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<document id="C1E63CE0C4E2D8C83586916EC81208E6" ID-DOI="10.5281/zenodo.213903" ID-GBIF-Dataset="ac31e0e8-6809-46b1-a792-0a0b143fd41e" ID-ISSN="1175-5326" ID-Zenodo-Dep="213903" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564814817" checkinUser="plazi" docAuthor="Gonzalez, Victor H., Sepúlveda, Paula A. &amp; Griswold, Terry L." docDate="2012" docId="03BF87B9FF9AFFA3FF59FA07BC78FE81" docLanguage="en" docName="zt03591p078.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heriades (Neotrypetes) tayrona Gonzalez &amp; Griswold 2011" docType="treatment" docVersion="9" lastPageNumber="78" masterDocId="FF86FFC1FF9AFFA0FFCEFF89B800FFD1" masterDocTitle="Taxonomic notes on American Heriades Spinola, 1808 and Leioproctus Smith, 1853 (Hymenoptera: Megachilidae, Colletidae)" masterLastPageNumber="78" masterPageNumber="75" pageNumber="75" updateTime="1698320843763" updateUser="plazi">
<mods:mods id="EDAB870200ECB94AD9E2DCD0E41E68BA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B6F02A203036F846F1AAEEF8C5579DCA">Taxonomic notes on American Heriades Spinola, 1808 and Leioproctus Smith, 1853 (Hymenoptera: Megachilidae, Colletidae)</mods:title>
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<mods:namePart id="266356AA768CB4D6ECAE4EC9F4B84113">Gonzalez, Victor H.</mods:namePart>
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<mods:namePart id="A0AE96E2854FF09F1963924887628CD4">Sepúlveda, Paula A.</mods:namePart>
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<mods:namePart id="B78D7525BCCA7DDA673C055BD1B99F6C">Griswold, Terry L.</mods:namePart>
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<mods:title id="31894D76988F2800431B2B6583B1F24C">Zootaxa</mods:title>
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<mods:date id="3C0C8C05C388D4571BC890628B669B10">2012</mods:date>
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<mods:number id="5B51CCD2915F2068644EA1C27F56AEC0">3591</mods:number>
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<mods:identifier id="25B8E9041E34D2CAFCAE7DB1C8DD706B" type="DOI">10.5281/zenodo.213903</mods:identifier>
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<treatment id="03BF87B9FF9AFFA3FF59FA07BC78FE81" ID-DOI="http://doi.org/10.5281/zenodo.6180108" ID-GBIF-Taxon="119557527" ID-Zenodo-Dep="6180108" LSID="urn:lsid:plazi:treatment:03BF87B9FF9AFFA3FF59FA07BC78FE81" httpUri="http://treatment.plazi.org/id/03BF87B9FF9AFFA3FF59FA07BC78FE81" lastPageId="3" lastPageNumber="78" pageId="0" pageNumber="75">
<subSubSection id="C30C6524FF9AFFA0FF59FA07B934FA17" pageId="0" pageNumber="75" type="nomenclature">
<paragraph id="8BA936AFFF9AFFA0FF59FA07BB2CFA77" blockId="0.[151,812,1422,1478]" box="[151,812,1422,1446]" pageId="0" pageNumber="75">
<heading id="D0E181C3FF9AFFA0FF59FA07BB2CFA77" bold="true" box="[151,812,1422,1446]" fontSize="10" level="2" pageId="0" pageNumber="75" reason="2">
<taxonomicName id="4C164D2CFF9AFFA0FF59FA07BB2CFA77" ID-CoL="8SNLF" authority="Gonzalez &amp; Griswold, 2011" authorityName="Gonzalez &amp; Griswold" authorityYear="2011" box="[151,812,1422,1446]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="tayrona" subGenus="Neotrypetes">
<emphasis id="B962EABDFF9AFFA0FF59FA07BB2CFA77" bold="true" box="[151,812,1422,1446]" pageId="0" pageNumber="75">
<emphasis id="B962EABDFF9AFFA0FF59FA07B8FAFA77" bold="true" box="[151,250,1422,1446]" italics="true" pageId="0" pageNumber="75">Heriades</emphasis>
(
<emphasis id="B962EABDFF9AFFA0FEC4FA07B98BFA77" bold="true" box="[266,395,1422,1446]" italics="true" pageId="0" pageNumber="75">Neotrypetes</emphasis>
)
<emphasis id="B962EABDFF9AFFA0FE54FA07B9EEFA77" bold="true" box="[410,494,1422,1446]" italics="true" pageId="0" pageNumber="75">tayrona</emphasis>
<bibRefCitation id="EF874B5EFF9AFFA0FE3AFA07BB2CFA77" author="Gonzalez" box="[500,812,1422,1446]" pageId="0" pageNumber="75" refString="Gonzalez, V. H. &amp; Griswold, T. (2011) Heriades tayrona n. sp., the first osmiine bee from South America (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society, 84, 255 - 259." type="journal article" year="2011">Gonzalez &amp; Griswold, 2011</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA936AFFF9AFFA0FF59FA26B934FA17" blockId="0.[151,812,1422,1478]" box="[151,308,1455,1478]" pageId="0" pageNumber="75">
(
<figureCitation id="132D2A2AFF9AFFA0FF50FA26B92CFA17" box="[158,300,1455,1478]" captionStart="FIGURES 1 – 14" captionStartId="1.[151,269,1881,1905]" captionTargetBox="[192,1417,219,1829]" captionTargetId="figure@1.[151,1436,194,1857]" captionTargetPageId="1" captionText="FIGURES 1 – 14. Male of Heriades tayrona Gonzalez &amp; Griswold. 1, Lateral habitus. 2, Dorsal habitus. 3, Facial view. 4, Metasoma in ventral view. 5, Seventh tergum. 6 – 11, Third to eighth sterna. 12 – 14, Genital capsule in lateral, dorsal and ventral views." httpUri="https://zenodo.org/record/213904/files/figure.png" pageId="0" pageNumber="75">Figures 1–14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30C6524FF9AFFA0FF59FA66B966F979" pageId="0" pageNumber="75" type="diagnosis">
<paragraph id="8BA936AFFF9AFFA0FF59FA66B966F979" blockId="0.[151,1436,1519,2026]" pageId="0" pageNumber="75">
<emphasis id="B962EABDFF9AFFA0FF59FA66B90CF9D6" bold="true" box="[151,268,1519,1543]" pageId="0" pageNumber="75">Diagnosis.</emphasis>
This species can be easily recognized by the following combination of characters: female clypeus with a narrow (about two-fifths of basal width of clypeus), shallow median emargination on the distal margin delimited laterally by a small tubercle; ocelli very small in both sexes, slightly larger (1.2–1.4 times) than the diameter of punctures in the interocellar area; and basal metasomal terga uniformly punctate, with punctures smaller than those on scutum and separated by a puncture width or less. Male sterna five and six are similar to those in
<taxonomicName id="4C164D2CFF9AFFA0FBE2F9F8BC9DF959" box="[1068,1181,1649,1672]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="0" pageNumber="75" phylum="Arthropoda" rank="species" species="currani">
<emphasis id="B962EABDFF9AFFA0FBE2F9F8BC9DF959" box="[1068,1181,1649,1672]" italics="true" pageId="0" pageNumber="75">H. currani</emphasis>
</taxonomicName>
; it can be distinguished by the larger ocelli.
</paragraph>
</subSubSection>
<subSubSection id="C30C6524FF9AFFA3FF09F938BA29FEDE" lastPageId="3" lastPageNumber="78" pageId="0" pageNumber="75" type="description">
<paragraph id="8BA936AFFF9AFFA0FF09F938B924F83B" blockId="0.[151,1436,1519,2026]" pageId="0" pageNumber="75">
<emphasis id="B962EABDFF9AFFA0FF09F938B9A5F918" bold="true" box="[199,421,1713,1737]" pageId="0" pageNumber="75">Description (Male).</emphasis>
As in the female (cf.
<bibRefCitation id="EF874B5EFF9AFFA0FD57F938BBDAF919" author="Gonzalez" box="[665,986,1713,1736]" pageId="0" pageNumber="75" refString="Gonzalez, V. H. &amp; Griswold, T. (2011) Heriades tayrona n. sp., the first osmiine bee from South America (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society, 84, 255 - 259." type="journal article" year="2011">Gonzalez and Griswold 2011</bibRefCitation>
), except: Body length,
<quantity id="4CEE9B4AFF9AFFA0FB16F938BD2FF919" box="[1240,1327,1713,1736]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="0" pageNumber="75" unit="mm" value="5.2">5.2 mm</quantity>
; forewing length,
<quantity id="4CEE9B4AFF9AFFA0FF29F958B93CF939" box="[231,316,1745,1768]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="0" pageNumber="75" unit="mm" value="3.5">3.5 mm</quantity>
. Compound eyes about twice as long as broad, two times wider than gena in profile; intertorular distance 1.8 times torulorbital distance; interocellar distance 4.3 times median ocellar diameter, about as long as ocelloccipital distance, slightly longer than ocellocular distance; labrum without small median tubercle basally; mandible with condylar ridge not abruptly angled, outer ridge and acetabular carina not abruptly ending in tubercle; clypeus 1.6 times broader than long, distal margin slightly concave, without shallow median emargination delimited by small tubercle; scape 1.9 times longer than wide, 0.6 times torulocellar distance; pedicel about as long as broad, slightly longer than first flagellomere; first flagellomere 1.4 times broader than long, about half length of second; second flagellomere 1.3 times longer than broad, about as long as third. Sixth and seventh terga, second to eighth sterna, and genital capsule as in figures 4–14.
</paragraph>
<caption id="DF696627FF9BFFA1FF59F8D0B95AF867" httpUri="https://zenodo.org/record/213904/files/figure.png" pageId="1" pageNumber="76" targetBox="[192,1417,219,1829]" targetPageId="1">
<paragraph id="8BA936AFFF9BFFA1FF59F8D0B95AF867" blockId="1.[151,1437,1881,1974]" pageId="1" pageNumber="76">
<emphasis id="B962EABDFF9BFFA1FF59F8D0B950F8A0" bold="true" box="[151,336,1881,1905]" pageId="1" pageNumber="76">FIGURES 1–14.</emphasis>
Male of
<taxonomicName id="4C164D2CFF9BFFA1FE7AF8D3BB69F8A0" authority="Gonzalez &amp; Griswold." authorityName="Gonzalez &amp; Griswold." box="[436,873,1882,1905]" class="Insecta" family="Megachilidae" genus="Heriades" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="76" phylum="Arthropoda" rank="species" species="tayrona">
<emphasis id="B962EABDFF9BFFA1FE7AF8D3BA6FF8A0" box="[436,623,1882,1905]" italics="true" pageId="1" pageNumber="76">Heriades tayrona</emphasis>
Gonzalez &amp; Griswold.
</taxonomicName>
1, Lateral habitus. 2, Dorsal habitus. 3, Facial view. 4, Metasoma in ventral view. 5, Seventh tergum. 6–11, Third to eighth sterna. 12–14, Genital capsule in lateral, dorsal and ventral views.
</paragraph>
</caption>
<caption id="DF696627FF98FFA2FF59F8DBBCE1F87B" httpUri="https://zenodo.org/record/213905/files/figure.png" pageId="2" pageNumber="77" targetBox="[199,1385,218,1818]" targetPageId="2">
<paragraph id="8BA936AFFF98FFA2FF59F8DBBCE1F87B" blockId="2.[151,1436,1874,1962]" pageId="2" pageNumber="77">
<emphasis id="B962EABDFF98FFA2FF59F8DBB961F8BB" bold="true" box="[151,353,1874,1898]" pageId="2" pageNumber="77">FIGURES 15–28.</emphasis>
Female (15–17) and male (18–28) of
<taxonomicName id="4C164D2CFF98FFA2FCDAF8DBBC69F8B8" authority="Gonzalez." authorityName="Gonzalez." box="[788,1129,1874,1897]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="rosellae">
<emphasis id="B962EABDFF98FFA2FCDAF8DBBBF2F8B8" box="[788,1010,1874,1897]" italics="true" pageId="2" pageNumber="77">Leioproctus rosellae</emphasis>
Gonzalez.
</taxonomicName>
15, 18, Dorsal habitus. 19, Lateral habitus. 16, 20, Facial views. 17, 21, Metasoma in dorsal views. 22, Sixth sternum. 23, Seventh sternum. 24, 25, Eighth sternum in lateral and ventral views. 26–28, Genital capsule in lateral, dorsal and ventral views.
</paragraph>
</caption>
<paragraph id="8BA936AFFF99FFA3FF09FF11BC53FF1E" blockId="3.[151,1436,152,336]" pageId="3" pageNumber="78">Integumental sculpture as in female except for first metasomal sternum and second laterally with punctures large, as those on scutum; median portion of second sternum with much smaller, denser punctures.</paragraph>
<paragraph id="8BA936AFFF99FFA3FF59FF51BA29FEDE" blockId="3.[151,1436,152,336]" pageId="3" pageNumber="78">Pubescence in general as in female, except for clypeus and supraclypeal area densely covered by short, minutelybranched hairs obscuring integument.</paragraph>
</subSubSection>
<subSubSection id="C30C6524FF99FFA3FF09FE91BC78FE81" pageId="3" pageNumber="78" type="materials_examined">
<paragraph id="8BA936AFFF99FFA3FF09FE91BC78FE81" blockId="3.[151,1436,152,336]" pageId="3" pageNumber="78">
<emphasis id="B962EABDFF99FFA3FF09FE91B9AEFEE1" bold="true" box="[199,430,280,304]" pageId="3" pageNumber="78">Examined material.</emphasis>
<collectingCountry id="F301763FFF99FFA3FE75FE90BA30FEE1" box="[443,560,281,304]" name="Colombia" pageId="3" pageNumber="78">COLOMBIA</collectingCountry>
: 23, Magdalena, Santa
<collectingCountry id="F301763FFF99FFA3FC85FE90BB8EFEE1" box="[843,910,281,304]" name="Malta" pageId="3" pageNumber="78">Marta</collectingCountry>
, on the road from Bastidas to
<taxonomicName id="4C164D2CFF99FFA3FB37FE90BD98FEE1" authority="Concha" authorityName="Concha" box="[1273,1432,281,304]" class="Magnoliopsida" family="Asteraceae" genus="Bahia" kingdom="Plantae" order="Asterales" pageId="3" pageNumber="78" phylum="Tracheophyta" rank="genus">Bahía Concha</taxonomicName>
,
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, 74º
<geoCoordinate id="EE225068FF99FFA3FE80FEB0B9B3FE81" box="[334,435,313,336]" direction="west" orientation="longitude" pageId="3" pageNumber="78" precision="55" value="-9.924">09.924W</geoCoordinate>
;
<date id="FFA8106FFF99FFA3FE70FEB0BA4EFE81" box="[446,590,313,336]" pageId="3" pageNumber="78" value="2011-12-18">Dec 18, 2011</date>
,
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<quantity id="4CEE9B4AFF99FFA3FD97FEB0BA94FE81" box="[601,660,313,336]" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="3" pageNumber="78" unit="m" value="99.0">99 m</quantity>
.
</specimenCount>
, V.H. Gonzalez, P. Sepúlveda (
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,
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).
</paragraph>
</subSubSection>
</treatment>
</document>
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