261 lines
31 KiB
XML
261 lines
31 KiB
XML
<document id="7E8D17C9F6CBD1C93D00B993062FA207" ID-CLB-Dataset="8059" ID-DOI="10.11646/zootaxa.4933.2.1" ID-GBIF-Dataset="1a2ba1ca-028f-4b95-8dda-5ddec4e63524" ID-ISSN="1175-5326" ID-Zenodo-Dep="4550146" ID-ZooBank="5D1D20A5-0F44-4AEF-AF5F-A758FC37D076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613716226573" checkinUser="plazi" docAuthor="Anistratenko, Vitaliy V., Neubauer, Thomas A., Anistratenko, Olga Yu., Kijashko, Pavel V. & Wesselingh, Frank P." docDate="2021" docId="03BF87A3C85AFFACFF3DFE94FCEAA1E6" docLanguage="en" docName="zootaxa.4933.2.1.pdf" docOrigin="Zootaxa 4933 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clathrocaspia knipowitschii subsp. aluschtensis" docType="treatment" docVersion="10" lastPageNumber="178" masterDocId="FF86FFDBC841FFB7FFAAFFDEFFA5A577" masterDocTitle="A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)" masterLastPageNumber="197" masterPageNumber="151" pageNumber="178" updateTime="1698930299740" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9E10E7BAD47A10152B80893BE13CC182">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="9ABC8737E45A9B0B8D1F1398B5BB12D6">Anistratenko, Vitaliy V.</mods:namePart>
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<mods:affiliation id="EE5CE7C3DC5877DAB7BF5EB50E3C3BCF">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="A47911BE4074B041FFB8CFB8C2913345">Neubauer, Thomas A.</mods:namePart>
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<mods:affiliation id="2648AF6B63BF50DED81A5C13A2FF6DEC">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:affiliation id="7B245ADE586EAE758DE49DEFFF29AFB0">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. & Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<mods:namePart id="3552894F80ED2DD412DEC2BE85ED5A64">Kijashko, Pavel V.</mods:namePart>
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<mods:affiliation id="920D9D406E3ACFF39BC9873DEE16E30F">Zoological Institute, Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<treatment id="03BF87A3C85AFFACFF3DFE94FCEAA1E6" ID-DOI="http://doi.org/10.5281/zenodo.4559946" ID-GBIF-Taxon="179906613" ID-Zenodo-Dep="4559946" LSID="urn:lsid:plazi:treatment:03BF87A3C85AFFACFF3DFE94FCEAA1E6" httpUri="http://treatment.plazi.org/id/03BF87A3C85AFFACFF3DFE94FCEAA1E6" lastPageNumber="178" pageId="27" pageNumber="178">
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<paragraph id="8BA936B5C85AFFACFF3DFE94FB3FA412" blockId="27.[151,1178,330,393]" box="[151,1178,330,357]" pageId="27" pageNumber="178">
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<emphasis id="B962EAA7C85AFFACFF3DFE94FB3FA412" bold="true" box="[151,1178,330,357]" pageId="27" pageNumber="178">
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†
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<taxonomicName id="4C164D36C85AFFACFF0CFE94FB8DA413" ID-CoL="86ZVG" authority="(Golikov & Starobogatov, 1966)" baseAuthorityName="Golikov & Starobogatov" baseAuthorityYear="1966" box="[166,1064,330,357]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="subSpecies" species="knipowitschii" subSpecies="aluschtensis">
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<emphasis id="B962EAA7C85AFFACFF0CFE94FD36A412" bold="true" box="[166,659,330,357]" italics="true" pageId="27" pageNumber="178">Clathrocaspia knipowitschii aluschtensis</emphasis>
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(
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<bibRefCitation id="EF874B44C85AFFACFD09FE94FB85A413" author="Golikov, A. N. & Starobogatov, Ya. I." box="[675,1056,330,356]" pageId="27" pageNumber="178" pagination="352 - 362" refId="ref30686" refString="Golikov, A. N. & Starobogatov, Ya. I. (1966) Ponto-kaspiyskiye bryukhonogiye mollyuski v Azovo-Chernomorskom basseyne. Zoologicheskiy Zhurnal, 45 (3), 352 - 362." type="journal article" year="1966">Golikov & Starobogatov, 1966</bibRefCitation>
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)
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</taxonomicName>
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comb. n.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C653EC85AFFACFF3DFEB1FF56A4FE" box="[151,243,367,393]" pageId="27" pageNumber="178" type="description">
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<paragraph id="8BA936B5C85AFFACFF3DFEB1FF56A4FE" blockId="27.[151,1178,330,393]" box="[151,243,367,393]" pageId="27" pageNumber="178">
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<figureCitation id="132D2A30C85AFFACFF3DFEB1FF56A4FE" box="[151,243,367,393]" captionStart="FIGURE 13" captionStartId="25.[151,250,1778,1802]" captionTargetBox="[254,1333,181,1754]" captionTargetId="figure-17@25.[254,1333,181,1754]" captionTargetPageId="25" captionText="FIGURE 13. Clathrocaspia knipowitschii (Makarov, 1938) from the Black Sea Basin. (a, b) Neotype, locality 30b (IZAN 522/1). (c, d) Specimen from locality 30b (IZAN 522/2). (e) C. knipowitschii aluschtensis (Golikov & Starobogatov, 1966), holotype, locality 29 (ZIN 4438/1). (f) Holotype of Pyrgula makarovi, locality 31 (ZIN 4492/1). (g, l, u) Specimen from Holocene deposits, locality 32 (IZAN, no number). (h, m) Specimen from same locality (IZAN, no number). (i, n) Paratype of C. brotzkajae from same locality (IZAN, no number). (j, k) Specimen from locality 30c (UGSB 25234). (o, s, t) Specimen identified as C. makarovi by Anistratenko (2013) from locality 30c (UGSB 25236). (p–r) Specimen from locality 30c (UGSB 25235). Scale bars = 1 mm (a–j, l–o), 500 μm (k), 100 μm (p–t), 50 μm (u)." figureDoi="http://doi.org/10.5281/zenodo.4550182" httpUri="https://zenodo.org/record/4550182/files/figure.png" pageId="27" pageNumber="178">Fig. 13e</figureCitation>
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</paragraph>
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<subSubSection id="C30C653EC85AFFACFF3DFE6DFB55A77E" pageId="27" pageNumber="178" type="reference_group">
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<paragraph id="8BA936B5C85AFFACFF3DFE6DFB30A4BC" blockId="27.[151,1371,435,521]" box="[151,1173,435,459]" pageId="27" pageNumber="178">
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<treatmentCitationGroup id="AB06119BC85AFFACFF3DFE6DFB30A4BC" box="[151,1173,435,459]" pageId="27" pageNumber="178">
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<taxonomicName id="4C164D36C85AFFACFF3DFE6DFE3CA4BC" ID-CoL="7SYLP" baseAuthorityName="Clessin & W.Dybowski" baseAuthorityYear="1887" box="[151,409,435,459]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="gmelini" subGenus="Caspia">
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<emphasis id="B962EAA7C85AFFACFF3DFE6DFF4DA4BC" box="[151,232,435,459]" italics="true" pageId="27" pageNumber="178">Pyrgula</emphasis>
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(
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<emphasis id="B962EAA7C85AFFACFF5DFE6AFE9AA4BC" box="[247,319,436,459]" italics="true" pageId="27" pageNumber="178">Caspia</emphasis>
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)
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<emphasis id="B962EAA7C85AFFACFEE7FE6DFE3CA4BC" box="[333,409,435,459]" italics="true" pageId="27" pageNumber="178">gmelini</emphasis>
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</taxonomicName>
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[sic]
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<emphasis id="B962EAA7C85AFFACFE79FE6DFDEAA4BC" box="[467,591,435,459]" italics="true" pageId="27" pageNumber="178">aluschtensis</emphasis>
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subsp. n.—
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<treatmentCitation id="0AB710A4C85AFFACFD6CFE6DFB94A4BC" author="Golikov, A. N. & Starobogatov, Ya. I." box="[710,1073,435,459]" page="354" pageId="27" pageNumber="178" year="1966">
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<bibRefCitation id="EF874B44C85AFFACFD6CFE6DFB94A4BC" author="Golikov, A. N. & Starobogatov, Ya. I." box="[710,1073,435,459]" pageId="27" pageNumber="178" pagination="352 - 362" refId="ref30686" refString="Golikov, A. N. & Starobogatov, Ya. I. (1966) Ponto-kaspiyskiye bryukhonogiye mollyuski v Azovo-Chernomorskom basseyne. Zoologicheskiy Zhurnal, 45 (3), 352 - 362." type="journal article" year="1966">Golikov & Starobogatov 1966: 354</bibRefCitation>
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</treatmentCitation>
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, fig. 1(8).
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</paragraph>
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<paragraph id="8BA936B5C85AFFACFF3DFE0CFB55A77E" blockId="27.[151,1371,435,521]" pageId="27" pageNumber="178">
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<emphasis id="B962EAA7C85AFFACFF3DFE0CFF0AA49E" box="[151,175,466,489]" italics="true" pageId="27" pageNumber="178">C.</emphasis>
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[
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<emphasis id="B962EAA7C85AFFACFF1DFE0CFF4BA49E" box="[183,238,466,489]" italics="true" pageId="27" pageNumber="178">aspia</emphasis>
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]
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<taxonomicName id="4C164D36C85AFFACFF57FE0CFEECA49D" ID-CoL="7SYLP" baseAuthorityName="Clessin & W.Dybowski" baseAuthorityYear="1887" box="[253,329,466,490]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="gmelini">
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<emphasis id="B962EAA7C85AFFACFF57FE0CFEECA49D" box="[253,329,466,490]" italics="true" pageId="27" pageNumber="178">gmelini</emphasis>
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||
</taxonomicName>
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||
[sic]
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||
<emphasis id="B962EAA7C85AFFACFE28FE0CFE5BA49D" box="[386,510,466,490]" italics="true" pageId="27" pageNumber="178">aluschtensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF874B44C85AFFACFDA1FE0CFCE7A49D" author="Golikov, A. N. & Starobogatov, Ya. I." box="[523,834,466,490]" pageId="27" pageNumber="178" pagination="352 - 362" refId="ref30686" refString="Golikov, A. N. & Starobogatov, Ya. I. (1966) Ponto-kaspiyskiye bryukhonogiye mollyuski v Azovo-Chernomorskom basseyne. Zoologicheskiy Zhurnal, 45 (3), 352 - 362." type="journal article" year="1966">Golikov et Starobogatov, 1966</bibRefCitation>
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)—
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||
<bibRefCitation id="EF874B44C85AFFACFCCEFE0CFB6BA49D" author="Golikov, A. N. & Starobogatov, Ya. I." box="[868,1230,466,490]" pageId="27" pageNumber="178" pagination="65 - 166" refId="ref30721" refString="Golikov, A. N. & Starobogatov, Ya. I. (1972) Klass bryukhonogiye mollyuski. In: Mordukhay-Boltovskoy, F. D. (Ed.), Opredelitel' fauny Chernogo i Azovskogo morey: Svobodnozhivushchiye bespozvonochnyye. T. 3. Chlenistonogiye (krome rakoobraznykh), mollyuski, iglokozhiye, shchetinkochelyustnyye, khordovyye. Naukova dumka, Kiev, pp. 65 - 166." type="book chapter" year="1972">Golikov & Starobogatov 1972: 100</bibRefCitation>
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||
, pl. 2, fig. 19.
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||
<emphasis id="B962EAA7C85AFFACFF3DFE2FFF0AA77F" box="[151,175,497,520]" italics="true" pageId="27" pageNumber="178">C.</emphasis>
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||
[
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<emphasis id="B962EAA7C85AFFACFF1DFE2FFF4BA77F" box="[183,238,497,520]" italics="true" pageId="27" pageNumber="178">aspia</emphasis>
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] (
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<emphasis id="B962EAA7C85AFFACFEAFFE2EFE81A77F" box="[261,292,496,520]" italics="true" pageId="27" pageNumber="178">Cl.</emphasis>
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[
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<emphasis id="B962EAA7C85AFFACFE86FE2EFE01A77F" box="[300,420,496,520]" italics="true" pageId="27" pageNumber="178">athrocaspia</emphasis>
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])
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<emphasis id="B962EAA7C85AFFACFE11FE2EFD39A77F" box="[443,668,496,520]" italics="true" pageId="27" pageNumber="178">
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||
<taxonomicName id="4C164D36C85AFFACFE11FE2EFDBFA77F" ID-CoL="7SYLR" authorityName="sensu Golikov & Starobogatov" authorityYear="1966" box="[443,538,496,520]" class="Gastropoda" family="Hydrobiidae" genus="Pyrgula" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="makarovi">makarovi</taxonomicName>
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aluschtensis
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</emphasis>
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(Gol. et St.)—
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<bibRefCitation id="EF874B44C85AFFACFC9AFE2EFB60A77E" author="Alexenko, T. L. & Starobogatov, Ya. I." box="[816,1221,496,521]" pageId="27" pageNumber="178" pagination="32 - 38" refId="ref28750" refString="Alexenko, T. L. & Starobogatov, Ya. I. (1987) Vidy Caspia i Turricaspia (Gastropoda, Pectinibranchia, Pyrgulidae) Azovo-Chernomorskogo basseyna. Vestnik Zoologii, 21 (3), 32 - 38." type="journal article" year="1987">Alexenko & Starobogatov 1987: 32–33</bibRefCitation>
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||
, 37.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C30C653EC85AFFACFF3DFDE9FE68A65E" pageId="27" pageNumber="178" type="materials_examined">
|
||
<paragraph id="8BA936B5C85AFFACFF3DFDE9FDBFA702" blockId="27.[151,1437,566,1169]" pageId="27" pageNumber="178">
|
||
<emphasis id="B962EAA7C85AFFACFF3DFDE9FEE1A727" bold="true" box="[151,324,566,593]" pageId="27" pageNumber="178">Type material.</emphasis>
|
||
<materialsCitation id="3B7E3CE8C85AFFACFEE0FDE9FDBFA702" ID-GBIF-Occurrence="3042095339" collectionCode="ZIN" collectorName="S. A. Zernov" pageId="27" pageNumber="178" specimenCode="ZIN 4438, ZIN 4439" specimenCount="18" typeStatus="holotype">
|
||
<typeStatus id="54AD8817C85AFFACFEE0FDE9FE17A726" box="[330,434,567,593]" pageId="27" pageNumber="178" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<specimenCode id="DBB09ECEC85AFFACFE15FDE9FD96A726" box="[447,563,567,593]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="178" type="Museum">ZIN 4438</specimenCode>
|
||
/1) and
|
||
<specimenCount id="9D10FD3CC85AFFACFD2BFDE9FCABA726" box="[641,782,567,593]" pageId="27" pageNumber="178" type="generic" typeStatus="paratypes">18 paratypes</specimenCount>
|
||
(
|
||
<specimenCode id="DBB09ECEC85AFFACFCB1FDE9FC29A725" box="[795,908,567,594]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="27" pageNumber="178" type="Museum">ZIN 4439</specimenCode>
|
||
/2–4443/6). Collected by
|
||
<collectorName id="26E35363C85AFFACFB0EFDE9FA88A726" box="[1188,1325,567,593]" pageId="27" pageNumber="178">S.A. Zernov</collectorName>
|
||
in 15/08– 15/09/1909, steamship “Meotida”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C85AFFACFF6DFDA1FC32A672" blockId="27.[151,1437,566,1169]" pageId="27" pageNumber="178">
|
||
<materialsCitation id="3B7E3CE8C85AFFACFF6DFDA1FC2AA672" ID-GBIF-Occurrence="3042095328" location="Crimea" pageId="27" pageNumber="178" specimenCount="2" stateProvince="Crimea" typeStatus="holotype">
|
||
<emphasis id="B962EAA7C85AFFACFF6DFDA1FEC0A7EF" bold="true" box="[199,357,638,665]" pageId="27" pageNumber="178">
|
||
<typeStatus id="54AD8817C85AFFACFF6DFDA1FEA7A7EE" box="[199,258,639,665]" pageId="27" pageNumber="178">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Black Sea, offshore of
|
||
<collectingRegion id="49D2F857C85AFFACFDC0FDA1FD19A7EE" box="[618,700,639,665]" country="Ukraine" name="Crimea" pageId="27" pageNumber="178">Crimea</collectingRegion>
|
||
near Alushta (
|
||
<tableCitation id="C694030EC85AFFACFCF7FDA1FC14A7EE" box="[861,945,639,665]" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="27" pageNumber="178" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
|
||
, locality 29). The material comes from phaseoline silts, which typically contain reworked Neoeuxinian (Late Pleistocene) shells (
|
||
<bibRefCitation id="EF874B44C85AFFACFBF1FD7DFAFAA7CA" author="Wesselingh, F. P. & Neubauer, T. A. & Anistratenko, V. V. & Vinarski, M. V. & Yanina, T. & ter Poorten, J. J. & Kijashko, P. V. & Albrecht, C. & Anistratenko, O. Yu. & D'Hont, A. & Frolov, P. & Martinez Gandara, A. & Gittenberger, A. & Gogaladze, A. & Karpinsky, M. & Lattuada, M. & Popa, L. & Sands, A. F. & van de Velde, S. & Vandendorpe, J. & Wilke, T." box="[1115,1375,675,701]" pageId="27" pageNumber="178" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. & Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
|
||
Wesselingh
|
||
<emphasis id="B962EAA7C85AFFACFB4EFD7DFAB0A7CA" box="[1252,1301,675,701]" italics="true" pageId="27" pageNumber="178">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
). The preservation of the
|
||
<typeStatus id="54AD8817C85AFFACFEC4FD19FE3BA796" box="[366,414,711,737]" pageId="27" pageNumber="178">type</typeStatus>
|
||
material is typical for this kind of reworked material (pers. obs.
|
||
<collectorName id="26E35363C85AFFACFBCCFD19FB0BA796" box="[1126,1198,711,737]" name="Frank P. Wesselingh" pageId="27" pageNumber="178">F.P.W.</collectorName>
|
||
06/2020).
|
||
<collectorName id="26E35363C85AFFACFA8FFD19FACAA796" box="[1317,1391,711,737]" pageId="27" pageNumber="178">Likely</collectorName>
|
||
, the subspecies is an extinct
|
||
<collectorName id="26E35363C85AFFACFE0EFD35FDF8A672" box="[420,605,747,773]" pageId="27" pageNumber="178">Late Pleistocene</collectorName>
|
||
lineage of
|
||
<taxonomicName id="4C164D36C85AFFACFD70FD35FC2AA672" authorityName="sensu Logvinenko & Starobogatov" authorityYear="1969" box="[730,911,747,773]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="knipowitschii">
|
||
<emphasis id="B962EAA7C85AFFACFD70FD35FC2AA672" box="[730,911,747,773]" italics="true" pageId="27" pageNumber="178">C. knipowitschii</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C85AFFACFF6DFCD0FE68A65E" blockId="27.[151,1437,566,1169]" box="[199,461,782,809]" pageId="27" pageNumber="178">
|
||
<emphasis id="B962EAA7C85AFFACFF6DFCD0FE24A65F" bold="true" box="[199,385,782,808]" pageId="27" pageNumber="178">Other material.</emphasis>
|
||
None.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC85AFFACFF6DFCECFBF6A606" pageId="27" pageNumber="178" type="description">
|
||
<paragraph id="8BA936B5C85AFFACFF6DFCECFBF6A606" blockId="27.[151,1437,566,1169]" pageId="27" pageNumber="178">
|
||
<emphasis id="B962EAA7C85AFFACFF6DFCECFEFCA63B" bold="true" box="[199,345,818,844]" pageId="27" pageNumber="178">Description.</emphasis>
|
||
Slender shell with five low convex whorls. Shell surface poorly preserved, only traces of reticulate sculpture are visible. Aperture ovoid, adjoined, leaving no umbilicus. Base steep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC85AFFACFF6DFCA5FA22A11A" pageId="27" pageNumber="178" type="discussion">
|
||
<paragraph id="8BA936B5C85AFFACFF6DFCA5FA22A11A" blockId="27.[151,1437,566,1169]" pageId="27" pageNumber="178">
|
||
<emphasis id="B962EAA7C85AFFACFF6DFCA5FE9EA6E2" bold="true" box="[199,315,891,917]" pageId="27" pageNumber="178">Remarks.</emphasis>
|
||
Originally this taxon was described as the Black Sea’s fossil subspecies of
|
||
<taxonomicName id="4C164D36C85AFFACFB38FCA5FA30A6E2" authorityName="Clessin et W. Dybowski" authorityYear="1888" box="[1170,1429,891,917]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="gmelinii">
|
||
<emphasis id="B962EAA7C85AFFACFB38FCA5FA30A6E2" box="[1170,1429,891,917]" italics="true" pageId="27" pageNumber="178">Clathrocaspia gmelinii</emphasis>
|
||
</taxonomicName>
|
||
, but later
|
||
<bibRefCitation id="EF874B44C85AFFACFF50FC41FDD6A6CE" author="Alexenko, T. L. & Starobogatov, Ya. I." box="[250,627,927,953]" pageId="27" pageNumber="178" pagination="32 - 38" refId="ref28750" refString="Alexenko, T. L. & Starobogatov, Ya. I. (1987) Vidy Caspia i Turricaspia (Gastropoda, Pectinibranchia, Pyrgulidae) Azovo-Chernomorskogo basseyna. Vestnik Zoologii, 21 (3), 32 - 38." type="journal article" year="1987">Alexenko & Starobogatov (1987)</bibRefCitation>
|
||
considered it to belong as subspecies in
|
||
<taxonomicName id="4C164D36C85AFFACFB95FC41FAEBA6CE" box="[1087,1358,927,953]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="makarovi">
|
||
<emphasis id="B962EAA7C85AFFACFB95FC41FAEBA6CE" box="[1087,1358,927,953]" italics="true" pageId="27" pageNumber="178">Clathrocaspia makarovi</emphasis>
|
||
</taxonomicName>
|
||
, based on the co-occurrence of that species in the Azov-Black Sea Basin.
|
||
<bibRefCitation id="EF874B44C85AFFACFC29FC1DFB66A6AA" author="Kantor, Yu. I. & Sysoev, A. V." box="[899,1219,963,989]" pageId="27" pageNumber="178" refId="ref31309" refString="Kantor, Yu. I. & Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor & Sysoev (2006: 88)</bibRefCitation>
|
||
went a step further and listed
|
||
<emphasis id="B962EAA7C85AFFACFEA4FC39FE32A176" box="[270,407,999,1025]" italics="true" pageId="27" pageNumber="178">aluschtensis</emphasis>
|
||
as a synonym of
|
||
<taxonomicName id="4C164D36C85AFFACFDCCFC39FD6AA176" box="[614,719,999,1025]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="makarovi">
|
||
<emphasis id="B962EAA7C85AFFACFDCCFC39FD6AA176" box="[614,719,999,1025]" italics="true" pageId="27" pageNumber="178">makarovi</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54AD8817C85AFFACFCB9FC39FCD0A176" box="[787,885,999,1025]" pageId="27" pageNumber="178" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="132D2A30C85AFFACFC2CFC39FC4DA176" box="[902,1000,999,1025]" captionStart="FIGURE 13" captionStartId="25.[151,250,1778,1802]" captionTargetBox="[254,1333,181,1754]" captionTargetId="figure-17@25.[254,1333,181,1754]" captionTargetPageId="25" captionText="FIGURE 13. Clathrocaspia knipowitschii (Makarov, 1938) from the Black Sea Basin. (a, b) Neotype, locality 30b (IZAN 522/1). (c, d) Specimen from locality 30b (IZAN 522/2). (e) C. knipowitschii aluschtensis (Golikov & Starobogatov, 1966), holotype, locality 29 (ZIN 4438/1). (f) Holotype of Pyrgula makarovi, locality 31 (ZIN 4492/1). (g, l, u) Specimen from Holocene deposits, locality 32 (IZAN, no number). (h, m) Specimen from same locality (IZAN, no number). (i, n) Paratype of C. brotzkajae from same locality (IZAN, no number). (j, k) Specimen from locality 30c (UGSB 25234). (o, s, t) Specimen identified as C. makarovi by Anistratenko (2013) from locality 30c (UGSB 25236). (p–r) Specimen from locality 30c (UGSB 25235). Scale bars = 1 mm (a–j, l–o), 500 μm (k), 100 μm (p–t), 50 μm (u)." figureDoi="http://doi.org/10.5281/zenodo.4550182" httpUri="https://zenodo.org/record/4550182/files/figure.png" pageId="27" pageNumber="178">Fig. 13e</figureCitation>
|
||
), which is illustrated here for the first time, indeed shows a close resemblance to the type of
|
||
<taxonomicName id="4C164D36C85AFFACFD47FBD5FC5EA152" box="[749,1019,1035,1061]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="makarovi">
|
||
<emphasis id="B962EAA7C85AFFACFD47FBD5FC5EA152" box="[749,1019,1035,1061]" italics="true" pageId="27" pageNumber="178">Clathrocaspia makarovi</emphasis>
|
||
</taxonomicName>
|
||
(=
|
||
<taxonomicName id="4C164D36C85AFFACFB8AFBD5FB10A152" authorityName="sensu Logvinenko & Starobogatov" authorityYear="1969" box="[1056,1205,1035,1061]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="knipowitschii">
|
||
<emphasis id="B962EAA7C85AFFACFB8AFBD5FB10A152" box="[1056,1205,1035,1061]" italics="true" pageId="27" pageNumber="178">knipowitschii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="132D2A30C85AFFACFB69FBD5FAB9A151" box="[1219,1308,1035,1062]" captionStart="FIGURE 13" captionStartId="25.[151,250,1778,1802]" captionTargetBox="[254,1333,181,1754]" captionTargetId="figure-17@25.[254,1333,181,1754]" captionTargetPageId="25" captionText="FIGURE 13. Clathrocaspia knipowitschii (Makarov, 1938) from the Black Sea Basin. (a, b) Neotype, locality 30b (IZAN 522/1). (c, d) Specimen from locality 30b (IZAN 522/2). (e) C. knipowitschii aluschtensis (Golikov & Starobogatov, 1966), holotype, locality 29 (ZIN 4438/1). (f) Holotype of Pyrgula makarovi, locality 31 (ZIN 4492/1). (g, l, u) Specimen from Holocene deposits, locality 32 (IZAN, no number). (h, m) Specimen from same locality (IZAN, no number). (i, n) Paratype of C. brotzkajae from same locality (IZAN, no number). (j, k) Specimen from locality 30c (UGSB 25234). (o, s, t) Specimen identified as C. makarovi by Anistratenko (2013) from locality 30c (UGSB 25236). (p–r) Specimen from locality 30c (UGSB 25235). Scale bars = 1 mm (a–j, l–o), 500 μm (k), 100 μm (p–t), 50 μm (u)." figureDoi="http://doi.org/10.5281/zenodo.4550182" httpUri="https://zenodo.org/record/4550182/files/figure.png" pageId="27" pageNumber="178">Fig. 13f</figureCitation>
|
||
) but differs by a slightly more slender shell and more flattened whorls. Given the differences in morphology and stratigraphy, we consider
|
||
<emphasis id="B962EAA7C85AFFACFF51FB8DFE24A11A" box="[251,385,1107,1133]" italics="true" pageId="27" pageNumber="178">aluschtensis</emphasis>
|
||
as an extinct subspecies of
|
||
<taxonomicName id="4C164D36C85AFFACFD05FB8DFCC4A11A" authorityName="sensu Logvinenko & Starobogatov" authorityYear="1969" box="[687,865,1107,1133]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="178" phylum="Mollusca" rank="species" species="knipowitschii">
|
||
<emphasis id="B962EAA7C85AFFACFD05FB8DFCC4A11A" box="[687,865,1107,1133]" italics="true" pageId="27" pageNumber="178">C. knipowitschii</emphasis>
|
||
</taxonomicName>
|
||
that inhabited the Black Sea during the Holocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC85AFFACFF6DFBA8FCEAA1E6" box="[199,847,1142,1169]" pageId="27" pageNumber="178" type="distribution">
|
||
<paragraph id="8BA936B5C85AFFACFF6DFBA8FCEAA1E6" blockId="27.[151,1437,566,1169]" box="[199,847,1142,1169]" pageId="27" pageNumber="178">
|
||
<emphasis id="B962EAA7C85AFFACFF6DFBA8FEC4A1E7" bold="true" box="[199,353,1142,1168]" pageId="27" pageNumber="178">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54AD8817C85AFFACFDCAFBA9FD35A1E6" box="[608,656,1143,1169]" pageId="27" pageNumber="178">type</typeStatus>
|
||
locality/stratum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |