FFECFBE8694FFB5974C8" box="[1099,1172,619,645]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1</figureCitation>
E-F). Disc of pronotum weakly faintly convex, not flattened along lateral margin, with large impression on each lateral side and two very small impressions at base. Puncturation of disc fine and sparse; puncture diameter 35 times as short as interpuncture distance, circular, coarser and denser in impressions (
<figureCitation id="137BCAA59375FFECFCAD69F3FC9074BC" box="[782,861,727,753]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1</figureCitation>
E-F). Prosternum with scattered superficial hairy puncturation (
<figureCitation id="137BCAA59375FFECFE9769DFFE477558" box="[308,394,763,789]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1B</figureCitation>
). Prothoracic hypomera flattened along margin, slightly narrower at basal half. Prohypomera with coarse and dense separate (not merged) punctures among weak wrinkles in anterior two thirds (
<figureCitation id="137BCAA59375FFECFA91683BFA427574" box="[1330,1423,799,825]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1G</figureCitation>
). Prosternal process smooth and shiny, with two terminal lines of punctures in basal half, with setose punctures in the anterior half, slightly convex and with a sharp bulge at apex (
<figureCitation id="137BCAA59375FFECFCE56843FC0175CC" box="[838,972,871,897]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1B, G</figureCitation>
).
</paragraph>
<paragraph id="8BFFD6209375FFECFF6468AFFC237214" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Pterothorax (
<figureCitation id="137BCAA59375FFECFEFB68AFFE6D75E8" box="[344,416,907,933]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1</figureCitation>
A-B). Scutellar shield with straight margins, finely and sparsely punctured. Elytra elongateoval, widest at middle; punctures in striae coarse, widely spaced; punctures in interstriae tiny and sparse, slightly lined parallel (
<figureCitation id="137BCAA59375FFECFEE368F7FE5175A0" box="[320,412,979,1005]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1A</figureCitation>
). Anterior half of mesoventrite pubescent, with coarse and dense rasp-like puncturation; mesepisterna with large and dense punctures, punctures with short setae; mesepimera and metepisterna with coarse and dense puncturation, punctures with very short setae; metaventrite and intercoxal part of mesoventrite sparsely punctured and covered with short setae, punctures in medium size (
<figureCitation id="137BCAA59375FFECFC2A6F1BFC2D7214" box="[905,992,1087,1113]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1B</figureCitation>
).
</paragraph>
<paragraph id="8BFFD6209375FFECFF646F47FCD27340" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Legs (
<figureCitation id="137BCAA59375FFECFEAE6F47FE5D7230" box="[269,400,1123,1149]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1B, C</figureCitation>
). Trochanters with brush of short dark-reddish setae and one long seta. Basal third of femora with the same brush on reflex side; another surface of femora with coarse puncturation and short setae (
<figureCitation id="137BCAA59375FFECFA956FA3FA4072EC" box="[1334,1421,1159,1185]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1B</figureCitation>
). Pro- and mesotibiae straight, metatibiae bent (
<figureCitation id="137BCAA59375FFECFD306F8FFD247288" box="[659,745,1195,1221]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1C</figureCitation>
). Pro- and mesotarsomeres 14 strongly expanded, transverse, pro- and mesotarsomere 1 wider than apex of protibia. Metatarsi on the ventral side with dense brown to light brown setae, metatarsomere 1 with very dense light brown setae.
</paragraph>
<paragraph id="8BFFD6209375FFECFF646E33FE797318" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Abdomen (
<figureCitation id="137BCAA59375FFECFEE56E33FE50737C" box="[326,413,1303,1329]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1B</figureCitation>
). Abdominal ventrites with fine and sparse puncturation and short setation. Ventrite 5 with sparse long setae at apex.
</paragraph>
<paragraph id="8BFFD6209375FFECFF646E7BFEB5738C" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Genitalia (
<figureCitation id="137BCAA59375FFECFE9C6E7BFE457334" box="[319,392,1375,1401]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Figs. 1</figureCitation>
H-K). Apical part of the basal piece of aedeagus with longitudinal rugosity dorsally. Apical piece of aedeagus weakly rounded and emarginated before apex, apical piece dorsally with longitudinal depression in middle (
<figureCitation id="137BCAA59375FFECFEB26E83FEA7738C" box="[273,362,1447,1473]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1K</figureCitation>
).
</paragraph>
<paragraph id="8BFFD6209375FFECFF646EEFFB1F7038" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
Female (
<figureCitation id="137BCAA59375FFECFE886EEFFE4A73A8" box="[299,391,1483,1509]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1D</figureCitation>
). Body much more robust, dull dorsally. Antennae shorter, not reaching base of pronotum. Lateral margins of pronotum widely beaded only at basal half. Postero-lateral corners of pronotum right. Metatibiae strongly bent. Elytra with keels on third and fifth interstriae, keel on third interstriae larger. Such keels can be found in large females of
<emphasis id="B9340A329375FFECFED36D13FE10701C" box="[368,477,1591,1617]" italics="true" pageId="2" pageNumber="298">Pelorinus</emphasis>
(
<bibRefCitation id="EFD1ABD19375FFECFE4F6D13FD4D701C" author="Soldati, F." box="[492,640,1591,1617]" pageId="2" pageNumber="298" pagination="221 - 225" refId="ref3019" refString="Soldati, F. (2012) A new species of the genus Probaticus Seidlitz, 1896 from Greece (Insecta: Coleoptera: Tenebrionidae). Annales zoologici, 62 (2), 221 - 225. https: // doi. org / 10.3161 / 000345412 X 652747" type="journal article" year="2012">Soldati 2012</bibRefCitation>
), but are not diagnostic characters. Measurements: Y= 1.4;
<collectingCountry id="F35796B09375FFECFABE6D13FA8C701C" box="[1309,1345,1591,1617]" name="Philippines" pageId="2" pageNumber="298">PH</collectingCountry>
<subScript id="17C4D4659375FFECFAE26D6DFA80701A" attach="left" box="[1345,1357,1609,1623]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.64; P
<subScript id="17C4D4659375FFECFF046D49FF7E7036" attach="both" box="[167,179,1645,1659]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
P
<subScript id="17C4D4659375FFECFF616D49FF0A7036" attach="left" box="[194,199,1645,1659]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
= 1.31; E
<subScript id="17C4D4659375FFECFE8D6D49FEFE7036" attach="both" box="[302,307,1645,1659]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
E
<subScript id="17C4D4659375FFECFEE76D49FE9D7036" attach="left" box="[324,336,1645,1659]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.47; EH
<subScript id="17C4D4659375FFECFE6F6D49FE157036" attach="right" box="[460,472,1645,1659]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 2.32; EP
<subScript id="17C4D4659375FFECFDEC6D49FD967036" attach="left" box="[591,603,1645,1659]" fontSize="6" pageId="2" pageNumber="298">w</subScript>
= 1.41; EP
<subScript id="17C4D4659375FFECFD716D49FD1A7036" attach="left" box="[722,727,1645,1659]" fontSize="6" pageId="2" pageNumber="298">l</subScript>
= 2.74. Body length
<quantity id="4CB87BC59375FFECFC1E6D7FFBEE7038" box="[957,1059,1627,1653]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="298" unit="mm" value="14.8">14.8 mm</quantity>
, width
<quantity id="4CB87BC59375FFECFBD66D7EFB037038" box="[1141,1230,1626,1653]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="298" unit="mm" value="6.5">6.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A85AB9375FFECFF646D5BFBEA70D4" box="[199,1063,1663,1689]" pageId="2" pageNumber="298" type="etymology">
<paragraph id="8BFFD6209375FFECFF646D5BFBEA70D4" blockId="2.[151,1437,150,2013]" box="[199,1063,1663,1689]" pageId="2" pageNumber="298">
<emphasis id="B9340A329375FFECFF646D5BFE8370D4" bold="true" box="[199,334,1663,1689]" pageId="2" pageNumber="298">Etymology.</emphasis>
The specific epithet refers to the
<typeStatus id="54FB68829375FFECFD666D5BFD3870D4" box="[709,757,1663,1689]" pageId="2" pageNumber="298">type</typeStatus>
locality: Kandovan tunnel.
</paragraph>
</subSubSection>
<subSubSection id="C35A85AB9375FFEDFF646D86FAA4779C" lastPageId="3" lastPageNumber="299" pageId="2" pageNumber="298" type="diagnosis">
<paragraph id="8BFFD6209375FFECFF646D86FAC87148" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
<emphasis id="B9340A329375FFECFF646D86FE1E70F1" bold="true" box="[199,467,1698,1724]" pageId="2" pageNumber="298">Differential diagnosis.</emphasis>
This species is very similar to
<taxonomicName id="4C40ADA39375FFECFCF46D87FB6370F0" authority="(Kulzer, 1964)" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[855,1198,1699,1725]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFCF46D87FC3570F0" box="[855,1016,1699,1725]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
(Kulzer, 1964)
</taxonomicName>
and to
<taxonomicName id="4C40ADA39375FFECFAB16D87FEA770AC" authority="Nabozhenko, 2022" authorityName="Nabozhenko" authorityYear="2022" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329375FFECFAB16D87FA5170F0" box="[1298,1436,1699,1725]" italics="true" pageId="2" pageNumber="298">E. kasatkini</emphasis>
Nabozhenko, 2022
</taxonomicName>
from Central
<collectingRegion id="498418C29375FFECFDA76DE3FD9F70AC" box="[516,594,1735,1761]" country="Iran" name="Alborz" pageId="2" pageNumber="298">Alborz</collectingRegion>
Mts. by the completely beaded pronotum (at least in male) and longitudinal rugosity in apical part of the basal piece of the aedeagus, but differs in the following characters from:
</paragraph>
<paragraph id="8BFFD6209375FFECFF646C2BFADF7190" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="298">
<emphasis id="B9340A329375FFECFF646C2BFEB27164" box="[199,383,1807,1833]" italics="true" pageId="2" pageNumber="298">
<taxonomicName id="4C40ADA39375FFECFF406C2BFEB27164" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[227,383,1807,1833]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">E. huedepohli</taxonomicName>
</emphasis>
: prothoracic hypomera with coarse and dense separate punctures among weak wrinkles in anterior two thirds (
<figureCitation id="137BCAA59375FFECFED26C17FE057100" box="[369,456,1843,1869]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1G</figureCitation>
), coarse longitudinal wrinkles in
<taxonomicName id="4C40ADA39375FFECFC9B6C17FC1F7100" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[824,978,1843,1869]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFC9B6C17FC1F7100" box="[824,978,1843,1869]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
</taxonomicName>
; lateral margins of prothoracic hypomera distinctly flattened (
<figureCitation id="137BCAA59375FFECFEDD6C73FE1A713C" box="[382,471,1879,1905]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1G</figureCitation>
) and widely clearly flattened in
<taxonomicName id="4C40ADA39375FFECFCF66C73FC3E713C" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[853,1011,1879,1905]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFCF66C73FC3E713C" box="[853,1011,1879,1905]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
</taxonomicName>
; metatibiae moderately bent in male (
<figureCitation id="137BCAA59375FFECFF3C6C5FFF3B71D8" box="[159,246,1915,1941]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1C</figureCitation>
), strongly bent in
<taxonomicName id="4C40ADA39375FFECFE656C5FFDAF71D8" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[454,610,1915,1941]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFE656C5FFDAF71D8" box="[454,610,1915,1941]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
</taxonomicName>
and strongly bent in female (
<figureCitation id="137BCAA59375FFECFC0C6C5FFBCB71D8" box="[943,1030,1915,1941]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1D</figureCitation>
) while is not bent in
<taxonomicName id="4C40ADA39375FFECFB5B6C5FFA5971D8" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[1272,1428,1915,1941]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFB5B6C5FFA5971D8" box="[1272,1428,1915,1941]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
</taxonomicName>
; apex of the apical piece of the aedeagus narrowed and emarginated before apex (
<figureCitation id="137BCAA59375FFECFB836CBBFBB571F4" box="[1056,1144,1951,1977]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="2" pageNumber="298">Fig. 1K</figureCitation>
), weakly rounded and not emarginated in
<taxonomicName id="4C40ADA39375FFECFEE66CE7FE2D7190" baseAuthorityName="Kulzer" baseAuthorityYear="1964" box="[325,480,1987,2013]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="298" phylum="Arthropoda" rank="species" species="huedepohli">
<emphasis id="B9340A329375FFECFEE66CE7FE2D7190" box="[325,480,1987,2013]" italics="true" pageId="2" pageNumber="298">E. huedepohli</emphasis>
</taxonomicName>
(see photographs and description of this species in
<bibRefCitation id="EFD1ABD19375FFECFB816CE7FAC37190" author="Nabozhenko M. V. &amp; Gagarina L. V. &amp; Chigray I. A. &amp; Nabozhenko S. V." box="[1058,1294,1987,2013]" pageId="2" pageNumber="298" pagination="16 - 34" refId="ref2558" refString="Nabozhenko M. V., Gagarina L. V., Chigray I. A. &amp; Nabozhenko S. V. (2022 b) Trophic relations and ecological niches of darkling beetles of the genus Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) in the Caucasus and description of a new species from Abkhazia. South of Russia: ecology, development, 17 (3), 16 - 34. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2022 - 3 - 16 - 34" type="journal article" year="2022">Nabozhenko (2022))</bibRefCitation>
.
</paragraph>
<paragraph id="8BFFD6209374FFEDFF646BB3FC0F77E0" blockId="3.[151,1436,151,465]" pageId="3" pageNumber="299">
<taxonomicName id="4C40ADA39374FFEDFF406BB3FEA676FC" authorityName="Nabozhenko" authorityYear="2022" box="[227,363,151,177]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFF406BB3FEA676FC" box="[227,363,151,177]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
: lateral margins of pronotum emarginated (
<figureCitation id="137BCAA59374FFEDFCCB6BB3FC0D76FC" box="[872,960,151,177]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1E</figureCitation>
), not emarginated in
<taxonomicName id="4C40ADA39374FFEDFB1F6BB3FA8976FC" authorityName="Nabozhenko" authorityYear="2022" box="[1212,1348,151,177]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFB1F6BB3FA8976FC" box="[1212,1348,151,177]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
; lateral margins of pronotum widely beaded in basal half (
<figureCitation id="137BCAA59374FFEDFD706B9FFCE57698" box="[723,808,187,213]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1E</figureCitation>
) and less beaded in
<taxonomicName id="4C40ADA39374FFEDFBB36B9FFB587698" authorityName="Nabozhenko" authorityYear="2022" box="[1040,1173,187,213]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFBB36B9FFB587698" box="[1040,1173,187,213]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
; prothoracic hypomera with coarse and dense separate punctures among weak wrinkles in anterior two thirds (
<figureCitation id="137BCAA59374FFEDFBDE6BFBFB1B76B4" box="[1149,1238,223,249]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1G</figureCitation>
) while punctures often merged and poorly visible among coarse wrinkles in
<taxonomicName id="4C40ADA39374FFEDFC986A27FC0C7750" authorityName="Nabozhenko" authorityYear="2022" box="[827,961,259,285]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFC986A27FC0C7750" box="[827,961,259,285]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
; lateral margins of prothoracic hypomera distinctly flattened (
<figureCitation id="137BCAA59374FFEDFED56A03FE01770C" box="[374,460,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1G</figureCitation>
) and weakly flattened in
<taxonomicName id="4C40ADA39374FFEDFD446A03FCA7770C" authorityName="Nabozhenko" authorityYear="2022" box="[743,874,295,321]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFD446A03FCA7770C" box="[743,874,295,321]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
; metatibiae moderately bent in male (
<figureCitation id="137BCAA59374FFEDFAB36A03FAA9770C" box="[1296,1380,295,321]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1C</figureCitation>
) and very weakly bent in
<taxonomicName id="4C40ADA39374FFEDFE216A6FFDC57728" authorityName="Nabozhenko" authorityYear="2022" box="[386,520,331,357]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFE216A6FFDC57728" box="[386,520,331,357]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
and strongly bent in female (
<figureCitation id="137BCAA59374FFEDFCF96A6FFC7F7728" box="[858,946,331,357]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1D</figureCitation>
) but not in
<taxonomicName id="4C40ADA39374FFEDFB986A6FFB0C7728" authorityName="Nabozhenko" authorityYear="2022" box="[1083,1217,331,357]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFB986A6FFB0C7728" box="[1083,1217,331,357]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
; apex of the apical piece of the aedeagus narrowed and emarginated before apex (
<figureCitation id="137BCAA59374FFEDFCF46A4BFC6277C4" box="[855,943,367,393]" captionStart="FIGURE 1" captionStartId="1.[151,250,1814,1838]" captionTargetBox="[350,1233,671,1775]" captionTargetId="figure-133@1.[346,1241,666,1787]" captionTargetPageId="1" captionText="FIGURE 1. Euboeus kandovanensis sp. nov., habitus, details of structure. A = male, dorsally; B = male, ventrally; C = male metatibia; D = female, dorsally; E = male, head and pronotum, dorsally; F = ditto, dorso-laterally; G = prothoracic hypomeron; H= male inner sternite VIII, ventrally; I = spiculum gastrale; J = aedeagus ventrally; K = apical piece and part of basal piece of aedeagus, dorsally." figureDoi="http://doi.org/10.5281/zenodo.10145776" httpUri="https://zenodo.org/record/10145776/files/figure.png" pageId="3" pageNumber="299">Fig. 1K</figureCitation>
), wider and not emarginated in
<taxonomicName id="4C40ADA39374FFEDFAB46A4BFA5177C4" authorityName="Nabozhenko" authorityYear="2022" box="[1303,1436,367,393]" class="Insecta" family="Tenebrionidae" genus="Euboeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="299" phylum="Arthropoda" rank="species" species="kasatkini">
<emphasis id="B9340A329374FFEDFAB46A4BFA5177C4" box="[1303,1436,367,393]" italics="true" pageId="3" pageNumber="299">E. kasatkini</emphasis>
</taxonomicName>
(see photographs and description of this species in
<bibRefCitation id="EFD1ABD19374FFEDFD716AB7FC7377E0" author="Nabozhenko M. V. &amp; Gagarina L. V. &amp; Chigray I. A. &amp; Nabozhenko S. V." box="[722,958,403,429]" pageId="3" pageNumber="299" pagination="16 - 34" refId="ref2558" refString="Nabozhenko M. V., Gagarina L. V., Chigray I. A. &amp; Nabozhenko S. V. (2022 b) Trophic relations and ecological niches of darkling beetles of the genus Nalassus Mulsant, 1854 (Coleoptera: Tenebrionidae) in the Caucasus and description of a new species from Abkhazia. South of Russia: ecology, development, 17 (3), 16 - 34. [in Russian] https: // doi. org / 10.18470 / 1992 - 1098 - 2022 - 3 - 16 - 34" type="journal article" year="2022">Nabozhenko (2022))</bibRefCitation>
.
</paragraph>
<paragraph id="8BFFD6209374FFEDFF646A92FAA4779C" blockId="3.[151,1436,151,465]" box="[199,1385,438,465]" pageId="3" pageNumber="299">
<emphasis id="B9340A329374FFEDFF646A92FE8E779D" bold="true" box="[199,323,438,464]" pageId="3" pageNumber="299">Bionomics</emphasis>
. Adults of this species were found under bushes and herbaceous plants during the day (
<figureCitation id="137BCAA59374FFEDFAB56A93FA91779C" box="[1302,1372,439,465]" captionStart="FIGURE 2" captionStartId="3.[151,250,1199,1223]" captionTargetBox="[336,1252,489,1176]" captionTargetId="figure-185@3.[336,1252,489,1176]" captionTargetPageId="3" captionText="FIGURE 2. Habitat of Euboeus (Pelorinus) kandovanensis sp. n. on the slope of the Mountain area in Chalus pass. Photographs by F. Eshraghi Mofrad." figureDoi="http://doi.org/10.5281/zenodo.10145778" httpUri="https://zenodo.org/record/10145778/files/figure.png" pageId="3" pageNumber="299">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>