307 lines
43 KiB
XML
307 lines
43 KiB
XML
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<mods:title id="0439B430CC277B2A20AB8A1DB97AFA31">A revision of the Pontocaspian gastropods of the subfamily Caspiinae (Caenogastropoda: Hydrobiidae)</mods:title>
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<mods:namePart id="0485D5345E4C02F661BBF180E36D4FC1">Anistratenko, Vitaliy V.</mods:namePart>
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<mods:affiliation id="8C23FE1C8DD7DEE0FB1A0311EE26B67A">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine.</mods:affiliation>
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<mods:affiliation id="7D31E414F21342C6491FB36C5615EB97">Department of Animal Ecology and Systematics, Justus Liebig University, Heinrich-Buff-Ring 26 - 32 IFZ, 35392 Giessen, Germany. tneub @ zo. jlug. de; https: // orcid. org / 0000 - 0002 - 1398 - 9941 & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, The Netherlands.</mods:affiliation>
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<mods:affiliation id="E8EE818FEFE8A24CFAA15FBD5B1F1278">Department of Invertebrate Fauna and Systematics, Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, B. Khmelnytsky Str. 15, 01030 Kiev, Ukraine. & Department of Cainozoic Deposits, Institute of Geological Sciences, National Academy of Sciences of Ukraine, O. Gontchar Str. 55 b, 01054 Kiev, Ukraine.</mods:affiliation>
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<taxonomicName id="4C164D36C856FFA0FF3DFE94FCECA413" ID-CoL="84WCV" authority="(Logvinenko & Starobogatov, 1969)" baseAuthorityName="Logvinenko & Starobogatov" baseAuthorityYear="1969" box="[151,841,330,357]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
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<emphasis id="B962EAA7C856FFA0FF3DFE94FE21A412" bold="true" box="[151,388,330,357]" italics="true" pageId="23" pageNumber="174">Clathrocaspia isseli</emphasis>
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(
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<bibRefCitation id="EF874B44C856FFA0FE3FFE94FCE4A413" author="Logvinenko, B. M. & Starobogatov, Ya. I." box="[405,833,330,356]" pageId="23" pageNumber="174" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov, 1969</bibRefCitation>
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)
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<paragraph id="8BA936B5C856FFA0FF3DFEB1FE85A4FE" blockId="23.[151,841,330,393]" box="[151,288,367,393]" pageId="23" pageNumber="174">
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<figureCitation id="132D2A30C856FFA0FF3DFEB1FF42A4FE" box="[151,231,367,393]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12</figureCitation>
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i–l, q
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</paragraph>
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<emphasis id="B962EAA7C856FFA0FF3DFE6DFF4DA4BC" box="[151,232,435,459]" italics="true" pageId="23" pageNumber="174">Pyrgula</emphasis>
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(
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<emphasis id="B962EAA7C856FFA0FF5DFE6AFE9AA4BC" box="[247,319,436,459]" italics="true" pageId="23" pageNumber="174">Caspia</emphasis>
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)
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<emphasis id="B962EAA7C856FFA0FEE7FE6DFE24A4BC" box="[333,385,435,459]" italics="true" pageId="23" pageNumber="174">isseli</emphasis>
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Logv. et Star.
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—
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<bibRefCitation id="EF874B44C856FFA0FDC8FE6DFC50A4BC" author="Logvinenko, B. M. & Starobogatov, Ya. I." box="[610,1013,435,459]" pageId="23" pageNumber="174" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov 1969: 378</bibRefCitation>
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, fig. 367(6).
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<emphasis id="B962EAA7C856FFA0FF3DFE0CFE87A49D" box="[151,290,466,490]" italics="true" pageId="23" pageNumber="174">Pyrgula isseli</emphasis>
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Logvinenko et Starobogatov, 1968
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</taxonomicName>
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—
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<treatmentCitation id="0AB710A4C856FFA0FD35FE0CFC1BA49D" author="Kantor, Yu. I. & Sysoev, A. V." box="[671,958,466,490]" page="100" pageId="23" pageNumber="174" year="2006">
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<bibRefCitation id="EF874B44C856FFA0FD35FE0CFC1BA49D" author="Kantor, Yu. I. & Sysoev, A. V." box="[671,958,466,490]" pageId="23" pageNumber="174" refId="ref31309" refString="Kantor, Yu. I. & Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor & Sysoev 2006: 100</bibRefCitation>
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, pl. 45, fig. P.
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<emphasis id="B962EAA7C856FFA0FF3DFE2EFEC7A77F" box="[151,354,496,520]" italics="true" pageId="23" pageNumber="174">Clathrocaspia isseli</emphasis>
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(
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<bibRefCitation id="EF874B44C856FFA0FEC5FE2EFD6BA77F" author="Logvinenko, B. M. & Starobogatov, Ya. I." box="[367,718,496,521]" pageId="23" pageNumber="174" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov, 1969</bibRefCitation>
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)
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—
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|
||
<bibRefCitation id="EF874B44C856FFA0FD45FE2EFBA4A77E" author="Wesselingh, F. P. & Neubauer, T. A. & Anistratenko, V. V. & Vinarski, M. V. & Yanina, T. & ter Poorten, J. J. & Kijashko, P. V. & Albrecht, C. & Anistratenko, O. Yu. & D'Hont, A. & Frolov, P. & Martinez Gandara, A. & Gittenberger, A. & Gogaladze, A. & Karpinsky, M. & Lattuada, M. & Popa, L. & Sands, A. F. & van de Velde, S. & Vandendorpe, J. & Wilke, T." box="[751,1025,496,521]" pageId="23" pageNumber="174" pagination="31 - 124" refId="ref34307" refString="Wesselingh, F. P., Neubauer, T. A., Anistratenko, V. V., Vinarski, M. V., Yanina, T., ter Poorten, J. J., Kijashko, P. V., Albrecht, C., Anistratenko, O. Yu., D'Hont, A., Frolov, P., Martinez Gandara, A., Gittenberger, A., Gogaladze, A., Karpinsky, M., Lattuada, M., Popa, L., Sands, A. F., van de Velde, S., Vandendorpe, J. & Wilke, T. (2019) Mollusc species from the Pontocaspian region - an expert opinion list. ZooKeys, 827, 31 - 124. https: // doi. org / 10.3897 / zookeys. 827.31365" type="journal article" year="2019">
|
||
Wesselingh
|
||
<emphasis id="B962EAA7C856FFA0FCC6FE2FFC3CA77F" box="[876,921,496,520]" italics="true" pageId="23" pageNumber="174">et al</emphasis>
|
||
. 2019: 70
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC856FFA0FF3DFDE9FAEBA6CE" pageId="23" pageNumber="174" type="materials_examined">
|
||
<paragraph id="8BA936B5C856FFA0FF3DFDE9FB1FA65E" blockId="23.[151,1437,566,1385]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF3DFDE9FEE0A727" bold="true" box="[151,325,566,593]" pageId="23" pageNumber="174">Type material.</emphasis>
|
||
<typeStatus id="54AD8817C856FFA0FEE1FDE9FE16A726" box="[331,435,567,593]" pageId="23" pageNumber="174" type="holotype">Holotype</typeStatus>
|
||
not traced, should be in
|
||
<collectionCode id="ED07AE70C856FFA0FD6FFDE9FD52A726" box="[709,759,567,593]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="23" pageNumber="174" type="Museum">ZIN</collectionCode>
|
||
. In contrast to
|
||
<bibRefCitation id="EF874B44C856FFA0FC31FDE9FB4DA726" author="Kantor, Yu. I. & Sysoev, A. V." box="[923,1256,567,593]" pageId="23" pageNumber="174" refId="ref31309" refString="Kantor, Yu. I. & Sysoev, A. V. (2006) Morskiye i solonovatovodnyye bryukhonogiye mollyuski Rossii i sopredel'nykh stran: illyustrirovannyy katalog. KMK Scientific Press, Moscow, 372 pp., 140 pls pp." type="book" year="2006">Kantor & Sysoev (2006: 100)</bibRefCitation>
|
||
, who stated that no identified material of this species is found in
|
||
<collectionCode id="ED07AE70C856FFA0FD12FD85FD4CA702" box="[696,745,603,629]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="23" pageNumber="174" type="Museum">ZIN</collectionCode>
|
||
, we could locate two lots with
|
||
<specimenCount id="9D10FD3CC856FFA0FBEDFD85FB45A702" box="[1095,1248,603,629]" pageId="23" pageNumber="174" type="generic">23 specimens</specimenCount>
|
||
in the collection of Starobogatov, which he labelled as “
|
||
<taxonomicName id="4C164D36C856FFA0FDE5FDA1FD45A7EE" box="[591,736,639,665]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FDE5FDA1FD45A7EE" box="[591,736,639,665]" italics="true" pageId="23" pageNumber="174">Caspia isseli</emphasis>
|
||
</taxonomicName>
|
||
”. Likely, these are
|
||
<typeStatus id="54AD8817C856FFA0FC13FDA1FB81A7EE" box="[953,1060,639,665]" pageId="23" pageNumber="174" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C164D36C856FFA0FBE0FDA1FB00A7EE" box="[1098,1189,639,665]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FBE0FDA1FB00A7EE" box="[1098,1189,639,665]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
. However, the collection includes a variety of morphologies, some of which match the typical
|
||
<taxonomicName id="4C164D36C856FFA0FC77FD7DFB9CA7CA" box="[989,1081,675,701]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FC77FD7DFB9CA7CA" box="[989,1081,675,701]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
as described by
|
||
<bibRefCitation id="EF874B44C856FFA0FB5DFD7DFEDAA795" author="Logvinenko, B. M. & Starobogatov, Ya. I." pageId="23" pageNumber="174" pagination="308 - 385" refId="ref32049" refString="Logvinenko, B. M. & Starobogatov, Ya. I. (1969 [" 1968 "]) Mollusca. In: Birshtein, Y. A., Vinogradov, L. G., Kondakov, N. N., Kuhn, M. S., Astakhova, T. V. & Romanova, N. N. (Eds.), Atlas bespozvonochnykh Kaspiyskogo morya. Pishchevaya Promyshlennost (Vsesoyuznyi Nauchno-issledovatel'skii Institut Morskogo Rybnogo Khozyaistva i Okeanografii), Moskva, pp. 308 - 385." type="book chapter" year="1969">Logvinenko & Starobogatov (1969)</bibRefCitation>
|
||
(e.g.
|
||
<figureCitation id="132D2A30C856FFA0FE6BFD19FD97A795" box="[449,562,711,738]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12i, j</figureCitation>
|
||
), while some resemble
|
||
<taxonomicName id="4C164D36C856FFA0FC94FD19FC11A796" authorityName="Clessin & W. Dybowski" authorityYear="1887" box="[830,948,711,737]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="pallasii">
|
||
<emphasis id="B962EAA7C856FFA0FC94FD19FC11A796" box="[830,948,711,737]" italics="true" pageId="23" pageNumber="174">C. pallasii</emphasis>
|
||
</taxonomicName>
|
||
closely and should be classified under that species (e.g.
|
||
<figureCitation id="132D2A30C856FFA0FE8DFD35FE06A671" box="[295,419,747,774]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">
|
||
Fig.
|
||
<quantity id="4CEE9B50C856FFA0FEF0FD35FE22A672" box="[346,391,747,773]" metricMagnitude="-2" metricUnit="kg" metricValue="1.2" pageId="23" pageNumber="174" unit="g" value="12.0">12g</quantity>
|
||
, h
|
||
</figureCitation>
|
||
). Given the lack of the
|
||
<typeStatus id="54AD8817C856FFA0FD0DFD35FCA9A672" box="[679,780,747,773]" pageId="23" pageNumber="174" type="holotype">holotype</typeStatus>
|
||
, the identity of
|
||
<taxonomicName id="4C164D36C856FFA0FC10FD35FBB1A672" box="[954,1044,747,773]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FC10FD35FBB1A672" box="[954,1044,747,773]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
is doubtful. For the time being, we use all of the presumed
|
||
<typeStatus id="54AD8817C856FFA0FE08FCD1FDA8A65E" box="[418,525,783,809]" pageId="23" pageNumber="174" type="paratype">paratypes</typeStatus>
|
||
that are not misidentified
|
||
<taxonomicName id="4C164D36C856FFA0FC9EFCD1FC0CA65E" authorityName="Clessin & W. Dybowski" authorityYear="1887" box="[820,937,783,809]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="pallasii">
|
||
<emphasis id="B962EAA7C856FFA0FC9EFCD1FC0CA65E" box="[820,937,783,809]" italics="true" pageId="23" pageNumber="174">C. pallasii</emphasis>
|
||
</taxonomicName>
|
||
to characterise
|
||
<taxonomicName id="4C164D36C856FFA0FBF3FCD1FB11A65E" box="[1113,1204,783,809]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FBF3FCD1FB11A65E" box="[1113,1204,783,809]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C856FFA0FF6DFCEDFC10A6E2" blockId="23.[151,1437,566,1385]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF6DFCEDFEC0A63B" bold="true" box="[199,357,818,845]" pageId="23" pageNumber="174">Type locality.</emphasis>
|
||
<materialsCitation id="3B7E3CE8C856FFA0FEC1FCEDFC14A6E2" ID-GBIF-Occurrence="3042095338" country="Azerbaijan" county="Cheleken Peninsula" elevation="58" location="Caspian Sea" municipality="Apsheron Peninsula" pageId="23" pageNumber="174" specimenCount="1" typeStatus="paratype">
|
||
Originally given as “Southern Caspian Sea between
|
||
<quantity id="4CEE9B50C856FFA0FC1AFCECFBB7A63A" box="[944,1042,818,845]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="23" pageNumber="174" unit="m" value="57.5" valueMax="75.0" valueMin="40.0">
|
||
<elevation id="003BD186C856FFA0FC1AFCECFBB7A63A" box="[944,1042,818,845]" metricMagnitude="1" metricUnit="m" metricValue="5.75" metricValueMax="7.5" metricValueMin="4.0" pageId="23" pageNumber="174" unit="m" value="57.5" valueMax="75.0" valueMin="40.0">40–75 m</elevation>
|
||
</quantity>
|
||
water depth” without details on the locality. The presumed
|
||
<typeStatus id="54AD8817C856FFA0FE35FC89FE5AA606" box="[415,511,855,881]" pageId="23" pageNumber="174" type="paratype">paratype</typeStatus>
|
||
material derives from the
|
||
<location id="8EC9606EC856FFA0FC81FC89FC10A606" LSID="urn:lsid:plazi:treatment:03BF87A3C856FFA0FF3DFE94FC24A01E:8EC9606EC856FFA0FC81FC89FC10A606" box="[811,949,855,881]" country="Azerbaijan" county="Cheleken Peninsula" municipality="Apsheron Peninsula" name="Caspian Sea" pageId="23" pageNumber="174">Caspian Sea</location>
|
||
off
|
||
<collectingMunicipality id="6BCDACCFC856FFA0FC4EFC89FB6DA606" box="[996,1224,855,881]" pageId="23" pageNumber="174">Apsheron Peninsula</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3017625C856FFA0FB78FC89FAE8A606" box="[1234,1357,855,881]" name="Azerbaijan" pageId="23" pageNumber="174">Azerbaijan</collectingCountry>
|
||
(
|
||
<tableCitation id="C694030EC856FFA0FAF6FC89FF00A6E2" captionStart="TABLE 1" captionStartId="3.[151,239,473,499]" captionTargetPageId="3" captionText="TABLE 1. Locality information of the studied material. Coordinates of collecting sites are given based on data in museum catalogue cards, labels and corresponding publications. Asterisks mark localities with approximate geographic coordinates based on indirect data provided in the ZIN catalogue. Abbreviations: q.—quadrant, st.—station." httpUri="http://table.plazi.org/id/DF69663DC842FFB4FF3DFE07F8C6A760" pageId="23" pageNumber="174" tableUuid="DF69663DC842FFB4FF3DFE07F8C6A760">Table 1</tableCitation>
|
||
, locality 14), and off
|
||
<collectingCounty id="62C84E39C856FFA0FE32FCA5FDDCA6E2" box="[408,633,891,917]" pageId="23" pageNumber="174">Cheleken Peninsula</collectingCounty>
|
||
,
|
||
<collectingCountry id="F3017625C856FFA0FD29FCA5FCBBA6E2" box="[643,798,891,917]" name="Turkmenistan" pageId="23" pageNumber="174">Turkmenistan</collectingCountry>
|
||
(locality 21)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA936B5C856FFA0FF6DFC40FAEBA6CE" blockId="23.[151,1437,566,1385]" box="[199,1358,926,953]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF6DFC40FE24A6CF" bold="true" box="[199,385,926,952]" pageId="23" pageNumber="174">Other material.</emphasis>
|
||
A single shell from Holocene deposits of the Kura delta (locality 25f,
|
||
<collectionCode id="ED07AE70C856FFA0FB3EFC41FB71A6CE" box="[1172,1236,927,953]" country="Netherlands" httpUri="http://grbio.org/cool/47nz-wbm0" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="174" type="Museum">RGM</collectionCode>
|
||
1310305).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC856FFA0FF6DFC1CFB71A11A" pageId="23" pageNumber="174" type="description">
|
||
<paragraph id="8BA936B5C856FFA0FF6DFC1CFB71A11A" blockId="23.[151,1437,566,1385]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF6DFC1CFEFCA6AB" bold="true" box="[199,345,962,988]" pageId="23" pageNumber="174">Description.</emphasis>
|
||
Slender, elongate, conical shell with up to 5 low convex, tightly coiled whorls. Protoconch bulbous, bears small wrinkles and thin spiral threads throughout (
|
||
<figureCitation id="132D2A30C856FFA0FCF5FC39FC67A176" box="[863,962,999,1025]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12q</figureCitation>
|
||
). Strong reticulate sculpture covers teleoconch, whereas spiral keels are more prominent than axial ribs (
|
||
<figureCitation id="132D2A30C856FFA0FCDFFBD5FC6DA151" box="[885,968,1035,1062]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12</figureCitation>
|
||
i–l). Small subsutural ramp occurs above first keel, sometimes resulting in slightly stepped outline (
|
||
<figureCitation id="132D2A30C856FFA0FC8BFBF1FCDEA13D" box="[801,891,1071,1098]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12l</figureCitation>
|
||
). First 1–2 keels are always more prominent and separated by wider interspaces than the ones following abapically. Aperture adjoined, base steep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC856FFA0FF6DFBA9FAADA056" pageId="23" pageNumber="174" type="discussion">
|
||
<paragraph id="8BA936B5C856FFA0FF6DFBA9FAADA056" blockId="23.[151,1437,566,1385]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF6DFBA9FE9EA1E6" bold="true" box="[199,315,1143,1169]" pageId="23" pageNumber="174">Remarks.</emphasis>
|
||
The material Starobogatov distinguished as
|
||
<taxonomicName id="4C164D36C856FFA0FC84FBA9FC2CA1E6" box="[814,905,1143,1169]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FC84FBA9FC2CA1E6" box="[814,905,1143,1169]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C164D36C856FFA0FC15FBA9FB91A1E6" authorityName="Clessin & W. Dybowski" authorityYear="1887" box="[959,1076,1143,1169]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="pallasii">
|
||
<emphasis id="B962EAA7C856FFA0FC15FBA9FB91A1E6" box="[959,1076,1143,1169]" italics="true" pageId="23" pageNumber="174">C. pallasii</emphasis>
|
||
</taxonomicName>
|
||
contains a mix of both. In addition to the mix up in the
|
||
<typeStatus id="54AD8817C856FFA0FE05FB45FE7AA1C2" box="[431,479,1179,1205]" pageId="23" pageNumber="174">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C164D36C856FFA0FDC0FB45FD60A1C2" box="[618,709,1179,1205]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FDC0FB45FD60A1C2" box="[618,709,1179,1205]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
(see above), Starobogatov’s material of
|
||
<taxonomicName id="4C164D36C856FFA0FB27FB45FAA6A1C2" authorityName="Clessin & W. Dybowski" authorityYear="1887" box="[1165,1283,1179,1205]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="pallasii">
|
||
<emphasis id="B962EAA7C856FFA0FB27FB45FAA6A1C2" box="[1165,1283,1179,1205]" italics="true" pageId="23" pageNumber="174">C. pallasii</emphasis>
|
||
</taxonomicName>
|
||
also contains specimens of
|
||
<taxonomicName id="4C164D36C856FFA0FE9AFB61FE2FA1AE" box="[304,394,1215,1241]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FE9AFB61FE2FA1AE" box="[304,394,1215,1241]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<figureCitation id="132D2A30C856FFA0FE63FB61FD8FA1AD" box="[457,554,1215,1242]" captionStart="FIGURE 12" captionStartId="24.[151,250,1720,1744]" captionTargetBox="[151,1436,181,1696]" captionTargetId="figure-22@24.[151,1436,181,1696]" captionTargetPageId="24" captionText="FIGURE 12. (a, b) Clathrocaspia pallasii (Clessin & Dybowski in Dybowski, 1887), paralectotype, locality 26 (ZB-M W.Dyb. 104). (c, d) C. pallasii, lectotype, locality 26 (ZB-M W.Dyb. 104). (e) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (f) C. pallasii, specimen labelled as “Caspia microcingulifera, paratype”, locality 9 (ZIN, no number). (g, h) C. pallasii, paratype of C. isseli (Logvinenko & Starobogatov, 1969), locality 14 (ZIN, no number). (i, j). C. isseli, paratype, locality 21 (ZIN, no number). (k) C. isseli, specimen misidentified as C. pallasii, locality 16 (ZIN, no number). (l, q) C. isseli, from Holocene deposits of the Kura delta (locality 25f, RGM 1310305). (m, n) C. pallasii, paratype of Pyrgula sowinskyi, locality 24 (ZIN, no number). (o, r) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 105). (p, s) C. pallasii, paralectotype, locality 26 (ZB-M W.Dyb. 74). Scale bars = 1 mm (a–n), 100 μm (o–s)." figureDoi="http://doi.org/10.5281/zenodo.4550180" httpUri="https://zenodo.org/record/4550180/files/figure.png" pageId="23" pageNumber="174">Fig. 12k</figureCitation>
|
||
). Despite their overall similarity in shell shape,
|
||
<taxonomicName id="4C164D36C856FFA0FB94FB61FB17A1AE" authorityName="Clessin & W. Dybowski" authorityYear="1887" box="[1086,1202,1215,1241]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="pallasii">
|
||
<emphasis id="B962EAA7C856FFA0FB94FB61FB17A1AE" box="[1086,1202,1215,1241]" italics="true" pageId="23" pageNumber="174">C. pallasii</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="4C164D36C856FFA0FF7FFB3DFE94A18A" box="[213,305,1251,1277]" class="Gastropoda" family="Hydrobiidae" genus="Caspia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FF7FFB3DFE94A18A" box="[213,305,1251,1277]" italics="true" pageId="23" pageNumber="174">C. isseli</emphasis>
|
||
</taxonomicName>
|
||
by the consistently lower base, the less convex whorls, the slightly detached aperture and the larger and more bulbous protoconch.
|
||
<taxonomicName id="4C164D36C856FFA0FE59FAD9FD76A056" baseAuthorityName="Logvinenko & Starobogatov" baseAuthorityYear="1969" box="[499,723,1287,1313]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="isseli">
|
||
<emphasis id="B962EAA7C856FFA0FE59FAD9FD76A056" box="[499,723,1287,1313]" italics="true" pageId="23" pageNumber="174">Clathrocaspia isseli</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C164D36C856FFA0FCCDFAD9FC46A056" authorityName="Clessin et W. Dybowski" authorityYear="1888" box="[871,995,1287,1313]" class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="174" phylum="Mollusca" rank="species" species="gmelinii">
|
||
<emphasis id="B962EAA7C856FFA0FCCDFAD9FC46A056" box="[871,995,1287,1313]" italics="true" pageId="23" pageNumber="174">C. gmelinii</emphasis>
|
||
</taxonomicName>
|
||
in its more slender shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30C653EC856FFA0FF6DFAF4FC24A01E" pageId="23" pageNumber="174" type="distribution">
|
||
<paragraph id="8BA936B5C856FFA0FF6DFAF4FC24A01E" blockId="23.[151,1437,566,1385]" pageId="23" pageNumber="174">
|
||
<emphasis id="B962EAA7C856FFA0FF6DFAF4FEC4A033" bold="true" box="[199,353,1322,1348]" pageId="23" pageNumber="174">Distribution.</emphasis>
|
||
Endemic to the Caspian Sea, only known from two localities in the South Basin (from depths of 40 and
|
||
<quantity id="4CEE9B50C856FFA0FF40FA91FE90A01E" box="[234,309,1359,1385]" metricMagnitude="2" metricUnit="m" metricValue="1.0699999999999998" pageId="23" pageNumber="174" unit="m" value="107.0">107 m</quantity>
|
||
). Also found in Holocene deposits of the Kura delta.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |