738 lines
98 KiB
XML
738 lines
98 KiB
XML
<document id="7F26B676333676BB606F87B2B143D236" ID-DOI="10.5281/zenodo.213893" ID-GBIF-Dataset="d47fdace-5cc7-448f-ac95-d444cb6f8ca7" ID-ISSN="1175-5326" ID-Zenodo-Dep="213893" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460553939180" checkinUser="plazi" docAuthor="Wong, Kingsley J. H., Shih, Hsi-Te & Chan, Benny K. K." docDate="2012" docId="03BF87DCFFB6B242FF44FF7BFBE9BB91" docLanguage="en" docName="zt03550p087.pdf" docOrigin="Zootaxa 3550" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ocypode mortoni George 1982" docType="treatment" docVersion="9" lastPageNumber="82" masterDocId="FF86FFA4FFB2B249FFD3FFA6FFFFBD56" masterDocTitle="The ghost crab Ocypode mortoni George, 1982 (Crustacea: Decapoda: Ocypodidae): redescription, distribution at its type locality, and the phylogeny of East Asian Ocypode species" masterLastPageNumber="87" masterPageNumber="71" pageNumber="75" updateTime="1698318788151" updateUser="plazi">
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<mods:title id="0D68F4FEFE583D63FEC2B4683E95F216">The ghost crab Ocypode mortoni George, 1982 (Crustacea: Decapoda: Ocypodidae): redescription, distribution at its type locality, and the phylogeny of East Asian Ocypode species</mods:title>
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<mods:namePart id="D9154D282259772A7A6E85EDFAFB9E80">Wong, Kingsley J. H.</mods:namePart>
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<mods:namePart id="A79F7B5F7A407EC4EBE52B305A9CEB7E">Shih, Hsi-Te</mods:namePart>
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<mods:namePart id="E9A1554BEB9E5F3D314303A0A736DA50">Chan, Benny K. K.</mods:namePart>
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<mods:date id="6F5B87E5F0EEB0EB21091808103D8F40">2012</mods:date>
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<treatment id="03BF87DCFFB6B242FF44FF7BFBE9BB91" ID-DOI="http://doi.org/10.5281/zenodo.6166333" ID-GBIF-Taxon="119555279" ID-Zenodo-Dep="6166333" LSID="urn:lsid:plazi:treatment:03BF87DCFFB6B242FF44FF7BFBE9BB91" httpUri="http://treatment.plazi.org/id/03BF87DCFFB6B242FF44FF7BFBE9BB91" lastPageId="11" lastPageNumber="82" pageId="4" pageNumber="75">
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<subSubSection id="C30C6541FFB6B24DFF44FF7BFEF5BC41" pageId="4" pageNumber="75" type="nomenclature">
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<paragraph id="8BA936CAFFB6B24DFF44FF7BFDEEBDA1" blockId="4.[151,529,221,279]" box="[151,529,221,247]" pageId="4" pageNumber="75">
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<heading id="D0E181A6FFB6B24DFF44FF7BFDEEBDA1" bold="true" box="[151,529,221,247]" fontSize="11" level="1" pageId="4" pageNumber="75" reason="1">
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<taxonomicName id="4C164D49FFB6B24DFF44FF7BFDEEBDA1" ID-CoL="48LWN" authority="George, 1982" authorityName="George" authorityYear="1982" box="[151,529,221,247]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="mortoni">
|
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<emphasis id="B962EAD8FFB6B24DFF44FF7BFDEEBDA1" bold="true" box="[151,529,221,247]" pageId="4" pageNumber="75">
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<emphasis id="B962EAD8FFB6B24DFF44FF7BFE99BDA1" bold="true" box="[151,358,221,247]" italics="true" pageId="4" pageNumber="75">Ocypode mortoni</emphasis>
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<bibRefCitation id="EF874B3BFFB6B24DFEBEFF7BFDEEBDA1" author="George" box="[365,529,221,247]" pageId="4" pageNumber="75" refString="George, R. W. (1982) The distribution and evolution of the ghost crabs (Ocypode spp.) of Hong Kong with a description of a new species. In: Morton, B. S. & Tseng, C. K. (Eds), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 185 - 194." type="book chapter" year="1982">George, 1982</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BA936CAFFB6B24DFF44FF59FEF5BC41" blockId="4.[151,529,221,279]" box="[151,266,255,279]" pageId="4" pageNumber="75">
|
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(
|
||
<figureCitation id="132D2A4FFFB6B24DFF4CFF59FEFDBC41" box="[159,258,255,279]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="4.[151,250,1535,1558]" captionStartId-1="5.[151,250,1963,1986]" captionStartId-2="6.[151,250,1952,1975]" captionStartId-3="7.[151,250,1734,1757]" captionTargetBox-0="[157,1433,635,1534]" captionTargetBox-1="[246,1338,320,1931]" captionTargetBox-2="[182,1385,193,1895]" captionTargetBox-3="[151,1435,193,1712]" captionTargetId-0="figure@4.[151,1436,601,1542]" captionTargetId-1="figure@5.[246,1338,317,1931]" captionTargetId-2="figure@6.[182,1387,193,1897]" captionTargetId-3="figure@7.[151,1436,193,1713]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." captionText-1="FIGURE 4. Morphology of male first pleopod (G 1) of O. mortoni: a, ventral view of left G 1; b, tip of left G 1, ventral view; c, dorsal view of right G 1; d, G 1 of type specimen, redrawn from George (1982)." captionText-2="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." captionText-3="FIGURE 6. Ocypode mortoni: ♂ from Sai Wan, carapace 24.5 mm x 21.2 mm (CEL-Ocy-HK- 003): a, dorsal view; b, frontal view." httpUri-0="https://zenodo.org/record/213896/files/figure.png" httpUri-1="https://zenodo.org/record/213897/files/figure.png" httpUri-2="https://zenodo.org/record/213898/files/figure.png" httpUri-3="https://zenodo.org/record/213899/files/figure.png" pageId="4" pageNumber="75">Figs 3–6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFB6B24DFF44FEEEFB19BCCC" pageId="4" pageNumber="75" type="reference_group">
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<paragraph id="8BA936CAFFB6B24DFF44FEEEFEC4BC2A" blockId="4.[151,1437,328,411]" pageId="4" pageNumber="75">
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<treatmentCitationGroup id="AB0611E4FFB6B24DFF44FEEEFEC4BC2A" pageId="4" pageNumber="75">
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<taxonomicName id="4C164D49FFB6B24DFF44FEEEFDEFBC0B" ID-CoL="48LWN" authority="George, 1982: 187" authorityName="George" authorityPageNumber="187" authorityYear="1982" box="[151,528,328,350]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB6B24DFF44FEEEFEB7BC08" box="[151,328,328,350]" italics="true" pageId="4" pageNumber="75">Ocypode mortoni</emphasis>
|
||
<treatmentCitation id="0AB710DBFFB6B24DFE9CFEEEFDEFBC0B" author="George" box="[335,528,328,349]" page="187" pageId="4" pageNumber="75" year="1982">
|
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<bibRefCitation id="EF874B3BFFB6B24DFE9CFEEEFE22BC0B" author="George" box="[335,477,328,349]" pageId="4" pageNumber="75" refString="George, R. W. (1982) The distribution and evolution of the ghost crabs (Ocypode spp.) of Hong Kong with a description of a new species. In: Morton, B. S. & Tseng, C. K. (Eds), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 185 - 194." type="book chapter" year="1982">George, 1982</bibRefCitation>
|
||
: 187
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, figs. 1C, 2C, pls. 3. ―
|
||
<treatmentCitation id="0AB710DBFFB6B24DFCD8FEEEFC09BC08" author="Dai" box="[779,1014,328,350]" page="457" pageId="4" pageNumber="75" year="1991">
|
||
<bibRefCitation id="EF874B3BFFB6B24DFCD8FEEEFC3CBC0B" author="Dai" box="[779,963,328,350]" pageId="4" pageNumber="75" refString="Dai, A. - Y. & Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 682 pp." type="book" year="1991">Dai & Yang 1991</bibRefCitation>
|
||
: 457
|
||
</treatmentCitation>
|
||
, fig. 203B. ―
|
||
<treatmentCitation id="0AB710DBFFB6B24DFB43FEEFFA9BBC08" author="Sakai" box="[1168,1380,328,350]" page="1159" pageId="4" pageNumber="75" year="2000">
|
||
<bibRefCitation id="EF874B3BFFB6B24DFB43FEEFFADABC0B" author="Sakai" box="[1168,1317,328,350]" pageId="4" pageNumber="75" refString="Sakai, K. (2000) On the occurrence of three species of crabs on Shikoku Island, Japan, and a new species, Pinnotheres taichungae nov. spec., from Taiwan (Decapoda, Brachyura). Crustaceana, 73, 1155 - 1162." type="journal article" year="2000">K. Sakai 2000</bibRefCitation>
|
||
: 1159
|
||
</treatmentCitation>
|
||
, figs. 2a–b, 3a–e.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFB6B24DFF44FE23FB19BCCC" blockId="4.[151,1437,328,411]" box="[151,1254,389,411]" pageId="4" pageNumber="75">
|
||
<treatmentCitationGroup id="AB0611E4FFB6B24DFF44FE23FB19BCCC" box="[151,1254,389,411]" pageId="4" pageNumber="75">
|
||
<taxonomicName id="4C164D49FFB6B24DFF44FE23FE9BBCCD" ID-CoL="48LWK" box="[151,356,389,411]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="macrocera">
|
||
<emphasis id="B962EAD8FFB6B24DFF44FE23FE9BBCCD" box="[151,356,389,411]" italics="true" pageId="4" pageNumber="75">Ocypode macrocera</emphasis>
|
||
</taxonomicName>
|
||
(not
|
||
<taxonomicName id="4C164D49FFB6B24DFE4AFE20FCD9BCCD" ID-CoL="48LWK" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[409,806,389,411]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="macrocera">
|
||
<emphasis id="B962EAD8FFB6B24DFE4AFE20FDDABCCD" box="[409,549,390,411]" italics="true" pageId="4" pageNumber="75">O. macrocera</emphasis>
|
||
H. Milne Edwards, 1837
|
||
</taxonomicName>
|
||
) ―
|
||
<treatmentCitation id="0AB710DBFFB6B24DFC80FE23FBE0BCCC" author="Dai" box="[851,1055,389,411]" page="370" pageId="4" pageNumber="75" year="1985">
|
||
<bibRefCitation id="EF874B3BFFB6B24DFC80FE23FC12BCCD" author="Dai" box="[851,1005,389,411]" pageId="4" pageNumber="75" refString="Dai, A. - Y., Song, Y. - Z & Yang, S. - L. (1985) A study on Ocypode from China. Acta Zootaxonomica Sinica, 10, 370 - 378. [in Chinese with English abstract]" type="journal article" year="1985">
|
||
Dai
|
||
<emphasis id="B962EAD8FFB6B24DFCADFE20FC55BCCD" box="[894,938,389,411]" italics="true" pageId="4" pageNumber="75">et al</emphasis>
|
||
. 1985
|
||
</bibRefCitation>
|
||
: 370
|
||
</treatmentCitation>
|
||
, figs. 1–7, pls. 1–2.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFB6B24DFF44FE61FAF8BF1A" pageId="4" pageNumber="75" type="materials_examined">
|
||
<paragraph id="8BA936CAFFB6B24DFF44FE61FAF8BF1A" blockId="4.[151,1436,455,589]" pageId="4" pageNumber="75">
|
||
<emphasis id="B962EAD8FFB6B24DFF44FE61FE7CBCB6" bold="true" box="[151,387,455,480]" pageId="4" pageNumber="75">Material examined.</emphasis>
|
||
63 2ƤƤ (CEL-Ocy-HK-003), Sai Wan,
|
||
<collectingCountry id="F301765AFFB6B24DFCBDFE6EFC08BCB6" box="[878,1015,456,480]" name="Hong Kong" pageId="4" pageNumber="75">Hong Kong</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFB6B24DFBD6FE61FB61BC89" box="[1029,1182,455,479]" pageId="4" pageNumber="75" value="2009-06-20">20 Jun. 2009</date>
|
||
, coll. K. Wong
|
||
<emphasis id="B962EAD8FFB6B24DFA8AFE6FFA72BCB6" box="[1369,1421,456,480]" italics="true" pageId="4" pageNumber="75">et al</emphasis>
|
||
.; 33 1Ƥ (CEL-Ocy-HK-021), Bui O,
|
||
<collectingCountry id="F301765AFFB6B24DFD91FE4BFD36BF53" box="[578,713,493,517]" name="Hong Kong" pageId="4" pageNumber="75">Hong Kong</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFB6B24DFD05FE4AFC94BF52" box="[726,875,492,516]" pageId="4" pageNumber="75" value="2009-06-21">21 Jun. 2009</date>
|
||
, coll. H.-N. Chen
|
||
<emphasis id="B962EAD8FFB6B24DFBEFFE4BFB91BF52" box="[1084,1134,492,516]" italics="true" pageId="4" pageNumber="75">et al</emphasis>
|
||
.; 1Ƥ (CEL-Ocy-HK-017), Fan Lau,
|
||
<collectingCountry id="F301765AFFB6B24DFED1FDB6FE76BF7E" box="[258,393,528,552]" name="Hong Kong" pageId="4" pageNumber="75">Hong Kong</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFB6B24DFE46FDA9FDD8BF71" box="[405,551,527,551]" pageId="4" pageNumber="75" value="2009-06-23">23 Jun. 2009</date>
|
||
, coll. K. Wong; 131Ƥ (ZRC-2012.0141), Sai Wan,
|
||
<collectingCountry id="F301765AFFB6B24DFBA1FDB6FB07BF7E" box="[1138,1272,528,552]" name="Hong Kong" pageId="4" pageNumber="75">Hong Kong</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFB6B24DFAD7FDA9FA67BF71" box="[1284,1432,527,551]" pageId="4" pageNumber="75" value="2009-06-20">20 Jun. 2009</date>
|
||
, coll. K. Wong
|
||
<emphasis id="B962EAD8FFB6B24DFEE9FD93FE94BF1A" box="[314,363,564,588]" italics="true" pageId="4" pageNumber="75">et al</emphasis>
|
||
.; 13 (NMNS-6879-001), Sai Wan,
|
||
<collectingCountry id="F301765AFFB6B24DFD2FFD93FC7DBF1B" box="[764,898,565,589]" name="Hong Kong" pageId="4" pageNumber="75">Hong Kong</collectingCountry>
|
||
,
|
||
<date id="FFA8100AFFB6B24DFC5DFD92FBDEBF1A" box="[910,1057,564,588]" pageId="4" pageNumber="75" value="2009-06-20">20 Jun. 2009</date>
|
||
, coll. K. Wong
|
||
<emphasis id="B962EAD8FFB6B24DFB1CFD93FAFFBF1A" box="[1231,1280,564,588]" italics="true" pageId="4" pageNumber="75">et al</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF696642FFB6B24DFF44FA59FF1EBB24" httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75" targetBox="[157,1433,635,1534]" targetPageId="4">
|
||
<paragraph id="8BA936CAFFB6B24DFF44FA59FF1EBB24" blockId="4.[151,1437,1535,1650]" pageId="4" pageNumber="75">
|
||
<emphasis id="B962EAD8FFB6B24DFF44FA59FEEDBB43" bold="true" box="[151,274,1535,1558]" pageId="4" pageNumber="75">FIGURE 3.</emphasis>
|
||
Morphology of
|
||
<taxonomicName id="4C164D49FFB6B24DFE6AF9A7FDD7BB40" box="[441,552,1537,1558]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="75" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB6B24DFE6AF9A7FDD7BB40" box="[441,552,1537,1558]" italics="true" pageId="4" pageNumber="75">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C30C6541FFB6B24DFF14F93FFF21BA17" pageId="4" pageNumber="75" type="diagnosis">
|
||
<paragraph id="8BA936CAFFB6B24DFF14F93FFF21BA17" blockId="4.[151,1437,1689,2037]" pageId="4" pageNumber="75">
|
||
<emphasis id="B962EAD8FFB6B24DFF14F93FFEBEBBE4" bold="true" box="[199,321,1689,1714]" pageId="4" pageNumber="75">Diagnosis.</emphasis>
|
||
Carapace subquadrate, regions defined by grooves; cornea with ocular stylets on the distal end, never longer than cornea, even in adult males (
|
||
<figureCitation id="132D2A4FFFB6B24DFD74F91BFD0ABB83" box="[679,757,1725,1749]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75">Figs. 3</figureCitation>
|
||
a, 5a–b). Outer surface of major cheliped palm often lemonyellow in color (
|
||
<figureCitation id="132D2A4FFFB6B24DFE8AF947FE60BBAF" box="[345,415,1761,1785]" captionStart="FIGURE 5" captionStartId="6.[151,250,1952,1975]" captionTargetBox="[182,1385,193,1895]" captionTargetId="figure@6.[182,1387,193,1897]" captionTargetPageId="6" captionText="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="4" pageNumber="75">Fig. 5</figureCitation>
|
||
b); stridulating ridge on inner surface of major cheliped consist of 44–65 evenly-spaced striation (
|
||
<figureCitation id="132D2A4FFFB6B24DFED6F8A3FEB4BA4B" box="[261,331,1797,1821]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75">Fig. 3</figureCitation>
|
||
c, i); base of immovable finger broad, compressed; finger tips of minor cheliped rounded (
|
||
<figureCitation id="132D2A4FFFB6B24DFABCF8A3FF5BBA17" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75">Fig. 3</figureCitation>
|
||
b–c).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFB6B241FF14F8EBFB36BEC6" lastPageId="8" lastPageNumber="79" pageId="4" pageNumber="75" type="description">
|
||
<paragraph id="8BA936CAFFB6B24CFF14F8EBFE86BC4A" blockId="4.[151,1437,1689,2037]" lastBlockId="5.[151,1436,151,284]" lastPageId="5" lastPageNumber="76" pageId="4" pageNumber="75">
|
||
<emphasis id="B962EAD8FFB6B24DFF14F8EBFEA6BA30" bold="true" box="[199,345,1869,1894]" pageId="4" pageNumber="75">Description.</emphasis>
|
||
Carapace subquadrate, broader than long, longitudinally convex, anterior 1/3 deflected, raised around branchial regions; surface evenly covered by beaded tubercles; regions markedly divided by grooves, those around cardiac region H-shaped, deeper; antero-lateral teeth absent; external orbital tooth forming right angle; anterior 1/3 of lateral margins roughly parallel, diverging posteriorly (
|
||
<figureCitation id="132D2A4FFFB6B24DFC6EF81FFBF2BA87" box="[957,1037,1977,2001]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75">Figs. 3</figureCitation>
|
||
a, 6a). Cornea on elongated stalks, bilobed, distal end extended with ocular stylets about half of cornea length in mature males (
|
||
<figureCitation id="132D2A4FFFB6B24DFB65F87BFAFABAA3" box="[1206,1285,2013,2037]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="4" pageNumber="75">Figs. 3</figureCitation>
|
||
a, 6a). Small, acute spine adjacent to antennule, suborbital ridge composed of 30–35 elongated granules; pterygostomian region sparsely covered by flattened granules (
|
||
<figureCitation id="132D2A4FFFB7B24CFD83FF1AFD6CBD82" box="[592,659,188,212]" captionStart="FIGURE 6" captionStartId="7.[151,250,1734,1757]" captionTargetBox="[151,1435,193,1712]" captionTargetId="figure@7.[151,1436,193,1713]" captionTargetPageId="7" captionText="FIGURE 6. Ocypode mortoni: ♂ from Sai Wan, carapace 24.5 mm x 21.2 mm (CEL-Ocy-HK- 003): a, dorsal view; b, frontal view." httpUri="https://zenodo.org/record/213899/files/figure.png" pageId="5" pageNumber="76">Fig. 6</figureCitation>
|
||
b). External maxillipeds flat, ischium longer than merus, both roughly quadrate (
|
||
<figureCitation id="132D2A4FFFB7B24CFED4FF79FEB4BDA1" box="[263,331,223,247]" captionStart="FIGURE 6" captionStartId="7.[151,250,1734,1757]" captionTargetBox="[151,1435,193,1712]" captionTargetId="figure@7.[151,1436,193,1713]" captionTargetPageId="7" captionText="FIGURE 6. Ocypode mortoni: ♂ from Sai Wan, carapace 24.5 mm x 21.2 mm (CEL-Ocy-HK- 003): a, dorsal view; b, frontal view." httpUri="https://zenodo.org/record/213899/files/figure.png" pageId="5" pageNumber="76">Fig. 6</figureCitation>
|
||
b), inner margins lined with long, light-colored stiff setae; exopod narrow, ill-developed, reaching 1/4 of merus length.
|
||
</paragraph>
|
||
<caption id="DF696642FFB7B24CFF44F80DFC4BBAB7" httpUri="https://zenodo.org/record/213897/files/figure.png" pageId="5" pageNumber="76" targetBox="[246,1338,320,1931]" targetPageId="5">
|
||
<paragraph id="8BA936CAFFB7B24CFF44F80DFC4BBAB7" blockId="5.[151,1437,1963,2017]" pageId="5" pageNumber="76">
|
||
<emphasis id="B962EAD8FFB7B24CFF44F80DFEECBA97" bold="true" box="[151,275,1963,1986]" pageId="5" pageNumber="76">FIGURE 4.</emphasis>
|
||
Morphology of male first pleopod (G1) of
|
||
<taxonomicName id="4C164D49FFB7B24CFD18F80AFCC5BA94" box="[715,826,1964,1986]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="76" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB7B24CFD18F80AFCC5BA94" box="[715,826,1964,1986]" italics="true" pageId="5" pageNumber="76">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
: a, ventral view of left G1; b, tip of left G1, ventral view; c, dorsal view of right G1; d, G1 of type specimen, redrawn from George (1982).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF696642FFB4B24FFF44F806FECCBA83" httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="6" pageNumber="77" targetBox="[182,1385,193,1895]" targetPageId="6">
|
||
<paragraph id="8BA936CAFFB4B24FFF44F806FECCBA83" blockId="6.[151,1436,1952,2005]" pageId="6" pageNumber="77">
|
||
<emphasis id="B962EAD8FFB4B24FFF44F806FEEABAE0" bold="true" box="[151,277,1952,1975]" pageId="6" pageNumber="77">FIGURE 5.</emphasis>
|
||
Life coloration of
|
||
<taxonomicName id="4C164D49FFB4B24FFE33F807FDADBAE0" box="[480,594,1953,1974]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="77" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB4B24FFE33F807FDADBAE0" box="[480,594,1953,1974]" italics="true" pageId="6" pageNumber="77">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF696642FFB5B24EFF44F960FF31BBAD" httpUri="https://zenodo.org/record/213899/files/figure.png" pageId="7" pageNumber="78" targetBox="[151,1435,193,1712]" targetPageId="7">
|
||
<paragraph id="8BA936CAFFB5B24EFF44F960FF31BBAD" blockId="7.[151,1435,1734,1787]" pageId="7" pageNumber="78">
|
||
<emphasis id="B962EAD8FFB5B24EFF44F960FEECBB8A" bold="true" box="[151,275,1734,1757]" pageId="7" pageNumber="78">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C164D49FFB5B24EFEC8F961FE33BB8B" box="[283,460,1735,1757]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="78" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB5B24EFEC8F961FE33BB8B" box="[283,460,1735,1757]" italics="true" pageId="7" pageNumber="78">Ocypode mortoni</emphasis>
|
||
</taxonomicName>
|
||
: ♂ from Sai Wan, carapace 24.5mm x 21.2 mm (CEL-Ocy-HK-003): a, dorsal view; b, frontal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA936CAFFB5B241FF14F881FE22BC3F" blockId="7.[151,1436,1831,2036]" lastBlockId="8.[151,1436,151,2030]" lastPageId="8" lastPageNumber="79" pageId="7" pageNumber="78">
|
||
Chelipeds asymmetrical; major merus triangular in latitudinal cross-section; lower fringe of inner surface leaflike, margin strongly serrated; carpus as long as broad, inner angle armed with strong tooth, outer-distal margin serrated; outer surface of palm covered with beaded tubercles of varies sizes, margins strongly serrated (
|
||
<figureCitation id="132D2A4FFFB5B24EFAEAF8C9FA82BAD1" box="[1337,1405,1903,1927]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="7" pageNumber="78">Fig. 3</figureCitation>
|
||
b); inner surface with latitudinal stridulating ridge composed of 44–65 evenly-spaced striations (
|
||
<figureCitation id="132D2A4FFFB5B24EFB70F832FB1ABAFA" box="[1187,1253,1940,1964]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="7" pageNumber="78">Fig. 3</figureCitation>
|
||
c, i); immovable finger triangular, high at base, which compressed and leaf-like, ridge of rounded tubercles lined subparallel to cutting edge, movable finger lined with ridge of large rounded tubercles; fingers shorter than palm, tips pointed (
|
||
<figureCitation id="132D2A4FFFBAB241FF4CFF31FF1DBDF9" box="[159,226,151,175]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Fig. 3</figureCitation>
|
||
b–c, i, 5b). Minor cheliped carpus slightly longer than broad, inner angle armed with strong tooth, fingers lined with ridge of rounded granules along respective median lines, tips rounded (
|
||
<figureCitation id="132D2A4FFFBAB241FBEDFF1AFB72BD82" box="[1086,1165,188,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Figs. 3</figureCitation>
|
||
d, 5b). Ambulatory legs slender, 2nd and 3rd subequal in total length, longest, 4th shortest; meri compressed, robust, distal end of anterior margin weakly armed with tooth (
|
||
<figureCitation id="132D2A4FFFBAB241FDFBFEA1FD8FBC49" box="[552,624,263,287]" captionStart="FIGURE 6" captionStartId="7.[151,250,1734,1757]" captionTargetBox="[151,1435,193,1712]" captionTargetId="figure@7.[151,1436,193,1713]" captionTargetPageId="7" captionText="FIGURE 6. Ocypode mortoni: ♂ from Sai Wan, carapace 24.5 mm x 21.2 mm (CEL-Ocy-HK- 003): a, dorsal view; b, frontal view." httpUri="https://zenodo.org/record/213899/files/figure.png" pageId="8" pageNumber="79">Fig. 6</figureCitation>
|
||
a). Propodi and dactyli of the first three pairs lined with rolls of soft plumose setae along median line on anterior surface; dactyli pointed, curved inwards for the first three pairs and outwards for the 4th (
|
||
<figureCitation id="132D2A4FFFBAB241FEADFEF7FE3DBC3F" box="[382,450,337,361]" captionStart="FIGURE 6" captionStartId="7.[151,250,1734,1757]" captionTargetBox="[151,1435,193,1712]" captionTargetId="figure@7.[151,1436,193,1713]" captionTargetPageId="7" captionText="FIGURE 6. Ocypode mortoni: ♂ from Sai Wan, carapace 24.5 mm x 21.2 mm (CEL-Ocy-HK- 003): a, dorsal view; b, frontal view." httpUri="https://zenodo.org/record/213899/files/figure.png" pageId="8" pageNumber="79">Fig. 6</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB241FF14FED3FC45BCE7" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
Urocardiac ossicle constricted beneath the T-shaped bifurcation (
|
||
<figureCitation id="132D2A4FFFBAB241FC02FED3FBE4BCDB" box="[977,1051,373,397]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Fig. 3</figureCitation>
|
||
h), and mildly expanded on the posterior half; zygocardiac ossicle bearing 16 comb-like teeth (
|
||
<figureCitation id="132D2A4FFFBAB241FC8AFE3FFC63BCE7" box="[857,924,409,433]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">
|
||
Fig.
|
||
<quantity id="4CEE9B2FFFBAB241FC5DFE3FFC63BCE7" box="[910,924,409,433]" metricMagnitude="-3" metricUnit="kg" metricValue="3.0" pageId="8" pageNumber="79" unit="g" value="3.0">3</quantity>
|
||
</figureCitation>
|
||
g).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB241FF14FE66FC36BFEE" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
Ventral surface of carapace glabrous except for tuft of soft setae between bases of 2nd and 3rd ambulatory legs. Male abdomen elongated; telson rounded, proximal margin convex; 6th somite as long as broad, lateral margins convex; 5th somite broader than long, roughly rectangular (
|
||
<figureCitation id="132D2A4FFFBAB241FC85FDA8FC60BF70" box="[854,927,526,550]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Fig. 3</figureCitation>
|
||
e). Male G1 slightly sinuous when viewed ventrally, not apparently tapering; distal part strongly curved outwards, tip divided into two chitinous beak-like lobes, upper lobe larger; neck of the distal curve fringed brating short setae (
|
||
<figureCitation id="132D2A4FFFBAB241FC3CFDF0FBCDBF38" box="[1007,1074,598,622]" captionStart="FIGURE 4" captionStartId="5.[151,250,1963,1986]" captionTargetBox="[246,1338,320,1931]" captionTargetId="figure@5.[246,1338,317,1931]" captionTargetPageId="5" captionText="FIGURE 4. Morphology of male first pleopod (G 1) of O. mortoni: a, ventral view of left G 1; b, tip of left G 1, ventral view; c, dorsal view of right G 1; d, G 1 of type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213897/files/figure.png" pageId="8" pageNumber="79">Fig. 4</figureCitation>
|
||
a–b, d). Female abdomen broad, telson as semicircle, narrow, proximal margin convex, 6th somite broader than long, lateral margins convex; preceding somites much broader than long, roughly rectangular (
|
||
<figureCitation id="132D2A4FFFBAB241FCBEFD06FC50BFEE" box="[877,943,672,696]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Fig. 3</figureCitation>
|
||
f).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB241FF14FD62FB36BEC6" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
<emphasis id="B962EAD8FFBAB241FF14FD62FEB0BF8B" bold="true" box="[199,335,708,733]" pageId="8" pageNumber="79">Coloration.</emphasis>
|
||
Dorsal surface of carapace dark brownish with light-colored thin sinuate lines (
|
||
<figureCitation id="132D2A4FFFBAB241FAE0FD62FA80BF8A" box="[1331,1407,708,732]" captionStart="FIGURE 5" captionStartId="6.[151,250,1952,1975]" captionTargetBox="[182,1385,193,1895]" captionTargetId="figure@6.[182,1387,193,1897]" captionTargetPageId="6" captionText="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
|
||
a); pterygostomian region and external maxillipeds creamish with microscopic dark dots; orbits grayish, ocular stylets yellowish (
|
||
<figureCitation id="132D2A4FFFBAB241FEC9FCAAFE9CBE72" box="[282,355,780,804]" captionStart="FIGURE 5" captionStartId="6.[151,250,1952,1975]" captionTargetBox="[182,1385,193,1895]" captionTargetId="figure@6.[182,1387,193,1897]" captionTargetPageId="6" captionText="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
|
||
b); ambulatory legs creamish with random faint blotches; outer surface of major cheliped distinctly yellow (
|
||
<figureCitation id="132D2A4FFFBAB241FEBEFC96FE49BE1E" box="[365,438,816,840]" captionStart="FIGURE 5" captionStartId="6.[151,250,1952,1975]" captionTargetBox="[182,1385,193,1895]" captionTargetId="figure@6.[182,1387,193,1897]" captionTargetPageId="6" captionText="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
|
||
b; might still be observable after alcohol preservation). Juveniles displaying a more conspicuous pattern with dark, sinuous markings on a pale yellowing background, on both dorsal surface of carapace and ambulatory leg meri (
|
||
<figureCitation id="132D2A4FFFBAB241FDF2FCDEFD9BBEC6" box="[545,612,888,912]" captionStart="FIGURE 5" captionStartId="6.[151,250,1952,1975]" captionTargetBox="[182,1385,193,1895]" captionTargetId="figure@6.[182,1387,193,1897]" captionTargetPageId="6" captionText="FIGURE 5. Life coloration of O. mortoni: a, dorsal view, adult; b, dorsal view, juvenile. c, frontal view, adult. Scale bars represent 1 cm." httpUri="https://zenodo.org/record/213898/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
|
||
c). Cheliped palm also yellowish, with whitish fingers.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFBAB241FF14FC3AFB69BE8E" pageId="8" pageNumber="79" type="distribution">
|
||
<paragraph id="8BA936CAFFBAB241FF14FC3AFB69BE8E" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
<emphasis id="B962EAD8FFBAB241FF14FC3AFE9FBEE3" bold="true" box="[199,352,924,949]" pageId="8" pageNumber="79">Distribution.</emphasis>
|
||
<collectingCountry id="F301765AFFBAB241FEBAFC3BFE12BEE3" box="[361,493,925,949]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
(
|
||
<typeStatus id="54AD8868FFBAB241FE2DFC3AFDCFBEE2" box="[510,560,924,948]" pageId="8" pageNumber="79">type</typeStatus>
|
||
locality), Guangdong, Guangxi, Hainan Island (
|
||
<bibRefCitation id="EF874B3BFFBAB241FB85FC3AFAFABEE2" author="Dai" box="[1110,1285,924,949]" pageId="8" pageNumber="79" refString="Dai, A. - Y., Song, Y. - Z & Yang, S. - L. (1985) A study on Ocypode from China. Acta Zootaxonomica Sinica, 10, 370 - 378. [in Chinese with English abstract]" type="journal article" year="1985">
|
||
Dai
|
||
<emphasis id="B962EAD8FFBAB241FB5BFC38FB44BEE3" box="[1160,1211,925,949]" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1985
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B3BFFBAB241FAC1FC3AFF2DBE8E" author="Dai" pageId="8" pageNumber="79" refString="Dai, A. - Y. & Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 682 pp." type="book" year="1991">Dai & Yang 1991</bibRefCitation>
|
||
), and the Pacific coast of Shikoku,
|
||
<collectingCountry id="F301765AFFBAB241FD8EFC67FD62BE8E" box="[605,669,961,984]" name="Japan" pageId="8" pageNumber="79">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FD7EFC66FCADBE8E" author="Sakai" box="[685,850,960,984]" pageId="8" pageNumber="79" refString="Sakai, K. (2000) On the occurrence of three species of crabs on Shikoku Island, Japan, and a new species, Pinnotheres taichungae nov. spec., from Taiwan (Decapoda, Brachyura). Crustaceana, 73, 1155 - 1162." type="journal article" year="2000">K. Sakai 2000</bibRefCitation>
|
||
). See also “Remarks” below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFBAB240FF14FC43FCDFBC99" lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79" type="discussion">
|
||
<paragraph id="8BA936CAFFBAB241FF14FC43FCF2B982" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
<emphasis id="B962EAD8FFBAB241FF14FC43FEC4BEAB" bold="true" box="[199,315,997,1021]" pageId="8" pageNumber="79">Remarks.</emphasis>
|
||
As suggested in previous works,
|
||
<taxonomicName id="4C164D49FFBAB241FD6DFC43FCC4BEAB" box="[702,827,997,1021]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FD6DFC43FCC4BEAB" box="[702,827,997,1021]" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
morphologically resembles
|
||
<taxonomicName id="4C164D49FFBAB241FBADFC43FB0ABEAB" box="[1150,1269,997,1021]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fabricii">
|
||
<emphasis id="B962EAD8FFBAB241FBADFC43FB0ABEAB" box="[1150,1269,997,1021]" italics="true" pageId="8" pageNumber="79">O. fabricii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFBAB241FAFCFC43FEBFB976" authority="George 1982" authorityName="George" authorityYear="1982" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="nobilii">
|
||
<emphasis id="B962EAD8FFBAB241FAFCFC43FA63BEAB" box="[1327,1436,997,1021]" italics="true" pageId="8" pageNumber="79">O. nobilii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FF4DFBAEFEC7B976" author="George" box="[158,312,1032,1056]" pageId="8" pageNumber="79" refString="George, R. W. (1982) The distribution and evolution of the ghost crabs (Ocypode spp.) of Hong Kong with a description of a new species. In: Morton, B. S. & Tseng, C. K. (Eds), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 185 - 194." type="book chapter" year="1982">George 1982</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, which all develop short ocular stylets above the cornea.
|
||
<taxonomicName id="4C164D49FFBAB241FC33FBAFFB58B977" box="[992,1191,1033,1057]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FC33FBAFFB58B977" box="[992,1191,1033,1057]" italics="true" pageId="8" pageNumber="79">Ocypode mortoni</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from the others based on the number of striations along the stridulating ridge on the inner surface of the major cheliped palm: up to 54 for
|
||
<taxonomicName id="4C164D49FFBAB241FE06FBF7FDAEB93F" box="[469,593,1105,1129]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FE06FBF7FDAEB93F" box="[469,593,1105,1129]" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
, 120 for
|
||
<taxonomicName id="4C164D49FFBAB241FD68FBF7FCD7B93F" box="[699,808,1105,1129]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="nobilii">
|
||
<emphasis id="B962EAD8FFBAB241FD68FBF7FCD7B93F" box="[699,808,1105,1129]" italics="true" pageId="8" pageNumber="79">O. nobilii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<quantity id="4CEE9B2FFFBAB241FCB2FBF6FC1EB93E" box="[865,993,1104,1128]" metricMagnitude="0" metricUnit="m" metricValue="3.1623" metricValueMax="3.5814" metricValueMin="2.7432" pageId="8" pageNumber="79" unit="in" value="124.5" valueMax="141.0" valueMin="108.0">108–141 in</quantity>
|
||
<taxonomicName id="4C164D49FFBAB241FC3AFBF7FBA1B93F" box="[1001,1118,1105,1129]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="fabricii">
|
||
<emphasis id="B962EAD8FFBAB241FC3AFBF7FBA1B93F" box="[1001,1118,1105,1129]" italics="true" pageId="8" pageNumber="79">O. fabricii</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF874B3BFFBAB241FB4FFBF6FA68B93E" author="George" box="[1180,1431,1104,1128]" pageId="8" pageNumber="79" refString="George, R. W. & Knott, M. E. (1965) The Ocypode ghost crabs of Western Australia. Journal of the Royal Society of Western Australia, 48, 15 - 21." type="journal article" year="1965">George & Knott 1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B3BFFBAB241FF44FBD2FECFB9DA" author="George" box="[151,304,1140,1164]" pageId="8" pageNumber="79" refString="George, R. W. (1982) The distribution and evolution of the ghost crabs (Ocypode spp.) of Hong Kong with a description of a new species. In: Morton, B. S. & Tseng, C. K. (Eds), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 185 - 194." type="book chapter" year="1982">George 1982</bibRefCitation>
|
||
). In our specimens, there were 44–65 striations on the stridulating ridges which conforms to
|
||
<taxonomicName id="4C164D49FFBAB241FA52FBD3FF0FB9E7" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FA52FBD3FF0FB9E7" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="132D2A4FFFBAB241FEFFFB3EFE92B9E6" box="[300,365,1176,1200]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="8" pageNumber="79">Fig. 3</figureCitation>
|
||
c, i). G1 morphology of our specimens, with two chitinous beak-like structures at the distal tip, also matches that of the original description (see
|
||
<figureCitation id="132D2A4FFFBAB241FD6FFB1AFCFFB982" box="[700,768,1212,1236]" captionStart="FIGURE 4" captionStartId="5.[151,250,1963,1986]" captionTargetBox="[246,1338,320,1931]" captionTargetId="figure@5.[246,1338,317,1931]" captionTargetPageId="5" captionText="FIGURE 4. Morphology of male first pleopod (G 1) of O. mortoni: a, ventral view of left G 1; b, tip of left G 1, ventral view; c, dorsal view of right G 1; d, G 1 of type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213897/files/figure.png" pageId="8" pageNumber="79">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB241FF14FB46FB8BBB4E" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
<bibRefCitation id="EF874B3BFFBAB241FF14FB46FEAEB9AE" author="Shen" box="[199,337,1248,1272]" pageId="8" pageNumber="79" refString="Shen, C. - J. (1940) The brachyuran fauna of Hong Kong. Hong Kong Fisheries Research Station Journal, 1, 211 - 242." type="journal article" year="1940">Shen (1940)</bibRefCitation>
|
||
recorded
|
||
<taxonomicName id="4C164D49FFBAB241FE17FB47FD5CB9AF" box="[452,675,1249,1273]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBAB241FE17FB47FD5CB9AF" box="[452,675,1249,1273]" italics="true" pageId="8" pageNumber="79">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D49FFBAB241FD61FB47FCA6B9AF" box="[690,857,1249,1273]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="cordimanus">
|
||
<emphasis id="B962EAD8FFBAB241FD61FB47FCA6B9AF" box="[690,857,1249,1273]" italics="true" pageId="8" pageNumber="79">O. cordimanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFBAB241FC42FB47FBDFB9AF" box="[913,1056,1249,1273]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FC42FB47FBDFB9AF" box="[913,1056,1249,1273]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
in his review of the brachyura of
|
||
<collectingCountry id="F301765AFFBAB241FF44FAA3FEDDB84B" box="[151,290,1285,1309]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
, based on his own collections, materials from other institutes and records in the earlier literature (
|
||
<bibRefCitation id="EF874B3BFFBAB241FF4DFA8EFEB3B816" author="Stimpson" box="[158,332,1320,1344]" pageId="8" pageNumber="79" refString="Stimpson, W. (1858) Crustacea Ocypodoidea. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit, Pars 5. Proceedings of the Academia of Natural Science, Philadelphia, 10, 93 - 110." type="journal article" year="1858">Stimpson 1858</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF874B3BFFBAB241FE8AFA8EFE6BB816" author="Stimpson" box="[345,404,1320,1344]" pageId="8" pageNumber="79" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49, 1 - 240." type="journal article" year="1907">1907</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B3BFFBAB241FE72FA8EFDDEB816" author="Balss" box="[417,545,1320,1344]" pageId="8" pageNumber="79" refString="Balss, H. (1922) Ostasiatische Decapoden. IV. Die Brachyrhynchen (Cancridea). Archiv fur Naturgeschichte, 88, 94 - 166." type="journal article" year="1922">Balss 1922</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B3BFFBAB241FDFDFA8EFD5FB816" author="Gee" box="[558,672,1320,1344]" pageId="8" pageNumber="79" refString="Gee, N. G., 1926. Tentative list of Chinese Decapod Crustacea. Lingnaam Agricultural Review, 3, 156 - 166. [dated 1925]" type="journal article" year="1926">Gee 1926</bibRefCitation>
|
||
). The presence of
|
||
<taxonomicName id="4C164D49FFBAB241FCA0FA8FFBADB817" box="[883,1106,1321,1345]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBAB241FCA0FA8FFBADB817" box="[883,1106,1321,1345]" italics="true" pageId="8" pageNumber="79">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
has long been validated (see
|
||
<bibRefCitation id="EF874B3BFFBAB241FF44FAEBFE5BB832" author="Morton" box="[151,420,1356,1380]" pageId="8" pageNumber="79" refString="Morton, B. & Morton, J. (1983) The Sea Shore Ecology of Hong Kong. Hong Kong University Press, Hong Kong. 350 pp." type="book" year="1983">Morton & Morton 1983</bibRefCitation>
|
||
), however the records of
|
||
<taxonomicName id="4C164D49FFBAB241FD6CFAEBFC9BB833" box="[703,868,1357,1381]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="cordimanus">
|
||
<emphasis id="B962EAD8FFBAB241FD6CFAEBFC9BB833" box="[703,868,1357,1381]" italics="true" pageId="8" pageNumber="79">O. cordimanus</emphasis>
|
||
</taxonomicName>
|
||
has been shown to represent
|
||
<taxonomicName id="4C164D49FFBAB241FB7FFAEBFEFEB8DE" authority="Huang et al. 1998" authorityName="Huang et al." authorityYear="1998" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B962EAD8FFBAB241FB7FFAEBFAD9B833" box="[1196,1318,1357,1381]" italics="true" pageId="8" pageNumber="79">O. sinensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FAE6FAEAFF06B8DE" author="Huang" pageId="8" pageNumber="79" refString="Huang, J. - F., Yang, S. - L. & Ng, P. K. L. (1998) Notes on the taxonomy and distribution of the two closely related species of ghost crabs, Ocypode sinensis and O. cordimanus (Decapoda, Brachyura, Ocypodidae). Crustaceana, 71, 942 - 954." type="journal article" year="1998">
|
||
Huang
|
||
<emphasis id="B962EAD8FFBAB241FA5BFAE8FF52B8DF" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1998
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The last species
|
||
<taxonomicName id="4C164D49FFBAB241FE1CFAD7FDA0B8DF" box="[463,607,1393,1417]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FE1CFAD7FDA0B8DF" box="[463,607,1393,1417]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
, appears to be a temperate species (
|
||
<bibRefCitation id="EF874B3BFFBAB241FBD0FAD6FB49B8DE" author="Dai" box="[1027,1206,1392,1417]" pageId="8" pageNumber="79" refString="Dai, A. - Y., Song, Y. - Z & Yang, S. - L. (1985) A study on Ocypode from China. Acta Zootaxonomica Sinica, 10, 370 - 378. [in Chinese with English abstract]" type="journal article" year="1985">
|
||
Dai
|
||
<emphasis id="B962EAD8FFBAB241FBE5FAD4FB95B8DF" box="[1078,1130,1393,1417]" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1985
|
||
</bibRefCitation>
|
||
), and reported from
|
||
<collectingCountry id="F301765AFFBAB241FF44FA33FF28B8FA" box="[151,215,1429,1452]" name="Japan" pageId="8" pageNumber="79">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FF35FA32FE7BB8FA" author="Sakai" box="[230,388,1428,1452]" pageId="8" pageNumber="79" refString="Sakai, T. (1939) Studies on the Crabs of Japan. IV. Brachygnatha, Brachyrhyncha. Yokendo, Tokyo, pp. 365 - 741." type="book chapter" year="1939">T. Sakai 1939</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F301765AFFBAB241FE44FA33FE23B8FA" box="[407,476,1429,1452]" name="South Korea" pageId="8" pageNumber="79">Korea</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FE39FA32FDA0B8FA" author="Kim" box="[490,607,1428,1452]" pageId="8" pageNumber="79" refString="Kim, H. S. (1973) Anomura-Brachyura. In: Illustrated Encyclopedia of Fauna & Flora of Korea, 14. Samhwa, Seoul, pp. 589 - 682 [in Korean]" type="book chapter" year="1973">Kim 1973</bibRefCitation>
|
||
), North
|
||
<collectingCountry id="F301765AFFBAB241FD68FA32FD01B8FA" box="[699,766,1428,1452]" name="China" pageId="8" pageNumber="79">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FCDFFA32FC7AB8FA" author="Shen" box="[780,901,1428,1452]" pageId="8" pageNumber="79" refString="Shen, C. - J. (1932) The brachyuran Crustacea of North China. Zoologia Sinica, Series A, 9, 1 - 300." type="journal article" year="1932">Shen 1932</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="F301765AFFBAB241FC10FA32FBEAB8FA" box="[963,1045,1428,1452]" name="Taiwan" pageId="8" pageNumber="79">Taiwan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FBF0FA32FB0FB8FA" author="Huang" box="[1059,1264,1428,1453]" pageId="8" pageNumber="79" refString="Huang, J. - F., Yu, H. - P. & Takeda, M. (1992) A review of the ocypodid and mictyrid crabs (Crustacea: Decapoda: Brachyura) in Taiwan. Bulletin of the Institute of Zoology, Academia Sinica, 31, 141 - 161." type="journal article" year="1992">
|
||
Huang
|
||
<emphasis id="B962EAD8FFBAB241FBA5FA30FB59B8FB" box="[1142,1190,1429,1453]" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1992
|
||
</bibRefCitation>
|
||
). Records from the coasts of South
|
||
<collectingCountry id="F301765AFFBAB241FEA2FA1EFE4BB886" box="[369,436,1464,1488]" name="China" pageId="8" pageNumber="79">China</collectingCountry>
|
||
(e.g.
|
||
<bibRefCitation id="EF874B3BFFBAB241FE20FA1EFD93B886" author="Shen" box="[499,620,1464,1488]" pageId="8" pageNumber="79" refString="Shen, C. - J. (1936) On a collection of brachyuran Decapoda from Hainan Island with descriptions of three new species. Chinese Journal of Zoology, 11, 63 - 80." type="journal article" year="1936">Shen 1936</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF874B3BFFBAB241FDA4FA1EFD4CB886" author="Shen" box="[631,691,1464,1488]" pageId="8" pageNumber="79" refString="Shen, C. - J. (1940) The brachyuran fauna of Hong Kong. Hong Kong Fisheries Research Station Journal, 1, 211 - 242." type="journal article" year="1940">1940</bibRefCitation>
|
||
), thus, represents the southern limits for
|
||
<taxonomicName id="4C164D49FFBAB241FBABFA1FFAFAB887" box="[1144,1285,1465,1489]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FBABFA1FFAFAB887" box="[1144,1285,1465,1489]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="EF874B3BFFBAB241FAA0FA1EFEEDB8A2" author="Dai" pageId="8" pageNumber="79" refString="Dai, A., Yang, S., Song, Y. & Chen, G. (1986) Crabs of the China Seas. China Ocean Press, Beijing, 568 pp. [in Chinese with English abstract]" type="book" year="1986">
|
||
Dai
|
||
<emphasis id="B962EAD8FFBAB241FF44FA78FF36B8A3" box="[151,201,1501,1525]" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1986
|
||
</bibRefCitation>
|
||
; Dai 1991;
|
||
<bibRefCitation id="EF874B3BFFBAB241FE45FA7AFD9EB8A2" author="Dai" box="[406,609,1500,1525]" pageId="8" pageNumber="79" refString="Dai, A. - Y. & Yang, S. - L. (1991) Crabs of the China Seas. China Ocean Press, Beijing, 682 pp." type="book" year="1991">Dai & Yang 1991</bibRefCitation>
|
||
). However, studies on the genus from
|
||
<collectingCountry id="F301765AFFBAB241FBC7FA7AFBA8B8A2" box="[1044,1111,1500,1524]" name="China" pageId="8" pageNumber="79">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF874B3BFFBAB241FBB5FA7AFAEEB8A2" author="Dai" box="[1126,1297,1500,1525]" pageId="8" pageNumber="79" refString="Dai, A. - Y., Song, Y. - Z & Yang, S. - L. (1985) A study on Ocypode from China. Acta Zootaxonomica Sinica, 10, 370 - 378. [in Chinese with English abstract]" type="journal article" year="1985">
|
||
Dai
|
||
<emphasis id="B962EAD8FFBAB241FB4BFA78FB35B8A3" box="[1176,1226,1501,1525]" italics="true" pageId="8" pageNumber="79">et al</emphasis>
|
||
. 1985
|
||
</bibRefCitation>
|
||
) recognized material from
|
||
<collectingCountry id="F301765AFFBAB241FEE4F9A7FE45BB4F" box="[311,442,1537,1561]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
as
|
||
<taxonomicName id="4C164D49FFBAB241FE33F9A7FDA4BB4F" box="[480,603,1537,1561]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FE33F9A7FDA4BB4F" box="[480,603,1537,1561]" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
(then as
|
||
<taxonomicName id="4C164D49FFBAB241FD13F9A7FCA4BB4F" box="[704,859,1537,1561]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="macrocera">
|
||
<emphasis id="B962EAD8FFBAB241FD13F9A7FCA4BB4F" box="[704,859,1537,1561]" italics="true" pageId="8" pageNumber="79">O. macrocera</emphasis>
|
||
</taxonomicName>
|
||
) instead of
|
||
<taxonomicName id="4C164D49FFBAB241FC33F9A7FB91BB4F" box="[992,1134,1537,1561]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FC33F9A7FB91BB4F" box="[992,1134,1537,1561]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB241FF14F982FCBFBAD6" blockId="8.[151,1436,151,2030]" pageId="8" pageNumber="79">
|
||
Our molecular phylogenetic analyses (
|
||
<figureCitation id="132D2A4FFFBAB241FDA7F982FD47BB6A" box="[628,696,1572,1596]" captionStart="FIGURE 9" captionStartId="11.[151,250,1067,1090]" captionTargetBox="[186,1402,186,1045]" captionTargetId="figure@11.[184,1412,167,1059]" captionTargetPageId="11" captionText="FIGURE 9. Bayesian Inference (BI) tree showing phylogenetic relationships of East Asian Ocypode spp., based on COI gene. Additional support values of BI, maximum likelihood (ML) maximum parsimony (MP) analyses printed at the nodes. For details of haplotypes see Table 1, with species and localities behind. Black bars represent “ ridge-bearing ” species, and gray bars represent “ ridge-less ” species (see Discussion)." httpUri="https://zenodo.org/record/213902/files/figure.png" pageId="8" pageNumber="79">Fig. 9</figureCitation>
|
||
), show
|
||
<taxonomicName id="4C164D49FFBAB241FCDDF983FC64BB6B" box="[782,923,1573,1597]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FCDDF983FC64BB6B" box="[782,923,1573,1597]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFBAB241FC02F983FBB3BB6B" box="[977,1100,1573,1597]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FC02F983FBB3BB6B" box="[977,1100,1573,1597]" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
to be closely related, which is not surprising as they are very similar morphologically except for the presence of ocular stylets in
|
||
<taxonomicName id="4C164D49FFBAB241FACEF9EFFA63BB37" box="[1309,1436,1609,1633]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FACEF9EFFA63BB37" box="[1309,1436,1609,1633]" italics="true" pageId="8" pageNumber="79">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
adults. It is very difficult to separate juveniles or younger individuals of the two species. As for the
|
||
<collectingCountry id="F301765AFFBAB241FAC5F9CBFA63BBD3" box="[1302,1436,1645,1669]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
records of “
|
||
<taxonomicName id="4C164D49FFBAB241FEF7F937FE49BBFF" box="[292,438,1681,1705]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FEF7F937FE49BBFF" box="[292,438,1681,1705]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
” by
|
||
<bibRefCitation id="EF874B3BFFBAB241FE26F936FD79BBFE" author="Shen" box="[501,646,1680,1704]" pageId="8" pageNumber="79" refString="Shen, C. - J. (1940) The brachyuran fauna of Hong Kong. Hong Kong Fisheries Research Station Journal, 1, 211 - 242." type="journal article" year="1940">Shen (1940)</bibRefCitation>
|
||
, all four sites where he collected material (Shatin, Ngau Chi Wan, Repulse Bay and Stanley) have been altered or urbanized over the last 70 years, and no further records have been found since then (see also
|
||
<bibRefCitation id="EF874B3BFFBAB241FE12F97EFDA9BBA6" author="George" box="[449,598,1752,1776]" pageId="8" pageNumber="79" refString="George, R. W. (1982) The distribution and evolution of the ghost crabs (Ocypode spp.) of Hong Kong with a description of a new species. In: Morton, B. S. & Tseng, C. K. (Eds), Proceedings of the First International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China. Hong Kong University Press, Hong Kong, pp. 185 - 194." type="book chapter" year="1982">George 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF874B3BFFBAB241FDB1F97FFC8DBBA6" author="Morton" box="[610,882,1752,1776]" pageId="8" pageNumber="79" refString="Morton, B. & Morton, J. (1983) The Sea Shore Ecology of Hong Kong. Hong Kong University Press, Hong Kong. 350 pp." type="book" year="1983">Morton & Morton 1983</bibRefCitation>
|
||
). Extensive searches performed in this study also failed to acquire any material of this species. While it is tempting to regard the earlier records of
|
||
<taxonomicName id="4C164D49FFBAB241FB3EF95BFA82BA43" box="[1261,1405,1789,1813]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FB3EF95BFA82BA43" box="[1261,1405,1789,1813]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
as probable misidentifications, the past presence and subsequent local extinction is nevertheless still possible. As
|
||
<collectingCountry id="F301765AFFBAB241FF44F8E3FEE1BA0B" box="[151,286,1861,1885]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
is within the range of the known southern distribution of
|
||
<taxonomicName id="4C164D49FFBAB241FC02F8E3FB9DBA0B" box="[977,1122,1861,1885]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFBAB241FC02F8E3FB9DBA0B" box="[977,1122,1861,1885]" italics="true" pageId="8" pageNumber="79">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
we tentatively continue to include it as known from there, though with some hesitation.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBAB240FF14F82BFCDFBC99" blockId="8.[151,1436,151,2030]" lastBlockId="9.[151,1436,151,832]" lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79">
|
||
For records in the East Asian region, aside from southern
|
||
<collectingCountry id="F301765AFFBAB241FC9DF82AFC6EBAF2" box="[846,913,1932,1956]" name="China" pageId="8" pageNumber="79">China</collectingCountry>
|
||
(including
|
||
<collectingCountry id="F301765AFFBAB241FBC1F82BFB66BAF2" box="[1042,1177,1932,1957]" name="Hong Kong" pageId="8" pageNumber="79">Hong Kong</collectingCountry>
|
||
),
|
||
<taxonomicName id="4C164D49FFBAB241FB7FF82BFA8FBAF3" box="[1196,1392,1933,1957]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="79" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBAB241FB7FF82BFA8FBAF3" box="[1196,1392,1933,1957]" italics="true" pageId="8" pageNumber="79">Ocypode mortoni</emphasis>
|
||
</taxonomicName>
|
||
has only been recorded on the Pacific coast of Shikoku,
|
||
<collectingCountry id="F301765AFFBAB241FD23F817FCCFBA9E" box="[752,816,1969,1992]" name="Japan" pageId="8" pageNumber="79">Japan</collectingCountry>
|
||
(see
|
||
<bibRefCitation id="EF874B3BFFBAB241FCBCF817FBE8BA9E" author="Sakai" box="[879,1047,1968,1992]" pageId="8" pageNumber="79" refString="Sakai, K. (2000) On the occurrence of three species of crabs on Shikoku Island, Japan, and a new species, Pinnotheres taichungae nov. spec., from Taiwan (Decapoda, Brachyura). Crustaceana, 73, 1155 - 1162." type="journal article" year="2000">K. Sakai 2000</bibRefCitation>
|
||
). The illustrations of the Japanese material agrees with our specimens, in terms of the number of striae along the stridulatory ridge (Shikoku specimens:
|
||
<quantity id="4CEE9B2FFFBBB240FEC9FF31FEACBDF9" box="[282,339,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="9" pageNumber="80" unit="in" value="60.0">60 in</quantity>
|
||
male and
|
||
<quantity id="4CEE9B2FFFBBB240FE16FF31FE01BDF9" box="[453,510,151,175]" metricMagnitude="0" metricUnit="m" metricValue="1.3208" pageId="9" pageNumber="80" unit="in" value="52.0">52 in</quantity>
|
||
female;
|
||
<collectingCountry id="F301765AFFBBB240FD8CFF3EFD1DBDE6" box="[607,738,152,176]" name="Hong Kong" pageId="9" pageNumber="80">Hong Kong</collectingCountry>
|
||
specimens: 44–65), and in the form of the zygocardiac ossicle (
|
||
<figureCitation id="132D2A4FFFBBB240FF4CFF1AFF1ABD82" box="[159,229,188,212]" captionStart="FIGURE 3" captionStartId="4.[151,250,1535,1558]" captionTargetBox="[157,1433,635,1534]" captionTargetId="figure@4.[151,1436,601,1542]" captionTargetPageId="4" captionText="FIGURE 3. Morphology of O. mortoni: a, dorsal surface of carapace; b, outer surface of major cheliped palm; c; inner surface of major cheliped palm, showing stridulating ridge; d, outer surface of minor cheliped; e, male abdomen; f, female abdomen; g, zygocardiac ossicle; h, urocardiac ossicle; i, inner surface of major cheliped palm of the type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213896/files/figure.png" pageId="9" pageNumber="80">Fig. 3</figureCitation>
|
||
h). However, the distal tip of the G
|
||
<quantity id="4CEE9B2FFFBBB240FD5AFF1AFD4BBD82" box="[649,692,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="80" unit="in" value="1.0">1 in</quantity>
|
||
material from
|
||
<collectingCountry id="F301765AFFBBB240FCB5FF1BFC0FBD83" box="[870,1008,189,213]" name="Hong Kong" pageId="9" pageNumber="80">Hong Kong</collectingCountry>
|
||
, when viewed ventrally, ends with a pronounced chitinous bi-lobed, pointed, beak-like structure, with the upper lobe larger than the lower; and fine setae on the “neck” of the curvature beneath (see
|
||
<figureCitation id="132D2A4FFFBBB240FD1EFEA2FCEFBC4A" box="[717,784,260,284]" captionStart="FIGURE 4" captionStartId="5.[151,250,1963,1986]" captionTargetBox="[246,1338,320,1931]" captionTargetId="figure@5.[246,1338,317,1931]" captionTargetPageId="5" captionText="FIGURE 4. Morphology of male first pleopod (G 1) of O. mortoni: a, ventral view of left G 1; b, tip of left G 1, ventral view; c, dorsal view of right G 1; d, G 1 of type specimen, redrawn from George (1982)." httpUri="https://zenodo.org/record/213897/files/figure.png" pageId="9" pageNumber="80">Fig. 4</figureCitation>
|
||
a–b, d). Both characters are somewhat wanting in Sakai’s (2000) illustrations (fig. 3d in
|
||
<bibRefCitation id="EF874B3BFFBBB240FE3AFE8EFD72BC69" author="Sakai" box="[489,653,295,319]" pageId="9" pageNumber="80" refString="Sakai, K. (2000) On the occurrence of three species of crabs on Shikoku Island, Japan, and a new species, Pinnotheres taichungae nov. spec., from Taiwan (Decapoda, Brachyura). Crustaceana, 73, 1155 - 1162." type="journal article" year="2000">K. Sakai 2000</bibRefCitation>
|
||
). The importance of these slight variations is uncertain, and we would opt to consider both the
|
||
<collectingCountry id="F301765AFFBBB240FE74FEEBFDD4BC33" box="[423,555,333,357]" name="Hong Kong" pageId="9" pageNumber="80">Hong Kong</collectingCountry>
|
||
and Japanese records conspecific, at least for now. While
|
||
<taxonomicName id="4C164D49FFBBB240FB69FEEBFACABC32" box="[1210,1333,333,356]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FB69FEEBFACABC32" box="[1210,1333,333,356]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
could be expected to occur in
|
||
<collectingCountry id="F301765AFFBBB240FE53FEC9FE2DBCD1" box="[384,466,367,391]" name="Taiwan" pageId="9" pageNumber="80">Taiwan</collectingCountry>
|
||
and adjacent areas, no specimens have been collected despite numerous efforts to find it, and none are present in museum collections. The biogeography of the species remains unclear, and needs further investigation based on more extensive targeted collecting.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C6541FFBBB242FF14FE7DFBE9BB91" lastPageId="11" lastPageNumber="82" pageId="9" pageNumber="80" type="distribution">
|
||
<paragraph id="8BA936CAFFBBB240FF14FE7DFC8EBF36" blockId="9.[151,1436,151,832]" pageId="9" pageNumber="80">
|
||
<emphasis id="B962EAD8FFBBB240FF14FE7DFCADBCA2" bold="true" box="[199,850,475,500]" pageId="9" pageNumber="80">
|
||
Distribution pattern of
|
||
<taxonomicName id="4C164D49FFBBB240FE3FFE7DFDAFBCA2" box="[492,592,475,500]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFBBB240FE3FFE7DFDAFBCA2" bold="true" box="[492,592,475,500]" italics="true" pageId="9" pageNumber="80">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
spp. in
|
||
<collectingCountry id="F301765AFFBBB240FD6FFE7AFCB3BCA2" box="[700,844,476,500]" name="Hong Kong" pageId="9" pageNumber="80">Hong Kong</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
From the seven sandy shores surveyed,
|
||
<taxonomicName id="4C164D49FFBBB240FAEBFE7AFEACBF4E" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FAEBFE7AFEACBF4E" italics="true" pageId="9" pageNumber="80">Ocypode ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
was collected in all sites while
|
||
<taxonomicName id="4C164D49FFBBB240FD67FDA6FCD0BF4E" box="[692,815,512,536]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FD67FDA6FCD0BF4E" box="[692,815,512,536]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
was only limited at Sai Wan, Tong Fuk and Bui O (
|
||
<figureCitation id="132D2A4FFFBBB240FABCFE59FF5BBF6A" captionStart="FIGURE 1" captionStartId="1.[151,250,1910,1933]" captionTargetBox="[175,1406,1003,1892]" captionTargetId="figure@1.[168,1412,1003,1901]" captionTargetPageId="1" captionText="FIGURE 1. Map of Hong Kong showing sandy shore study sites. Squares indicate the sites visited by George (1982) and circles indicate the sites visited in the present study. Closed circle / square indicates the presence of O. mortoni. Empty circle / square indicated absence of O. mortoni." httpUri="https://zenodo.org/record/213894/files/figure.png" pageId="9" pageNumber="80">Fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="4C164D49FFBBB240FF6EFD82FE7EBF6A" box="[189,385,548,572]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FF6EFD82FE7EBF6A" box="[189,385,548,572]" italics="true" pageId="9" pageNumber="80">Ocypode mortoni</emphasis>
|
||
</taxonomicName>
|
||
was found at the same tidal levels as
|
||
<taxonomicName id="4C164D49FFBBB240FCFDFD83FBF2BF6A" box="[814,1037,548,572]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FCFDFD83FBF2BF6A" box="[814,1037,548,572]" italics="true" pageId="9" pageNumber="80">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
(2.0–
|
||
<quantity id="4CEE9B2FFFBBB240FB83FD82FB6EBF6A" box="[1104,1169,548,572]" metricMagnitude="0" metricUnit="m" metricValue="2.3" pageId="9" pageNumber="80" unit="m" value="2.3">2.3 m</quantity>
|
||
above C.D.) but not on higher shores, where the zone is solely dominated by
|
||
<taxonomicName id="4C164D49FFBBB240FD3CFDEEFC95BF36" box="[751,874,584,608]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B962EAD8FFBBB240FD3CFDEEFC95BF36" box="[751,874,584,608]" italics="true" pageId="9" pageNumber="80">O. sinensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFBBB240FF14FDCDFE21BE16" blockId="9.[151,1436,151,832]" pageId="9" pageNumber="80">
|
||
<emphasis id="B962EAD8FFBBB240FF14FDCDFA63BFD2" bold="true" box="[199,1436,619,644]" pageId="9" pageNumber="80">
|
||
Vertical distribution pattern at Sai Wan and relative abundance of
|
||
<taxonomicName id="4C164D49FFBBB240FC21FDCDFB8DBFD2" box="[1010,1138,619,644]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FC21FDCDFB8DBFD2" bold="true" box="[1010,1138,619,644]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B962EAD8FFBBB240FB7CFDCDFA63BFD2" bold="true" box="[1199,1436,619,644]" italics="true" pageId="9" pageNumber="80">
|
||
<taxonomicName id="4C164D49FFBBB240FB7CFDCDFA67BFD2" box="[1199,1432,619,644]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">O. ceratophthalmus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</emphasis>
|
||
At Sai Wan, along the transect close to the stream (T1), average density of
|
||
<taxonomicName id="4C164D49FFBBB240FBF5FD36FAF5BFFE" box="[1062,1290,656,680]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FBF5FD36FAF5BFFE" box="[1062,1290,656,680]" italics="true" pageId="9" pageNumber="80">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
reached 2.2 individuals m -2, while for
|
||
<taxonomicName id="4C164D49FFBBB240FE66FD1EFDCFBF99" box="[437,560,696,719]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FE66FD1EFDCFBF99" box="[437,560,696,719]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
only occurred at 0.4 individuals m -2. On the transect further beyond the stream (T2),
|
||
<taxonomicName id="4C164D49FFBBB240FF05FD78FE52BFA3" box="[214,429,733,757]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FF05FD78FE52BFA3" box="[214,429,733,757]" italics="true" pageId="9" pageNumber="80">O ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
maintained an average density of 1.8 individuals m -2, and
|
||
<taxonomicName id="4C164D49FFBBB240FBEBFD78FB4CBFA0" box="[1080,1203,734,758]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FBEBFD78FB4CBFA0" box="[1080,1203,734,758]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
at 0.8 individuals m - 2 (
|
||
<figureCitation id="132D2A4FFFBBB240FF62FCA2FF07BE4A" box="[177,248,772,796]" captionStart="FIGURE 7" captionStartId="9.[151,250,1920,1943]" captionTargetBox="[225,1341,895,1869]" captionTargetId="figure@9.[217,1351,885,1881]" captionTargetPageId="9" captionText="FIGURE 7. Density of O. mortoni and O. ceratophthalmus along T 1 and T 2 in Sai Wan (refer to Y-axis on left hand side); and the total number of crabs collected in the transects at Fan Lau and Bui O along the 30 meter-transects (refer to the Y axis on the right hand side)." httpUri="https://zenodo.org/record/213900/files/figure.png" pageId="9" pageNumber="80">Fig. 7</figureCitation>
|
||
). In Fan Lau and Bui O, the relative abundance of
|
||
<taxonomicName id="4C164D49FFBBB240FC82FCA3FBCDBE4A" box="[849,1074,772,796]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FC82FCA3FBCDBE4A" box="[849,1074,772,796]" italics="true" pageId="9" pageNumber="80">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C164D49FFBBB240FB8FFCA3FB25BE4A" box="[1116,1242,773,796]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FB8FFCA3FB25BE4A" box="[1116,1242,773,796]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
was 96:4%, and 83:17% respectively (
|
||
<figureCitation id="132D2A4FFFBBB240FE58FC8EFE2FBE16" box="[395,464,808,832]" captionStart="FIGURE 7" captionStartId="9.[151,250,1920,1943]" captionTargetBox="[225,1341,895,1869]" captionTargetId="figure@9.[217,1351,885,1881]" captionTargetPageId="9" captionText="FIGURE 7. Density of O. mortoni and O. ceratophthalmus along T 1 and T 2 in Sai Wan (refer to Y-axis on left hand side); and the total number of crabs collected in the transects at Fan Lau and Bui O along the 30 meter-transects (refer to the Y axis on the right hand side)." httpUri="https://zenodo.org/record/213900/files/figure.png" pageId="9" pageNumber="80">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF696642FFBBB240FF44F826FEC2BA85" httpUri="https://zenodo.org/record/213900/files/figure.png" pageId="9" pageNumber="80" targetBox="[225,1341,895,1869]" targetPageId="9">
|
||
<paragraph id="8BA936CAFFBBB240FF44F826FEC2BA85" blockId="9.[151,1436,1920,2003]" pageId="9" pageNumber="80">
|
||
<emphasis id="B962EAD8FFBBB240FF44F826FEEDBAC0" bold="true" box="[151,274,1920,1943]" pageId="9" pageNumber="80">FIGURE 7.</emphasis>
|
||
Density of
|
||
<taxonomicName id="4C164D49FFBBB240FE5AF827FE07BAC0" box="[393,504,1921,1942]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFBBB240FE5AF827FE07BAC0" box="[393,504,1921,1942]" italics="true" pageId="9" pageNumber="80">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFBBB240FDFAF827FD0DBAC0" box="[553,754,1920,1942]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="80" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFBBB240FDFAF827FD0DBAC0" box="[553,754,1920,1942]" italics="true" pageId="9" pageNumber="80">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
along T1 and T2 in Sai Wan (refer to Y-axis on left hand side); and the total number of crabs collected in the transects at Fan Lau and Bui O along the 30 meter-transects (refer to the Y axis on the right hand side).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF696642FFB8B243FF44F8DFFCB8BABA" httpUri="https://zenodo.org/record/213901/files/figure.png" pageId="10" pageNumber="81" targetBox="[198,1402,193,1896]" targetPageId="10">
|
||
<paragraph id="8BA936CAFFB8B243FF44F8DFFCB8BABA" blockId="10.[151,1436,1913,2029]" pageId="10" pageNumber="81">
|
||
<emphasis id="B962EAD8FFB8B243FF44F8DFFEEDBAD9" bold="true" box="[151,274,1913,1936]" pageId="10" pageNumber="81">FIGURE 8.</emphasis>
|
||
Burrow morphology of
|
||
<taxonomicName id="4C164D49FFB8B243FDD9F8DCFD9BBAC6" box="[522,612,1914,1936]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB8B243FDD9F8DCFD9BBAC6" box="[522,612,1914,1936]" italics="true" pageId="10" pageNumber="81">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
spp.: Burrow opening: a–b from Sai Wan (
|
||
<taxonomicName id="4C164D49FFB8B243FBCAF8DDFB1DBAC6" box="[1049,1250,1914,1936]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFB8B243FBCAF8DDFB1DBAC6" box="[1049,1250,1914,1936]" italics="true" pageId="10" pageNumber="81">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C164D49FFB8B243FB26F8DDFA9BBAC6" box="[1269,1380,1915,1936]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB8B243FB26F8DDFA9BBAC6" box="[1269,1380,1915,1936]" italics="true" pageId="10" pageNumber="81">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
were both found in these two types of burrows, see Discussion); c–d from Bui O (
|
||
<taxonomicName id="4C164D49FFB8B243FC72F83CFB95BAF9" box="[929,1130,1945,1967]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFB8B243FC72F83CFB95BAF9" box="[929,1130,1945,1967]" italics="true" pageId="10" pageNumber="81">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C164D49FFB8B243FBACF83CFB11BAF9" box="[1151,1262,1946,1967]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB8B243FBACF83CFB11BAF9" box="[1151,1262,1946,1967]" italics="true" pageId="10" pageNumber="81">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
were both found in these two types of burrows). Note the variation of arrangements of sediment piles around the opening; e, burrow opening of
|
||
<taxonomicName id="4C164D49FFB8B243FF44F871FEF9BABA" box="[151,262,2007,2028]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="81" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB8B243FF44F871FEF9BABA" box="[151,262,2007,2028]" italics="true" pageId="10" pageNumber="81">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
redrawn from George (1982). Scale bars represent 5 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF696642FFB9B242FF44FB8DFD89B9CB" httpUri="https://zenodo.org/record/213902/files/figure.png" pageId="11" pageNumber="82" targetBox="[186,1402,186,1045]" targetPageId="11">
|
||
<paragraph id="8BA936CAFFB9B242FF44FB8DFD89B9CB" blockId="11.[151,1437,1067,1181]" pageId="11" pageNumber="82">
|
||
<emphasis id="B962EAD8FFB9B242FF44FB8DFEECB916" bold="true" box="[151,275,1067,1090]" pageId="11" pageNumber="82">FIGURE 9.</emphasis>
|
||
Bayesian Inference (BI) tree showing phylogenetic relationships of East Asian
|
||
<taxonomicName id="4C164D49FFB9B242FBEEFB8DFB68B917" box="[1085,1175,1067,1089]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FBEEFB8DFB68B917" box="[1085,1175,1067,1089]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
spp., based on COI gene. Additional support values of BI, maximum likelihood (ML) maximum parsimony (MP) analyses printed at the nodes. For details of haplotypes see Table 1, with species and localities behind. Black bars represent “ridge-bearing” species, and gray bars represent “ridge-less” species (see Discussion).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA936CAFFB9B242FF14FB6DFB4DB871" blockId="11.[151,1436,1226,1735]" pageId="11" pageNumber="82">
|
||
<emphasis id="B962EAD8FFB9B242FF14FB6DFE4DB9B5" bold="true" box="[199,434,1226,1251]" pageId="11" pageNumber="82">Molecular analysis.</emphasis>
|
||
From 28 individuals of
|
||
<taxonomicName id="4C164D49FFB9B242FD02FB6CFCCAB9B4" box="[721,821,1226,1250]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FD02FB6CFCCAB9B4" box="[721,821,1226,1250]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
spp., 24 haplotypes (see Table 1) of COI sequences, each of 636 basepairs were obtained. The alignment are AT-rich (in average A: 27.6%, T: 34.3%, G: 17.0%, C: 21.2%), within which 156 sites are variable, and 130 of these are parsimoniously informative.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFB9B242FF14FA94FB26B8E4" blockId="11.[151,1436,1226,1735]" pageId="11" pageNumber="82">
|
||
Pairwise nucleotide divergences (K2P distance) and the mean number of differences of the COI sequences between five East Asian
|
||
<taxonomicName id="4C164D49FFB9B242FE66FAF3FDE6B83B" box="[437,537,1365,1389]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FE66FAF3FDE6B83B" box="[437,537,1365,1389]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
species are displayed in Table 2. Apart from
|
||
<taxonomicName id="4C164D49FFB9B242FBF4FAF0FB31B83B" box="[1063,1230,1365,1389]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="cordimanus">
|
||
<emphasis id="B962EAD8FFB9B242FBF4FAF0FB31B83B" box="[1063,1230,1365,1389]" italics="true" pageId="11" pageNumber="82">O. cordimanus</emphasis>
|
||
</taxonomicName>
|
||
which involved a single haplotype, the remaining 4 species show a certain degree of intraspecific variation. The percentage of bp differences and K2P can be as much as 1.57% (10/636) and 1.60% (for
|
||
<taxonomicName id="4C164D49FFB9B242FC6AFA3DFBCAB8E4" box="[953,1077,1435,1458]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB9B242FC6AFA3DFBCAB8E4" box="[953,1077,1435,1458]" italics="true" pageId="11" pageNumber="82">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
), respectively.
|
||
</paragraph>
|
||
<paragraph id="8BA936CAFFB9B242FF14FA1BFBE9BB91" blockId="11.[151,1436,1226,1735]" pageId="11" pageNumber="82">
|
||
A phylogenetic (
|
||
<collectingCountry id="F301765AFFB9B242FE5AFA1BFE58B883" box="[393,423,1469,1493]" name="Belgium" pageId="11" pageNumber="82">BI</collectingCountry>
|
||
) tree of COI of East Asian
|
||
<taxonomicName id="4C164D49FFB9B242FD27FA1BFCA7B883" box="[756,856,1469,1493]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FD27FA1BFCA7B883" box="[756,856,1469,1493]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
species is shown in
|
||
<figureCitation id="132D2A4FFFB9B242FB9CFA1BFB4CB883" box="[1103,1203,1469,1493]" captionStart="FIGURE 9" captionStartId="11.[151,250,1067,1090]" captionTargetBox="[186,1402,186,1045]" captionTargetId="figure@11.[184,1412,167,1059]" captionTargetPageId="11" captionText="FIGURE 9. Bayesian Inference (BI) tree showing phylogenetic relationships of East Asian Ocypode spp., based on COI gene. Additional support values of BI, maximum likelihood (ML) maximum parsimony (MP) analyses printed at the nodes. For details of haplotypes see Table 1, with species and localities behind. Black bars represent “ ridge-bearing ” species, and gray bars represent “ ridge-less ” species (see Discussion)." httpUri="https://zenodo.org/record/213902/files/figure.png" pageId="11" pageNumber="82">Figure 9</figureCitation>
|
||
, with the additional support values of ML and MP analyses beside the nodes. Results show all the studied
|
||
<taxonomicName id="4C164D49FFB9B242FB50FA79FB18B8A1" box="[1155,1255,1503,1527]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FB50FA79FB18B8A1" box="[1155,1255,1503,1527]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
species formed well-supported, distinct clades at the species level.
|
||
<taxonomicName id="4C164D49FFB9B242FD33F9A4FC5ABB4C" box="[736,933,1538,1562]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB9B242FD33F9A4FC5ABB4C" box="[736,933,1538,1562]" italics="true" pageId="11" pageNumber="82">Ocypode mortoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFB9B242FC0DF9A5FB92BB4C" box="[990,1133,1539,1562]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFB9B242FC0DF9A5FB92BB4C" box="[990,1133,1539,1562]" italics="true" pageId="11" pageNumber="82">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
are genetically the closest related (average K2P distance: 6.25%, see Table 2). Of the 24 haplotype analysed, the East Asian
|
||
<taxonomicName id="4C164D49FFB9B242FB3BF983FAB3BB6B" box="[1256,1356,1573,1597]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962EAD8FFB9B242FB3BF983FAB3BB6B" box="[1256,1356,1573,1597]" italics="true" pageId="11" pageNumber="82">Ocypode</emphasis>
|
||
</taxonomicName>
|
||
can be divided into two main clades with robust clade support: one contains the species with a stridulating ridge on the inner surface of major cheliped, i.e.
|
||
<emphasis id="B962EAD8FFB9B242FD9BF9CDFC92BBD4" box="[584,877,1643,1666]" italics="true" pageId="11" pageNumber="82">
|
||
<taxonomicName id="4C164D49FFB9B242FD9BF9CDFD34BBD4" box="[584,715,1643,1666]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="mortoni">O. mortoni</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C164D49FFB9B242FD08F9CDFC92BBD4" box="[731,877,1643,1666]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="stimpsoni">O. stimpsoni</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C164D49FFB9B242FC7EF9CDFB6FBBD4" box="[941,1168,1642,1666]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFB9B242FC7EF9CDFB6FBBD4" box="[941,1168,1642,1666]" italics="true" pageId="11" pageNumber="82">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
; while the other clade contains species that lack ridges on the inner side of the chelae:
|
||
<taxonomicName id="4C164D49FFB9B242FCADF928FBDABBF3" box="[894,1061,1677,1701]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="cordimanus">
|
||
<emphasis id="B962EAD8FFB9B242FCADF928FBDABBF3" box="[894,1061,1677,1701]" italics="true" pageId="11" pageNumber="82">O. cordimanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFB9B242FBB3F928FB21BBF3" box="[1120,1246,1678,1701]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="sinensis">
|
||
<emphasis id="B962EAD8FFB9B242FBB3F928FB21BBF3" box="[1120,1246,1678,1701]" italics="true" pageId="11" pageNumber="82">O. sinensis</emphasis>
|
||
</taxonomicName>
|
||
. Within the first clade,
|
||
<taxonomicName id="4C164D49FFB9B242FF33F916FE41BB91" box="[224,446,1711,1735]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="ceratophthalmus">
|
||
<emphasis id="B962EAD8FFB9B242FF33F916FE41BB91" box="[224,446,1711,1735]" italics="true" pageId="11" pageNumber="82">O. ceratophthalmus</emphasis>
|
||
</taxonomicName>
|
||
is sister to the
|
||
<taxonomicName id="4C164D49FFB9B242FDB4F916FD1DBB91" box="[615,738,1712,1735]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="mortoni">
|
||
<emphasis id="B962EAD8FFB9B242FDB4F916FD1DBB91" box="[615,738,1712,1735]" italics="true" pageId="11" pageNumber="82">O. mortoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D49FFB9B242FCCBF916FC59BB91" box="[792,934,1712,1735]" class="Malacostraca" family="Ocypodidae" genus="Ocypode" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="82" phylum="Arthropoda" rank="species" species="stimpsoni">
|
||
<emphasis id="B962EAD8FFB9B242FCCBF916FC59BB91" box="[792,934,1712,1735]" italics="true" pageId="11" pageNumber="82">O. stimpsoni</emphasis>
|
||
</taxonomicName>
|
||
subclade.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |