treatments-xml/data/03/BF/BE/03BFBE01131FFFCAFF751E862FC97540.xml
2024-06-21 12:22:17 +02:00

201 lines
19 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="39FEBE570D858F8AAE031CDC6F08D647" ID-DOI="10.5281/zenodo.193221" ID-GBIF-Dataset="b0a2eb6d-be07-47b1-a8c7-652397391ca5" ID-ISSN="1175-5326" ID-Zenodo-Dep="193221" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460126632289" checkinUser="plazi" docAuthor="Flechtmann, Carlos H. W. &amp; De, Dalva L." docDate="2010" docId="03BFBE01131FFFCAFF751E862FC97540" docLanguage="en" docName="zt02337p030.pdf" docOrigin="Zootaxa 2337" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Procalacarus perporosus Flechtmann &amp; De, 2010, n. sp." docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FF86C679131CFFC9FFE21E112E427357" masterDocTitle="New taxa in the Eriophyidae (Acari, Prostigmata) from forest trees in southern Brazil" masterLastPageNumber="30" masterPageNumber="18" pageNumber="21" updateTime="1698235033353" updateUser="plazi">
<mods:mods id="E94B56BD1363ED000C96B2A4B2CF34B2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4C2347A98B10F4C4F87A542EE0987AF2">
<mods:title id="676FFCE3838F41DA595127B19A98832E">New taxa in the Eriophyidae (Acari, Prostigmata) from forest trees in southern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="71DF2A3F175834CFC3DD32E7A8223CC0" type="personal">
<mods:role id="21B214D59A79E93A9B19213CCA2F965F">
<mods:roleTerm id="1078286A74472C572D6691EF129E7582">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E62AB9578C68699771817D26B07C911">Flechtmann, Carlos H. W.</mods:namePart>
</mods:name>
<mods:name id="97FF65733139046260C9D89A070EC39A" type="personal">
<mods:role id="A27ED00146800C3DE53DC5F9359ECF92">
<mods:roleTerm id="1BCE026F2B253B3587CBBEE36384DE63">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AD3CD6EE3658F238E98083C929384948">De, Dalva L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="485DEFE69C2EF9A118794CA3ECE0993B">text</mods:typeOfResource>
<mods:relatedItem id="A769288D12D762466A3BFC0AF8D83173" type="host">
<mods:titleInfo id="B362AD3C7FB695B84EE55ED2E640063A">
<mods:title id="E3957019BA81598D909BA3D3E26B4014">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C357EED84FD663DCDF823BCEA525E986">
<mods:date id="CC1D46BC2FEF0148601F9DB3E5278BF0">2010</mods:date>
<mods:detail id="AB3E64116157373F2F0BFF9EB748EE4D" type="volume">
<mods:number id="DC00B2C622DCBF50722FF1598C1BFEC3">2337</mods:number>
</mods:detail>
<mods:extent id="37C915E7631432B99D0BE9406F2B4E64" unit="page">
<mods:start id="BF8D141E1729BF8B02C540C2C1F66C86">18</mods:start>
<mods:end id="6AD21773C5DACB0EADE9D1F6EC56DD87">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="39E656FA8ECFF740BB2C63E863035EEF">journal article</mods:classification>
<mods:identifier id="2222812511B51B932194579EF44FA943" type="DOI">10.5281/zenodo.193221</mods:identifier>
<mods:identifier id="AC5085E0DBC9E34FE7D193FD8256E5FF" type="GBIF-Dataset">b0a2eb6d-be07-47b1-a8c7-652397391ca5</mods:identifier>
<mods:identifier id="4692EFC26DB5EE3343BB8307313B9E7B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5C6113E27EF4824706290F1F972CFEC3" type="Zenodo-Dep">193221</mods:identifier>
</mods:mods>
<treatment id="03BFBE01131FFFCAFF751E862FC97540" ID-DOI="http://doi.org/10.5281/zenodo.6197506" ID-GBIF-Taxon="124955565" ID-Zenodo-Dep="6197506" LSID="urn:lsid:plazi:treatment:03BFBE01131FFFCAFF751E862FC97540" httpUri="http://treatment.plazi.org/id/03BFBE01131FFFCAFF751E862FC97540" lastPageNumber="21" pageId="3" pageNumber="21">
<subSubSection id="C30C5C9C131FFFCAFF751E862EAD7380" pageId="3" pageNumber="21" type="nomenclature">
<paragraph id="8BA90F17131FFFCAFF751E862C4C73E6" blockId="3.[151,526,151,215]" box="[151,526,151,177]" pageId="3" pageNumber="21">
<heading id="D0E1B87B131FFFCAFF751E862C4C73E6" bold="true" box="[151,526,151,177]" fontSize="11" level="1" pageId="3" pageNumber="21" reason="1">
<emphasis id="B962D305131FFFCAFF751E862C4C73E6" bold="true" box="[151,526,151,177]" pageId="3" pageNumber="21">
<taxonomicName id="4C167494131FFFCAFF751E862F8673E6" box="[151,452,151,177]" class="Arachnida" family="Eriophyidae" genus="Procalacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="perporosus" status="sp. nov.">
<emphasis id="B962D305131FFFCAFF751E862F8673E6" bold="true" box="[151,452,151,177]" italics="true" pageId="3" pageNumber="21">Procalacarus perporosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2516E7E131FFFCAFE2E1E862C4C73E6" box="[460,526,151,177]" pageId="3" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA90F17131FFFCAFF751EAC2EAD7380" blockId="3.[151,526,151,215]" box="[151,239,189,215]" pageId="3" pageNumber="21">
(
<figureCitation id="132D1392131FFFCAFF7D1EAC2EA77380" box="[159,229,189,215]" captionStart="FIGURE 2" captionStartId="4.[151,255,1952,1976]" captionTargetBox="[199,1406,227,1908]" captionTargetId="figure@4.[154,1432,194,1930]" captionTargetPageId="4" captionText="FIGURE 2. Procalacarus perporosus n. sp., female. D—dorsal aspect; L 1 — leg I; L 2 — leg II; VA—ventral anterior or coxigenital region." httpUri="https://zenodo.org/record/193223/files/figure.png" pageId="3" pageNumber="21">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C5C9C131FFFCAFF751F162B1B72E0" pageId="3" pageNumber="21" type="diagnosis">
<paragraph id="8BA90F17131FFFCAFF751F162B1B72E0" blockId="3.[151,1437,263,1559]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF751F162F507276" bold="true" box="[151,274,263,289]" pageId="3" pageNumber="21">Diagnosis</emphasis>
.
<taxonomicName id="4C167494131FFFCAFEC01F192F847276" box="[290,454,264,289]" class="Arachnida" family="Eriophyidae" genus="Procalacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="perporosus">
<emphasis id="B962D305131FFFCAFEC01F192F847276" box="[290,454,264,289]" italics="true" pageId="3" pageNumber="21">P. perporosus</emphasis>
</taxonomicName>
is the second species assigned to this genus and is distinctive in presenting eight longitudinal rows of slit “wax” producing pores on anterior one third of opisthosomal dorsum, which merge into five rows on posterior two thirds (five rows throughout entire opisthosoma in
<taxonomicName id="4C167494131FFFCAFB081F432F8E72C5" authority="Mohanasundaram, 1983" authorityName="Mohanasundaram" authorityYear="1983" class="Arachnida" family="Eriophyidae" genus="Procalacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="aliyarensis">
<emphasis id="B962D305131FFFCAFB081F432BDF723C" box="[1258,1437,338,363]" italics="true" pageId="3" pageNumber="21">P. aliyarensis</emphasis>
<bibRefCitation id="EF8772E6131FFFCAFF751F692F8E72C5" author="Mohanasundaram" box="[151,460,376,402]" pageId="3" pageNumber="30" refString="Mohanasundaram, M. (1983) Indian Eriophyid studies IV. Record of new Phyllocoptine Mites (Phyllocoptinae: Eriophyidae: Acarina) from South India. Acarologia, 34 (1), 13 - 35." type="journal article" year="1983">Mohanasundaram, 1983</bibRefCitation>
</taxonomicName>
) and in having the median line on the posterior half of the prodorsal shield ornamentation and surrounded by a clear geometric pattern as figured (no median line in
<taxonomicName id="4C167494131FFFCAFB4E1F8C2B0572E1" box="[1196,1351,413,438]" class="Arachnida" family="Eriophyidae" genus="Procalacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="aliyarensis">
<emphasis id="B962D305131FFFCAFB4E1F8C2B0572E1" box="[1196,1351,413,438]" italics="true" pageId="3" pageNumber="21">P. aliyarensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30C5C9C131FFFCAFF241FD32FC377BB" pageId="3" pageNumber="21" type="description">
<paragraph id="8BA90F17131FFFCAFF241FD32C947790" blockId="3.[151,1437,263,1559]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF241FD32F60728B" bold="true" box="[198,290,450,476]" pageId="3" pageNumber="21">Female</emphasis>
(n = 6). Body spindleform, robust, 179(173216), 69(6580) wide. Dorsum of opisthosoma entirely covered with five thick longitudinal white wax-like strands and prodorsal shield with irregular “waxy” masses.
<emphasis id="B962D305131FFFCAFEBC1C1C2C427170" bold="true" box="[350,512,525,551]" pageId="3" pageNumber="21">Gnathosoma</emphasis>
: apical seta (
<emphasis id="B962D305131FFFCAFD421C1C2CED7171" box="[672,687,525,550]" italics="true" pageId="3" pageNumber="21">d</emphasis>
) 11(1012); basal seta (
<emphasis id="B962D305131FFFCAFC311C1C2DAD7171" box="[979,1007,525,550]" italics="true" pageId="3" pageNumber="21">ep</emphasis>
) robust, spine-like, 3(3); chelicerae 17(1619).
<emphasis id="B962D305131FFFCAFEC61C232FBA711B" bold="true" box="[292,504,562,588]" pageId="3" pageNumber="21">Prodorsal shield</emphasis>
47(4147), 61(5975) wide, shield with five cells on each side; median line present on posterior half and inserted in a lozenge design as figured. Scapular tubercles 26(2628) apart; scapular setae missing. Shield with a broad, rounded anterior lobe, 7(67) and 26(2331) basal width.
<emphasis id="B962D305131FFFCAFABB1C6C2BD671C0" bold="true" box="[1369,1428,637,663]" pageId="3" pageNumber="21">Legs</emphasis>
: tibia I and genu II setae missing. Leg
<date id="FFA829D7131FFFCAFDBC1CB32CD271EB" box="[606,656,674,700]" pageId="3" pageNumber="21" value="1930-01">I 30</date>
(2931); femur 9(910), femoral seta (
<emphasis id="B962D305131FFFCAFB8A1CB32AC671EC" box="[1128,1156,674,699]" italics="true" pageId="3" pageNumber="21">bv</emphasis>
) 11(1114); genu 4(4), genual seta (
<emphasis id="B962D305131FFFCAFED31CD92F0971B6" box="[305,331,712,737]" italics="true" pageId="3" pageNumber="21">l”</emphasis>
) 31(2632); tibia 6(68), tibial seta (
<emphasis id="B962D305131FFFCAFCF01CD92D6771B6" box="[786,805,712,737]" italics="true" pageId="3" pageNumber="21">l</emphasis>
) missing; tarsus 5(57), dorsal seta (
<emphasis id="B962D305131FFFCAFB0B1CD92B4671B6" box="[1257,1284,712,737]" italics="true" pageId="3" pageNumber="21">ft</emphasis>
) 19(1719), lateral seta (
<emphasis id="B962D305131FFFCAFECA1CFC2F087051" box="[296,330,749,774]" italics="true" pageId="3" pageNumber="21">ft”</emphasis>
) 22(2123), unguinal seta (
<emphasis id="B962D305131FFFCAFD761CFC2CEF7051" box="[660,685,749,774]" italics="true" pageId="3" pageNumber="21">u</emphasis>
) 5(46), solenidion 7(57), distally expanded, empodium 5(5), 5-rayed. Leg
<date id="FFA829D7131FFFCAFED61D032F2F707B" box="[308,365,786,812]" pageId="3" pageNumber="21" value="1928-02">II 28</date>
(2730); femur 9(910),
<emphasis id="B962D305131FFFCAFD731D032CEE707C" box="[657,684,786,811]" italics="true" pageId="3" pageNumber="21">bv</emphasis>
11(1011); genu 4(4),
<emphasis id="B962D305131FFFCAFC591D032D91707C" box="[955,979,786,811]" italics="true" pageId="3" pageNumber="21">l”</emphasis>
missing; tibia 6(-6); tarsus 6(67),
<emphasis id="B962D305131FFFCAFA601D032BDE707C" box="[1410,1436,786,811]" italics="true" pageId="3" pageNumber="21">ft</emphasis>
8(68),
<emphasis id="B962D305131FFFCAFF141D292F5A7006" box="[246,280,824,849]" italics="true" pageId="3" pageNumber="21">ft”</emphasis>
18(1821),
<emphasis id="B962D305131FFFCAFE4E1D292F877006" box="[428,453,824,849]" italics="true" pageId="3" pageNumber="21">u</emphasis>
5(45), solenidion 5(57), expanded distally, empodium 5(45), 5-rayed.
<emphasis id="B962D305131FFFCAFAAF1D262BDE7006" bold="true" box="[1357,1436,823,849]" pageId="3" pageNumber="21">Coxae</emphasis>
smooth, with all usual setae. Coxal seta I (
<emphasis id="B962D305131FFFCAFD6D1D4C2CEE7021" box="[655,684,861,886]" italics="true" pageId="3" pageNumber="21">1b</emphasis>
) 9(-11), 12(1114) apart; coxal seta II (
<emphasis id="B962D305131FFFCAFB641D4C2AE17021" box="[1158,1187,861,886]" italics="true" pageId="3" pageNumber="21">1a</emphasis>
) 14(1214), 9(810) apart; coxal seta III (
<emphasis id="B962D305131FFFCAFE771D932FF170CC" box="[405,435,898,923]" italics="true" pageId="3" pageNumber="21">2a</emphasis>
) 33(3140), 26(2339) apart. Sternal line 8(810). Coxisternal region with 5(56) annuli, microtuberculate.
<emphasis id="B962D305131FFFCAFE3B1DB62C117096" bold="true" box="[473,595,935,961]" pageId="3" pageNumber="21">Genitalia</emphasis>
28(2729) wide, 16(1621) long; epigynum with short radial lines distally, otherwise smooth. Genital seta (
<emphasis id="B962D305131FFFCAFD651DDC2CE770B1" box="[647,677,973,998]" italics="true" pageId="3" pageNumber="21">3a</emphasis>
) 14(1114).
<emphasis id="B962D305131FFFCAFCA61DDC2DAF70B0" bold="true" box="[836,1005,973,999]" pageId="3" pageNumber="21">Opisthosoma</emphasis>
with eight longitudinal rows of slit pores, producing a white wax-like substance, on anterior one third opisthosoma dorsum, which merge into five rows of pores on posterior two thirds. Lateral seta (
<emphasis id="B962D305131FFFCAFCD11A092D0D7766" box="[819,847,1048,1073]" italics="true" pageId="3" pageNumber="21">c2</emphasis>
) 21(1825), on annulus 1(13). Ventral seta I (
<emphasis id="B962D305131FFFCAFA611A092BD07766" box="[1411,1426,1048,1073]" italics="true" pageId="3" pageNumber="21">d</emphasis>
) 48(4555), longer than distance
<emphasis id="B962D305131FFFCAFDFD1A2C2C077701" box="[543,581,1085,1111]" italics="true" pageId="3" pageNumber="21">d -e</emphasis>
, 31(3135) apart, on annulus 16(1620); ventral seta II (
<emphasis id="B962D305131FFFCAFB1A1A2C2B477701" box="[1272,1285,1085,1110]" italics="true" pageId="3" pageNumber="21">e</emphasis>
) 20(1720), 21(1822) apart, on annulus 35(3540); ventral seta III (
<emphasis id="B962D305131FFFCAFCCC1A722D74772B" box="[814,822,1123,1148]" italics="true" pageId="3" pageNumber="21">f</emphasis>
) 22(2123), 23(2326) apart, on annulus 53(5359) or 6(67)th from rear. Total dorsal annuli 66(6673); total ventral annuli 59(5966), microtuberculate. Caudal seta (
<emphasis id="B962D305131FFFCAFF371ABC2EB07791" box="[213,242,1197,1222]" italics="true" pageId="3" pageNumber="21">h2</emphasis>
) 42(4248); accessory seta (
<emphasis id="B962D305131FFFCAFDA71ABC2C217791" box="[581,611,1197,1222]" italics="true" pageId="3" pageNumber="21">h1</emphasis>
) missing.
</paragraph>
<paragraph id="8BA90F17131FFFCAFF261AC32FC377BB" blockId="3.[151,1437,263,1559]" box="[196,385,1234,1260]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF261AC32F4177BB" bold="true" box="[196,259,1234,1260]" pageId="3" pageNumber="21">Male</emphasis>
. Not seen.
</paragraph>
</subSubSection>
<subSubSection id="C30C5C9C131FFFCAFF261AE62D3076F0" pageId="3" pageNumber="21" type="materials_examined">
<paragraph id="8BA90F17131FFFCAFF261AE62A0976D5" blockId="3.[151,1437,263,1559]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF261AE62F3F7646" bold="true" box="[196,381,1271,1297]" pageId="3" pageNumber="21">
<typeStatus id="54ADB1B5131FFFCAFF261AE62F467646" box="[196,260,1271,1297]" pageId="3" pageNumber="21">Type</typeStatus>
material
</emphasis>
. Female
<typeStatus id="54ADB1B5131FFFCAFE101AE92C1D7645" box="[498,607,1272,1298]" pageId="3" pageNumber="21" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D10C49E131FFFCAFD7F1AE92D5B7645" box="[669,793,1272,1298]" pageId="3" pageNumber="21" type="female">35 female</specimenCount>
<typeStatus id="54ADB1B5131FFFCAFCC01AE92DDF7645" box="[802,925,1272,1298]" pageId="3" pageNumber="21" type="paratype">paratypes</typeStatus>
, from
<taxonomicName id="4C167494131FFFCAFC0D1AE92BDA7645" authority="Sw." authorityName="Sw." box="[1007,1432,1272,1298]" class="Aves" family="Bernieridae" genus="Randia" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="21" phylum="Chordata" rank="species" species="armata">
<emphasis id="B962D305131FFFCAFC0D1AE92AE87646" box="[1007,1194,1272,1297]" italics="true" pageId="3" pageNumber="21">Randia armata</emphasis>
(Sw.) (Rubiaceae)
</taxonomicName>
, Campus Embrapa Florestas, Colombo, Paraná,
<collectingCountry id="F3014F87131FFFCAFD5D1B0C2D487660" box="[703,778,1309,1335]" name="Brazil" pageId="3" pageNumber="21">Brazil</collectingCountry>
,
<geoCoordinate id="EE2269D0131FFFCAFCF61B0C2D307660" box="[788,882,1309,1335]" direction="south" orientation="latitude" pageId="3" pageNumber="21" precision="925" value="-25.316668">25º19S</geoCoordinate>
and
<geoCoordinate id="EE2269D0131FFFCAFC481B0C2A517660" box="[938,1043,1309,1335]" direction="west" orientation="longitude" pageId="3" pageNumber="21" precision="925" value="-49.15">49º09W</geoCoordinate>
,
<date id="FFA829D7131FFFCAFBFC1B0C2AF57660" box="[1054,1207,1309,1335]" pageId="3" pageNumber="21" value="2008-08">August 2008</date>
, collected by Dalva L. de Queiroz, on 8 microscopic preparations, in the collection of Departamento de Entomologia e Acarologia,
<collectionCode id="ED0797D2131FFFCAFECA1B792FC576D5" box="[296,391,1384,1410]" pageId="3" pageNumber="21">ESALQ</collectionCode>
Universidade de São Paulo, Piracicaba, São Paulo,
<collectingCountry id="F3014F87131FFFCAFC1E1B792A0A76D5" box="[1020,1096,1384,1410]" name="Brazil" pageId="3" pageNumber="21">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BA90F17131FFFCAFF261B9C2D3076F0" blockId="3.[151,1437,263,1559]" box="[196,882,1421,1447]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF261B9C2FCA76F0" bold="true" box="[196,392,1421,1447]" pageId="3" pageNumber="21">Relation to host</emphasis>
. Vagrant on leaves; no apparent damage.
</paragraph>
</subSubSection>
<subSubSection id="C30C5C9C131FFFCAFF261BA32FC97540" pageId="3" pageNumber="21" type="etymology">
<paragraph id="8BA90F17131FFFCAFF261BA32FC97540" blockId="3.[151,1437,263,1559]" pageId="3" pageNumber="21">
<emphasis id="B962D305131FFFCAFF261BA32F08769B" bold="true" box="[196,330,1458,1484]" pageId="3" pageNumber="21">Etymology</emphasis>
. The specific designation
<taxonomicName id="4C167494131FFFCAFD661BA22D4A769B" box="[644,776,1459,1484]" class="Arachnida" family="Eriophyidae" genus="Procalacarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="21" phylum="Arthropoda" rank="species" species="perporosus">
<emphasis id="B962D305131FFFCAFD661BA22D4A769B" box="[644,776,1459,1484]" italics="true" pageId="3" pageNumber="21">perporosus</emphasis>
</taxonomicName>
is from the Latin,
<emphasis id="B962D305131FFFCAFC0A1BA22A4C769B" box="[1000,1038,1459,1484]" italics="true" pageId="3" pageNumber="21">per</emphasis>
, meaning very, exceedingly, plus
<emphasis id="B962D305131FFFCAFF751BC92EB976A6" box="[151,251,1496,1521]" italics="true" pageId="3" pageNumber="21">porosus</emphasis>
, Latin, meaning full of pores and refers to the large number of slit-like pores on the dorsal opisthosomal annuli.
</paragraph>
</subSubSection>
</treatment>
</document>