340 lines
25 KiB
XML
340 lines
25 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6094551" ID-GBIF-Dataset="18bd5418-ed8c-45a5-a662-5d2ebde01c75" ID-GBIF-Taxon="117135279" ID-Zenodo-Dep="6094551" checkinTime="1456484105736" checkinUser="donat" docAuthor="Ivan Löbl" docDate="2015" docId="03BF87FBFF91FFFF99E5942AFD09FBC3" docLanguage="en" docName="367-370_Löbl.pdf.imf" docOrigin="Mitteilungen Der Schweizerischen Entomologischen Gesellschaft 88" docTitle="Scaphisoma poussereaui Löbl, 2015, sp. nov." docType="treatment" docVersion="6" lastPageId="3" lastPageNumber="370" masterDocId="FF86FF83FF90FFFC9879957AFF96FF9F" masterDocTitle="Scaphisoma poussereaui sp. nov. from La Réunion (Coleoptera: Staphylinidae: Scaphidiinae), arange extension of the S. tricolor species group" masterLastPageNumber="370" masterPageNumber="367" pageId="1" pageNumber="368" updateTime="1644953490652" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Scaphisoma poussereaui sp. nov. from La Réunion (Coleoptera: Staphylinidae: Scaphidiinae), arange extension of the S. tricolor species group</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ivan Löbl</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Mitteilungen Der Schweizerischen Entomologischen Gesellschaft</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>88</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>367</mods:start>
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<mods:end>370</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.34001</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6094551" ID-GBIF-Taxon="117135279" ID-Zenodo-Dep="6094551" LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3" httpUri="http://treatment.plazi.org/id/03BF87FBFF91FFFF99E5942AFD09FBC3" lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368">
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<paragraph box="[412,779,336,360]" pageId="1" pageNumber="368">
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<emphasis bold="true" box="[412,779,336,360]" pageId="1" pageNumber="368">
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<heading bold="true" box="[412,779,336,360]" centered="true" fontSize="10" level="1" pageId="1" pageNumber="368" reason="4">
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<taxonomicName box="[412,684,336,360]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
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<emphasis bold="true" box="[412,684,336,360]" italics="true" pageId="1" pageNumber="368">Scaphisoma poussereaui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[692,779,336,360]" pageId="1" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection box="[550,641,370,389]" pageId="1" pageNumber="368" type="description">
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<paragraph box="[550,641,370,389]" pageId="1" pageNumber="368">
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(
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<figureCitation box="[556,633,370,389]" captionStart="Figs 1 – 3" captionStartId="2.[138,175,1120,1139]" captionTargetBox="[245,915,123,1102]" captionTargetId="figure@2.[245,915,122,1102]" captionTargetPageId="2" captionText="Figs 1 – 3. Aedeagus of Scaphisoma pousserreaui sp. nov., in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm." pageId="1" pageNumber="368">
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Figs
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<number box="[601,633,370,389]" pageId="1" pageNumber="368" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
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</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="368" type="materials_examined">
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<paragraph pageId="1" pageNumber="368">
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<materialsCitation ID-GBIF-Occurrence="1249305262" collectingDate="2015-02-19" collectionCode="MNHN" collectorName="Coll. J. Poussereau" country="Réunion" location="/ Chemin de Centure Maison Boyer J. P. Souche Jamblon" pageId="1" pageNumber="368" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis bold="true" box="[153,251,409,433]" italics="true" pageId="1" pageNumber="368">
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<typeStatus box="[153,251,409,433]" pageId="1" pageNumber="368">Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount box="[268,291,408,432]" pageId="1" pageNumber="368" type="male">♂</specimenCount>
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,
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<collectingCountry box="[296,430,407,433]" name="Réunion" pageId="1" pageNumber="368" value="Réunion">La Réunion</collectingCountry>
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||
<number box="[440,480,409,433]" pageId="1" pageNumber="368" value="974.0">974</number>
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<collectorName box="[490,699,409,433]" pageId="1" pageNumber="368">Coll. J. Poussereau</collectorName>
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<location LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3:8EC96036FF91FFFD9ABC94E3FE56FE51" country="Réunion" name="/ Chemin de Centure Maison Boyer J. P. Souche Jamblon" pageId="1" pageNumber="368">/Chemin de Centure Maison Boyer J.P. Souche Jamblon</location>
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<collectingDate box="[455,576,438,462]" pageId="1" pageNumber="368">
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<number box="[455,482,438,462]" pageId="1" pageNumber="368" value="19.0">19</number>
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||
0
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<number box="[502,516,438,462]" pageId="1" pageNumber="368" value="2.0">2</number>
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||
<number box="[523,576,438,462]" pageId="1" pageNumber="368" value="2015.0">2015</number>
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</collectingDate>
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(
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<collectionCode box="[592,679,438,462]" collectionName="France, Paris, Museum National d'Histoire Naturelle" pageId="1" pageNumber="368">MNHN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph pageId="1" pageNumber="368">
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<materialsCitation ID-GBIF-Occurrence="1249305263" collectingDate="2015-02-24" collectionCode="CPJP, MHNG" collectorName="Coll. J. Poussereau" country="Réunion" location="Le Tampon 800 m maison Lebeau M. tronc grand natte" pageId="1" pageNumber="368" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
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<emphasis bold="true" box="[153,261,468,491]" italics="true" pageId="1" pageNumber="368">
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<typeStatus box="[153,261,468,491]" pageId="1" pageNumber="368">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount box="[266,341,467,491]" pageId="1" pageNumber="368" type="male">
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<number box="[266,282,468,491]" pageId="1" pageNumber="368" value="3.0">3</number>
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♂♂
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</specimenCount>
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,
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<specimenCount box="[346,420,465,491]" pageId="1" pageNumber="368" type="female">
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<number box="[346,362,468,491]" pageId="1" pageNumber="368" value="4.0">4</number>
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♀♀
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</specimenCount>
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,
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<collectingCountry box="[425,553,465,491]" name="Réunion" pageId="1" pageNumber="368" value="Réunion">La Réunion</collectingCountry>
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<number box="[560,600,468,491]" pageId="1" pageNumber="368" value="974.0">974</number>
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<collectorName box="[607,811,467,491]" pageId="1" pageNumber="368">Coll. J. Poussereau</collectorName>
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/
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<location LSID="urn:lsid:plazi:treatment:03BF87FBFF91FFFF99E5942AFD09FBC3:8EC96036FF91FFFD9B4394A9FD89FD96" country="Réunion" name="Le Tampon 800 m maison Lebeau M. tronc grand natte" pageId="1" pageNumber="368">
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Le Tampon
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<quantity box="[957,1024,468,491]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="368" unit="m" value="800.0">800 m</quantity>
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maison Lebeau M. tronc grand natte
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</location>
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<collectingDate box="[550,672,497,521]" pageId="1" pageNumber="368">
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<number box="[550,577,497,521]" pageId="1" pageNumber="368" value="24.0">24</number>
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||
0
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||
<number box="[598,611,497,521]" pageId="1" pageNumber="368" value="2.0">2</number>
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<number box="[618,672,497,521]" pageId="1" pageNumber="368" value="2015.0">2015</number>
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</collectingDate>
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(
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<collectionCode box="[687,748,497,521]" pageId="1" pageNumber="368">CPJP</collectionCode>
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,
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<collectionCode box="[760,846,497,521]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="1" pageNumber="368">MHNG</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368" type="description">
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<paragraph lastPageId="3" lastPageNumber="370" pageId="1" pageNumber="368">
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<emphasis bold="true" box="[214,340,540,563]" italics="true" pageId="1" pageNumber="368">Description</emphasis>
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.Male. Length
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<quantity box="[491,647,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="1" pageNumber="368" unit="mm" value="1.95" valueMax="2.05" valueMin="1.85">1.85–2.05 mm</quantity>
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, width
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<quantity box="[721,878,540,563]" metricMagnitude="-3" metricUnit="m" metricValue="1.365" metricValueMax="1.46" metricValueMin="1.27" pageId="1" pageNumber="368" unit="mm" value="1.365" valueMax="1.46" valueMin="1.27">1.27–1.46 mm</quantity>
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. Head, mouthparts, antennae, pronotum, hypomera, and tarsi ochreous. Anterior two thirds of elytra dark brown to blackish, apical third of elytra yellowish. Mesoventrite and metaventrite with epimera and anepisterna reddish-brown. Coxae, femora, tibiae and abdomen light brown, slightly lighter than metaventrite. Antennae with relative length/width of antennomeres as:
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<date box="[522,608,686,710]" pageId="1" pageNumber="368" value="2013-03-08">
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III
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<number box="[557,585,686,710]" pageId="1" pageNumber="368" value="13.0">13</number>
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/
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<number box="[594,608,686,710]" pageId="1" pageNumber="368" value="8.0">8</number>
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</date>
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:
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||
<date box="[620,708,686,710]" pageId="1" pageNumber="368" value="1950-04-09">
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IV
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<number box="[657,686,686,710]" pageId="1" pageNumber="368" value="50.0">50</number>
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||
/
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<number box="[695,708,686,710]" pageId="1" pageNumber="368" value="9.0">9</number>
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||
</date>
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:
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||
<date box="[720,792,686,710]" pageId="1" pageNumber="368" value="1973-05-09">
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V
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||
<number box="[742,770,686,710]" pageId="1" pageNumber="368" value="73.0">73</number>
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||
/
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||
<number box="[779,792,686,710]" pageId="1" pageNumber="368" value="9.0">9</number>
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</date>
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||
:
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||
<date box="[804,905,686,710]" pageId="1" pageNumber="368" value="1960-06-10">
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VI
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||
<number box="[841,870,686,710]" pageId="1" pageNumber="368" value="60.0">60</number>
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||
/
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||
<number box="[879,905,686,710]" pageId="1" pageNumber="368" value="10.0">10</number>
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||
</date>
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||
:
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||
<date box="[917,1027,686,710]" pageId="1" pageNumber="368" value="1965-07-12">
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VII
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<number box="[963,991,686,710]" pageId="1" pageNumber="368" value="65.0">65</number>
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/
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<number box="[1000,1027,686,710]" pageId="1" pageNumber="368" value="12.0">12</number>
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</date>
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:
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<date box="[153,258,715,739]" pageId="1" pageNumber="368" value="1956-08-09">
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VIII
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<number box="[206,235,715,739]" pageId="1" pageNumber="368" value="56.0">56</number>
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/
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<number box="[244,258,715,739]" pageId="1" pageNumber="368" value="9.0">9</number>
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</date>
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||
:
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||
<date box="[268,368,715,739]" pageId="1" pageNumber="368" value="1968-09-12">
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IX
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<number box="[304,332,715,739]" pageId="1" pageNumber="368" value="68.0">68</number>
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/
|
||
<number box="[341,368,715,739]" pageId="1" pageNumber="368" value="12.0">12</number>
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</date>
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||
:
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||
<date box="[379,463,715,739]" pageId="1" pageNumber="368" value="1965-10-15">
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X
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<number box="[401,429,715,739]" pageId="1" pageNumber="368" value="65.0">65</number>
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/
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<number box="[438,463,715,739]" pageId="1" pageNumber="368" value="15.0">15</number>
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</date>
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:
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<date box="[474,577,715,739]" pageId="1" pageNumber="368" value="1968-11-14">
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XI
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<number box="[509,538,715,739]" pageId="1" pageNumber="368" value="68.0">68</number>
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/
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<number box="[546,573,715,739]" pageId="1" pageNumber="368" value="14.0">14</number>
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.
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</date>
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Pronotum and elytra not microsculptured. Pronotum with lateral margins evenly rounded, lateral margin carinae not visible in dorsal view, lateral striae very densely and distinctly punctate, disc with very fine and fairly dense punctation. Hypomera not microsculptured, with punctation very fine, hardly visible at
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<number box="[380,407,833,857]" pageId="1" pageNumber="368" value="25.0">25</number>
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times magnification. Exposed part of scutellum fairly large, about
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<quantity box="[220,315,862,886]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="368" unit="mm" value="0.05">0.05 mm</quantity>
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long. Elytra with lateral margins evenly rounded, lateral carinae in dorsal view concealed near bases, exposed posterior basal fourth to third; lateral striae very densely and fairly coarsely punctate; apical margins rounded, indistinctly crenulated near inner angles, inner apical angles situated posterior level of outer apical angles; sutural margins not raised, sutural striae fairly shallow, starting posterior to level of scutellar tip, subparallel; adsutural areas flat, with single puncture row; discal punctation of elytra dense and coarse except on yellowish apical area, fairly irregular, with punctures distinctly delimited, puncture intervals mostly distinctly larger than puncture diameters; punctation on light apical area very fine. Hind wings appearing fully developed. Mesanepisterna not microsculptured. Mesepimera as interval to mesocoxae and about
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<number box="[634,645,1155,1179]" pageId="1" pageNumber="368" value="3.0">3</number>
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times as long as wide. Metaventrite with strigulate microsculpture; punctation very fine, distinct punctures in impressed antecoxal rows and punctures bordering submesocoxal lines excepted; punctures also fairly coarse on small areas near inner margin of metacoxae. Middle part of metaventrite convex, shallowly impressed apically. Submesocoxal lines parallel, submesocoxal areas about
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<quantity box="[443,539,1302,1326]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="368" unit="mm" value="0.03">0.03 mm</quantity>
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long, about as sixth of interval to metacoxae. Metanepisterna convex, impressed toward suture, below level of metaventrite along suture, narrowed anteriad, with margin mostly oblique, rounded near anterior angles. Protibiae and metatibiae straight, mesotibiae distinctly curved posterior between basal third and apex, about as thick as protibiae, thicker than metatibiae. Protarsomeres
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<number box="[314,322,1449,1473]" pageId="1" pageNumber="368" value="1.0">1</number>
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strongly widened, about as wide as apices of protibiae. Protarsomeres
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<number box="[227,240,1478,1502]" pageId="1" pageNumber="368" value="2.0">2</number>
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and
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<number box="[291,302,1478,1502]" pageId="1" pageNumber="368" value="3.0">3</number>
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much narrower than protarsomere
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<number box="[694,708,1478,1502]" pageId="1" pageNumber="368" value="1.0">1</number>
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. Mesotarsomeres
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<number box="[911,919,1478,1502]" pageId="1" pageNumber="368" value="1.0">1</number>
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strongly widened, wider than apices of tibiae. Mesotarsomere
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<number box="[729,741,1214,1238]" pageId="2" pageNumber="369" value="2.0">2</number>
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slightly narrower than
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<number box="[996,1009,1214,1238]" pageId="2" pageNumber="369" value="1.0">1</number>
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, mesotarsomeres
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||
<number box="[317,328,1243,1268]" pageId="2" pageNumber="369" value="3.0">3</number>
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weakly widened. Abdomen with strigulate microsculpture, very finely punctate except on anarrow area posterior to the intercoxal process and at margins of submetaxocal line. Submetacoxal lines convex, submetacoxal areas
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<quantity metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="369" unit="mm" value="0.06">0.06 mm</quantity>
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, almost as third of interval to apical margin of sternite
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<number box="[766,780,1332,1356]" pageId="2" pageNumber="369" value="1.0">1</number>
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. Sternite
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<number box="[882,894,1331,1356]" pageId="2" pageNumber="369" value="6.0">6</number>
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flattened in middle, extended apically to form large, broadly rounded lobe. Aedeagus (
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<figureCitation box="[921,1013,1361,1385]" captionStart="Figs 1 – 3" captionStartId="2.[138,175,1120,1139]" captionTargetBox="[245,915,123,1102]" captionTargetId="figure@2.[245,915,122,1102]" captionTargetPageId="2" captionText="Figs 1 – 3. Aedeagus of Scaphisoma pousserreaui sp. nov., in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm." pageId="2" pageNumber="369">
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Figs
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<number box="[974,1013,1361,1385]" pageId="2" pageNumber="369" value="2.0" valueMax="3.0" valueMin="1.0">1–3</number>
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</figureCitation>
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||
)
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||
<quantity box="[138,234,1390,1414]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="369" unit="mm" value="1.28">1.28 mm</quantity>
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long. Basal bulb weakly sclerotized. Ventral branch of apical process of median lobe strongly inflexed lateroventrally, somewhat flattened, weakly narrowed toward tip in dorsal view. Parameres each with large and strongly sclerotized subbasal apophyses bearing membranous lobe. Internal sac with proximal, strongly bent, large tooth joined to moderately sclerotized area bearing three rods; apical part of internal sac very densely and very finely denticulate.
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</paragraph>
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<caption pageId="2" pageNumber="369" targetBox="[245,915,123,1102]" targetPageId="2">
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<paragraph pageId="2" pageNumber="369">
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Figs 1–3. Aedeagus of
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<taxonomicName box="[350,577,1120,1139]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="poussereaui" status="sp. nov.">
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<emphasis bold="true" box="[350,577,1120,1139]" italics="true" pageId="2" pageNumber="369">Scaphisoma pousserreaui</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[585,659,1120,1139]" pageId="2" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
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, in ventral view (1), in lateral view (2), median lobe, without parameres in dorsal view (3). Scale bar = 0.2 mm.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="370">
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Female. With most characters as in male, but: Pronotum slightly lighter than or as dark as anterior two thirds of elytra. Apical elytral margins almost oblique, with inner angles slightly prominent, tooth-like and very finely crenulated. Mesotibiae weakly bent, about as thick as metatibiae. All tarsomeres narrow. Sternite
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<number box="[1024,1038,271,294]" pageId="3" pageNumber="370" value="6.0">6</number>
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not lobed.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="370" type="etymology">
|
||
<paragraph pageId="3" pageNumber="370">
|
||
<emphasis bold="true" box="[214,328,329,353]" italics="true" pageId="3" pageNumber="370">Etymology</emphasis>
|
||
.The new species is named in honour of its collector, Jacques Poussereau., Dax, France
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="370" type="discussion">
|
||
<paragraph pageId="3" pageNumber="370">
|
||
<emphasis bold="true" box="[214,326,388,412]" italics="true" pageId="3" pageNumber="370">Comments</emphasis>
|
||
.The species is amember of the
|
||
<emphasis bold="true" box="[684,794,388,412]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
|
||
group defined in Löbl (
|
||
<number box="[162,219,417,441]" pageId="3" pageNumber="370" value="1975.0">1975</number>
|
||
:
|
||
<number box="[230,271,417,441]" pageId="3" pageNumber="370" value="272.0">272</number>
|
||
) containing the following species:
|
||
<emphasis bold="true" box="[650,770,417,441]" italics="true" pageId="3" pageNumber="370">S. affectum</emphasis>
|
||
Löbl,
|
||
<number box="[842,897,417,441]" pageId="3" pageNumber="370" value="2015.0">2015</number>
|
||
,
|
||
<emphasis bold="true" box="[909,1038,417,441]" italics="true" pageId="3" pageNumber="370">S. aspectum</emphasis>
|
||
Löbl,
|
||
<number box="[218,274,447,470]" pageId="3" pageNumber="370" value="2015.0">2015</number>
|
||
,
|
||
<emphasis bold="true" box="[286,424,447,470]" italics="true" pageId="3" pageNumber="370">S. borneense</emphasis>
|
||
Pic,
|
||
<number box="[480,536,447,470]" pageId="3" pageNumber="370" value="1916.0">1916</number>
|
||
,
|
||
<emphasis bold="true" box="[548,643,447,470]" italics="true" pageId="3" pageNumber="370">S. chujoi</emphasis>
|
||
Löbl,
|
||
<number box="[716,771,447,470]" pageId="3" pageNumber="370" value="1982.0">1982</number>
|
||
,
|
||
<emphasis bold="true" box="[784,877,447,470]" italics="true" pageId="3" pageNumber="370">S. hajeki</emphasis>
|
||
Löbl,
|
||
<number box="[950,1005,447,470]" pageId="3" pageNumber="370" value="2012.0">2012</number>
|
||
,
|
||
<emphasis bold="true" italics="true" pageId="3" pageNumber="370">S. luctans</emphasis>
|
||
Löbl,
|
||
<number box="[299,355,476,500]" pageId="3" pageNumber="370" value="1986.0">1986</number>
|
||
,
|
||
<emphasis bold="true" box="[365,499,476,500]" italics="true" pageId="3" pageNumber="370">S. maculiger</emphasis>
|
||
Löbl,
|
||
<number box="[568,624,476,500]" pageId="3" pageNumber="370" value="1875.0">1875</number>
|
||
,
|
||
<emphasis bold="true" box="[634,802,476,500]" italics="true" pageId="3" pageNumber="370">S. renominatum</emphasis>
|
||
Löbl,
|
||
<number box="[872,927,476,500]" pageId="3" pageNumber="370" value="1975.0">1975</number>
|
||
,
|
||
<emphasis bold="true" box="[937,1038,476,500]" italics="true" pageId="3" pageNumber="370">S. tarsale</emphasis>
|
||
Löbl,
|
||
<number box="[220,276,505,529]" pageId="3" pageNumber="370" value="2015.0">2015</number>
|
||
,
|
||
<emphasis bold="true" box="[290,525,505,529]" italics="true" pageId="3" pageNumber="370">S. testaceomaculatum</emphasis>
|
||
(Pic,
|
||
<number box="[595,649,505,529]" pageId="3" pageNumber="370" value="1915.0">1915</number>
|
||
) and
|
||
<emphasis bold="true" box="[715,826,505,529]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
|
||
Heller,
|
||
<number box="[919,976,505,529]" pageId="3" pageNumber="370" value="1917.0">1917</number>
|
||
. Two additional species,
|
||
<emphasis bold="true" box="[360,505,535,558]" italics="true" pageId="3" pageNumber="370">S. dentipenne</emphasis>
|
||
Löbl,
|
||
<number box="[578,632,535,558]" pageId="3" pageNumber="370" value="1971.0">1971</number>
|
||
and
|
||
<emphasis bold="true" box="[686,858,535,558]" italics="true" pageId="3" pageNumber="370">S. philippinense</emphasis>
|
||
Oberthür,
|
||
<number box="[978,1033,535,558]" pageId="3" pageNumber="370" value="1883.0">1883</number>
|
||
, known only from females, may also be members of the group. The group was defined mainly by having conspicuously large aedeagi with moderately sclerotized basal bulb, usually bearing aventral ridge supporting robust external muscles, the apical process of median lobe is bifid, strongly sclerotized and asymmetrical, the parameres are complex and the expanded membranous apex is supported by strongly sclerotized apophyses, and the internal sac is complex, often containing a curved proximal tooth. The species of the
|
||
<emphasis bold="true" box="[592,699,740,764]" italics="true" pageId="3" pageNumber="370">S. tricolor</emphasis>
|
||
group have comparatively large bodies, long antennae, the metaventrite with antecoxal puncture rows, the abdominal microsculpture strigulate, the male sternite
|
||
<number box="[628,640,798,822]" pageId="3" pageNumber="370" value="6.0">6</number>
|
||
with alarge, broadly rounded lobe. Having shortened sutural striae of the elytra is another diagnostic feature for some species and in anumber of examined undescribed Philippine species (the striae are elongate in
|
||
<emphasis bold="true" box="[278,397,887,910]" italics="true" pageId="3" pageNumber="370">S. affectum</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[410,538,887,910]" italics="true" pageId="3" pageNumber="370">S. aspectum</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[551,645,887,910]" italics="true" pageId="3" pageNumber="370">S. chujoi</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[659,751,887,910]" italics="true" pageId="3" pageNumber="370">S. hajeki</emphasis>
|
||
,
|
||
<emphasis bold="true" box="[764,899,887,910]" italics="true" pageId="3" pageNumber="370">S. maculiger</emphasis>
|
||
and
|
||
<emphasis bold="true" italics="true" pageId="3" pageNumber="370">S. testaceomaculatum</emphasis>
|
||
). The apical margins of the elytra are sexually dimorphic for some species, and males may also differ from females by their pronotal coloration.
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="poussereaui">
|
||
<emphasis bold="true" italics="true" pageId="3" pageNumber="370">Scaphisoma poussereaui</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguished from all members of the group by its elytral coloration and the aedeagal characters, in particular by the shape of the ventral branch of the apical process inflexed strongly ventrolaterally. In addition to these characters,
|
||
<taxonomicName box="[342,501,1063,1086]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="poussereaui">
|
||
<emphasis bold="true" box="[342,501,1063,1086]" italics="true" pageId="3" pageNumber="370">S. poussereaui</emphasis>
|
||
</taxonomicName>
|
||
may be easily distinguished from its Mascarene congeners by the elytra with short sutural striae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |