217 lines
29 KiB
XML
217 lines
29 KiB
XML
<document id="D464F6D0C8E287DDAC881E431C4FBBDB" ID-CLB-Dataset="31519" ID-DOI="10.11646/zootaxa.4079.2.5" ID-GBIF-Dataset="4b323554-9845-4c4b-b4f7-73b88422eaae" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050645" ID-ZooBank="91DF9041-B3CD-49E9-8250-D0C251938B45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510784073639" checkinUser="plazi" docAuthor="Yoshizawa, Kazunori & Lienhard, Charles" docDate="2016" docId="03BF294EFFC29F08FF38FAA65A99F990" docLanguage="en" docName="zootaxa.4079.2.5.pdf" docOrigin="Zootaxa 4079 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mydiognathus eviohlhoffae Yoshizawa & Lienhard, 2016, n. sp." docType="treatment" docUuid="FE221270-C927-43B7-915B-2010671DA809" docUuidSource="ZooBank" docVersion="8" lastPageNumber="235" masterDocId="FF865136FFC09F0EFFAFFFD1586FFFF7" masterDocTitle="Bridging the gap between chewing and sucking in the hemipteroid insects: new insights from Cretaceous amber" masterLastPageNumber="245" masterPageNumber="229" pageNumber="231" updateTime="1698491536162" updateUser="ExternalLinkService">
|
||
<mods:mods id="680C615B25DE975274AF8759D3AAB404" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="C3F148FEDAF6BD00C2D2353E253900B3">
|
||
<mods:title id="F195FD929FA52310625E0355A481C298">Bridging the gap between chewing and sucking in the hemipteroid insects: new insights from Cretaceous amber</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FD0EE4B4BB62416DDDC485CB31329FC0" type="personal">
|
||
<mods:role id="929E446E0817A31CEE9F70DB1905E3F4">
|
||
<mods:roleTerm id="231EFF6947CDE1B9175AA64E9632CE3B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1E06CD5AB015971D6B26DE7EBAA4D0D3">Yoshizawa, Kazunori</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="EFF6684FF9CA068AF629054C20F4E842" type="personal">
|
||
<mods:role id="05750CA10489CE512B1913C6466B7C39">
|
||
<mods:roleTerm id="96D5CB90BF3A04454F32F742979F534C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A608C4401C7246594E7786D568EA72C8">Lienhard, Charles</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FC1018F7FFD94899C41F77D84F554869">text</mods:typeOfResource>
|
||
<mods:relatedItem id="D2539A4DC9C15B08600E5CCF76581301" type="host">
|
||
<mods:titleInfo id="689BBACEFEBBD15FCF4E656553BD2FD9">
|
||
<mods:title id="3FB0E58A6A3F71CAF16D3562AD5664AE">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="9D012B6ED9DE04583A41297162E3E736">
|
||
<mods:date id="4AB9DF2DB30FA37B3D471302D4CE5FE8">2016</mods:date>
|
||
<mods:detail id="86C04B02CE63A40054938C88580D3FE6" type="volume">
|
||
<mods:number id="0A66AC787953694B5CB9788F68E872B4">4079</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F3CF8836F5B1652AAD9509E4E72EFCB9" type="issue">
|
||
<mods:number id="1A7AA461662D7228AE65F075BC1C664C">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5F2B71EBAD4E20B28CE2FC8A121F1CFE" unit="page">
|
||
<mods:start id="48CD6D15A104D41A8526C587B23788AF">229</mods:start>
|
||
<mods:end id="C5BD6960114725665E6428713325778E">245</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="AC1F5F2FB58B0E2C77B3F8C74FB11445">journal article</mods:classification>
|
||
<mods:identifier id="12A1E6F6CA4D3845F79F6143C0B9D9C0" type="CLB-Dataset">31519</mods:identifier>
|
||
<mods:identifier id="E0B76A22A72C1004D6473B1DA823EE11" type="DOI">10.11646/zootaxa.4079.2.5</mods:identifier>
|
||
<mods:identifier id="4A5AEA7BC4481C83ED94BB168A9D67C8" type="GBIF-Dataset">4b323554-9845-4c4b-b4f7-73b88422eaae</mods:identifier>
|
||
<mods:identifier id="84776D8FFFC6C680FFC1249932F4AA07" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="A895359DC494A31E0FB8B6882487A95D" type="Zenodo-Dep">1050645</mods:identifier>
|
||
<mods:identifier id="3A285B41B8B5694C4BB33F503241AB61" type="ZooBank">91DF9041-B3CD-49E9-8250-D0C251938B45</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03BF294EFFC29F08FF38FAA65A99F990" ID-DOI="http://doi.org/10.5281/zenodo.6079966" ID-GBIF-Taxon="137307197" ID-Zenodo-Dep="6079966" LSID="urn:lsid:zoobank.org:act:FE221270-C927-43B7-915B-2010671DA809" httpUri="http://treatment.plazi.org/id/03BF294EFFC29F08FF38FAA65A99F990" lastPageId="6" lastPageNumber="235" pageId="2" pageNumber="231">
|
||
<subSubSection id="C30CCBD3FFC29F0CFF38FAA65914FA44" pageId="2" pageNumber="231" type="nomenclature">
|
||
<paragraph id="8BA99858FFC29F0CFF38FAA65A42FA65" blockId="2.[151,557,1399,1459]" box="[151,557,1399,1426]" pageId="2" pageNumber="231">
|
||
<heading id="D0E12F34FFC29F0CFF38FAA65A42FA65" bold="true" box="[151,557,1399,1426]" fontSize="11" level="1" pageId="2" pageNumber="231" reason="1">
|
||
<emphasis id="B962444AFFC29F0CFF38FAA65A42FA65" bold="true" box="[151,557,1399,1426]" pageId="2" pageNumber="231">
|
||
<taxonomicName id="4C16E3DBFFC29F0CFF38FAA6598CFA66" box="[151,483,1399,1425]" class="Insecta" family="Archipsyllidae" genus="Mydiognathus" kingdom="Animalia" order="Psocodea" pageId="2" pageNumber="231" phylum="Arthropoda" rank="species" species="eviohlhoffae" status="sp. nov.">
|
||
<emphasis id="B962444AFFC29F0CFF38FAA6598CFA66" bold="true" box="[151,483,1399,1425]" italics="true" pageId="2" pageNumber="231">Mydiognathus eviohlhoffae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A251F931FFC29F0CFE44FAA95A42FA65" box="[491,557,1400,1426]" pageId="2" pageNumber="231" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BA99858FFC29F0CFF38FA4B5914FA44" blockId="2.[151,557,1399,1459]" box="[151,379,1434,1459]" pageId="2" pageNumber="231">
|
||
(
|
||
<figureCitation id="132D84DDFFC29F0CFF30FA4B5883FA44" box="[159,236,1434,1459]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="2" pageNumber="231">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="132D84DDFFC29F0CFF55FA4A5967FA44" box="[250,264,1435,1459]" captionStart="FIGURE 2" captionStartId="4.[151,250,1200,1222]" captionTargetBox="[153,1434,211,1154]" captionTargetId="figure@4.[151,1436,211,1155]" captionTargetPageId="4" captionText="FIGURE 2. Key head structures of Mydiognathus eviohlhoffae (a – c) and Neotrogla aurora (d). (a) Head. Dotted gray lines indicate alignment of mouthpart structures against the labrum (Lr) or lacinia (Lc, only for galea: Ga) (Acl: anteclypeus; At: antenna; ATP: anterior tentorial pit; Ca: cardo; E: eye; ES: epistomal suture; FGR: frontogenal suture; Fr: frons; Ga: galea; Ge: gena; Lc: lacinia; LiP: labial palpus; Lr: labrum; Md: mandible; MxP: maxillary palpus; Oc: ocellus; Pcl: postclypeus; Pf: palpifer; Pg: paraglossa; St: stipes). (b) Head, magnified. White arrowheads indicate the internally expanded lacinial base. (c) Tip of the mouth, magnified. (d) Antennal flagellum of Neotrogla aurora (Psocodea: Trogiomorpha: Prionoglarididae), showing the rupturing mechanism observed in extant Psocodea." httpUri="https://zenodo.org/record/1050649/files/figure.png" pageId="2" pageNumber="231">2</figureCitation>
|
||
a–c, 3a–c)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CCBD3FFC29F08FF38FA335C8DFA43" lastPageId="6" lastPageNumber="235" pageId="2" pageNumber="231" type="description">
|
||
<paragraph id="8BA99858FFC29F0DFF38FA335C17FF22" blockId="2.[151,1437,1506,2035]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="232" pageId="2" pageNumber="231">
|
||
<emphasis id="B962444AFFC29F0CFF38FA33594EFA0C" bold="true" box="[151,289,1506,1531]" pageId="2" pageNumber="231">Description</emphasis>
|
||
.
|
||
<emphasis id="B962444AFFC29F0CFE9EFA335902FA0C" box="[305,365,1506,1531]" italics="true" pageId="2" pageNumber="231">Head</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC29F0CFED2FA3359AEFA0C" box="[381,449,1506,1531]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="2" pageNumber="231">Figs 1</figureCitation>
|
||
a–d, 2a–c). Hypognathous, detached from thorax. Vertex rounded, protruding dorsally; medially concave along occipital suture. Frons bulbous. Frontogenal suture directed backward from anterior tentorial pit along ventral head capsule then strongly curved dorsally, ending between eye and antennal socket. Gena expanded laterally. Eye large; three ocelli, each on a tubercle. Postclypeus not expanded dorsally, not bulged; anteclypeus less sclerotized, with pair of swellings laterally. Antennal socket strongly pedicellate. Antenna filiform, with at least 16 articles; scape nearly cylindrical; pedicel strongly constricted basally, cylindrical distally, with bifurcated tip; flagellum long, far exceeding forewing length, no rupturing mechanism recognizable; flagellomeres annulate except for first one (f1) and basal half of f2. Labrum much longer than wide, apically rounded, covering entire elongated mandibles anteriorly. Mandibles elongate, apically curved inwardly, with two subapical conical teeth; basally broadened, with asymmetrical molar area internally. Maxillae elongate, symmetrical; cardo strip-like, projecting outward posteriorly; stipes bulged posteriorly and laterally; galea clearly separated from stipes, flattened apically and curved internally; lacinia stylet-like, apically curved posterointernally, basally extended deeply into head capsule; palpus four segmented, second and fourth articles especially long, inner surface of terminal article flattened; palpifer well developed. Hypopharynx elongate, apically exceeding base of labial palpus; anterior surface with keel along median line, placed between lacinial stylets. Labium elongate, medially concave forming groove housing elongated hypopharynx and lacinial stylets; paraglossa present; glossa absent; palpus four segmented (or three if the most basal tiny article represents palpifer).
|
||
</paragraph>
|
||
<paragraph id="8BA99858FFC39F0DFF68FF0E5D12FEB7" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="232">
|
||
<emphasis id="B962444AFFC39F0DFF68FF0E5977FF0F" box="[199,280,223,248]" italics="true" pageId="3" pageNumber="232">Thorax</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC39F0DFE85FF0E5903FF0F" box="[298,364,223,248]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="3" pageNumber="232">Fig. 1</figureCitation>
|
||
a,b). Prothorax small, detached from mesothorax. Meso- and metathorax fused, analogous in structures, well developed, bulged dorsally; scutum bilobed; scutellum pentagonal in shape; basalare well differentiated from episternum; anapleural suture not visible; posterior margin of epimeron protruding medially.
|
||
</paragraph>
|
||
<caption id="DF69C8D0FFC39F0DFF38F8755CF2F82E" httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="3" pageNumber="232" targetBox="[243,1343,375,1931]" targetPageId="3">
|
||
<paragraph id="8BA99858FFC39F0DFF38F8755CF2F82E" blockId="3.[151,1436,1956,2009]" pageId="3" pageNumber="232">
|
||
<emphasis id="B962444AFFC39F0DFF38F8755976F84E" bold="true" box="[151,281,1956,1978]" pageId="3" pageNumber="232">FIGURE 1.</emphasis>
|
||
Photographs of
|
||
<taxonomicName id="4C16E3DBFFC39F0DFE7BF8755A80F84D" box="[468,751,1956,1978]" class="Insecta" family="Archipsyllidae" genus="Mydiognathus" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="232" phylum="Arthropoda" rank="species" species="eviohlhoffae">
|
||
<emphasis id="B962444AFFC39F0DFE7BF8755A80F84D" box="[468,751,1956,1978]" italics="true" pageId="3" pageNumber="232">Mydiognathus eviohlhoffae</emphasis>
|
||
</taxonomicName>
|
||
. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69C8D0FFC49F0AFF38FB615B72FA89" httpUri="https://zenodo.org/record/1050649/files/figure.png" pageId="4" pageNumber="233" targetBox="[153,1434,211,1154]" targetPageId="4">
|
||
<paragraph id="8BA99858FFC49F0AFF38FB615B72FA89" blockId="4.[151,1437,1200,1406]" pageId="4" pageNumber="233">
|
||
<emphasis id="B962444AFFC49F0AFF38FB61597BFB32" bold="true" box="[151,276,1200,1222]" pageId="4" pageNumber="233">FIGURE 2.</emphasis>
|
||
Key head structures of
|
||
<taxonomicName id="4C16E3DBFFC49F0AFDBFFB615B48FB31" box="[528,807,1200,1222]" class="Insecta" family="Archipsyllidae" genus="Mydiognathus" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="eviohlhoffae">
|
||
<emphasis id="B962444AFFC49F0AFDBFFB615B48FB31" box="[528,807,1200,1222]" italics="true" pageId="4" pageNumber="233">Mydiognathus eviohlhoffae</emphasis>
|
||
</taxonomicName>
|
||
(a–c) and
|
||
<taxonomicName id="4C16E3DBFFC49F0AFC37FB615C23FB30" box="[920,1100,1200,1223]" class="Insecta" family="Prionoglarididae" genus="Neotrogla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="aurora">
|
||
<emphasis id="B962444AFFC49F0AFC37FB615C23FB30" box="[920,1100,1200,1223]" italics="true" pageId="4" pageNumber="233">Neotrogla aurora</emphasis>
|
||
</taxonomicName>
|
||
(d). (a) Head. Dotted gray lines indicate alignment of mouthpart structures against the labrum (Lr) or lacinia (Lc, only for galea: Ga) (Acl: anteclypeus; At: antenna; ATP: anterior tentorial pit; Ca: cardo; E: eye; ES: epistomal suture; FGR: frontogenal suture; Fr: frons; Ga: galea; Ge: gena; Lc: lacinia; LiP: labial palpus; Lr: labrum; Md: mandible; MxP: maxillary palpus; Oc: ocellus; Pcl: postclypeus; Pf: palpifer; Pg: paraglossa; St: stipes). (b) Head, magnified. White arrowheads indicate the internally expanded lacinial base. (c) Tip of the mouth, magnified. (d) Antennal flagellum of
|
||
<taxonomicName id="4C16E3DBFFC49F0AFCAFFA9B5BD8FA97" box="[768,951,1354,1376]" class="Insecta" family="Prionoglarididae" genus="Neotrogla" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="species" species="aurora">
|
||
<emphasis id="B962444AFFC49F0AFCAFFA9B5BD8FA97" box="[768,951,1354,1376]" italics="true" pageId="4" pageNumber="233">Neotrogla aurora</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C16E3DBFFC49F0AFC64FA9B5C5FFA97" box="[971,1072,1354,1376]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
: Trogiomorpha:
|
||
<taxonomicName id="4C16E3DBFFC49F0AFB4BFA9B5DFEFA97" box="[1252,1425,1354,1376]" class="Insecta" family="Prionoglarididae" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="family">Prionoglarididae</taxonomicName>
|
||
), showing the rupturing mechanism observed in extant
|
||
<taxonomicName id="4C16E3DBFFC49F0AFD19FAB95B76FA89" box="[694,793,1384,1406]" class="Insecta" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="233" phylum="Arthropoda" rank="order">Psocodea</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA99858FFC49F0AFF68FA7D5D33F9A3" blockId="4.[151,1437,1451,1980]" pageId="4" pageNumber="233">
|
||
<emphasis id="B962444AFFC49F0AFF68FA7D5965FA34" box="[199,266,1452,1475]" italics="true" pageId="4" pageNumber="233">Wings</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC49F0AFEB2FA7A590BFA33" box="[285,356,1451,1476]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="4" pageNumber="233">Figs 1</figureCitation>
|
||
a,b, 3a). Membrane and veins of both wings evenly covered by microtrichia; margins setose. Forewing transparent with broad blackish brown region anteroproximally; Sc short, ending on anterior margin; pterostigma darkly pigmented but not significantly thickened; nodus and nodulus (fore- and hindwings coupling mechanism) absent; venation as indicated for family (
|
||
<bibRefCitation id="EF87E5A9FFC49F0AFD43F9C65BD3F9C7" author="Vishniakova" box="[748,956,1559,1584]" pageId="4" pageNumber="233" refString="Vishniakova, V. N. (1976) Relict Archipsyllidae (Insecta: Psocoptera) in the Mesozoic fauna. Paleontology Journal, 2, 76 - 84." type="journal article" year="1976">Vishniakova 1976</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF87E5A9FFC49F0AFC68F9C95CFFF9D8" author="Huang" box="[967,1168,1559,1584]" pageId="4" pageNumber="233" refString="Huang, D. Y., Nel, A., Azar, D. & Nel, P. (2008) Phylogenetic relationships of the Mesozoic paraneopteran family Archipsyllidae (Insecta: Psocodea). Geobios, 41, 461 - 464. http: // dx. doi. org / 10.1016 / j. geobios. 2007.11.003" type="journal article" year="2008">
|
||
Huang
|
||
<emphasis id="B962444AFFC49F0AFBB7F9C95C27F9C7" box="[1048,1096,1559,1584]" italics="true" pageId="4" pageNumber="233">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
) and genus (see above); bases of Cu veins obscure. Hindwing slightly smaller and very similar to forewing, but areola postica absent.
|
||
</paragraph>
|
||
<paragraph id="8BA99858FFC49F0AFF68F9B15A08F8DB" blockId="4.[151,1437,1451,1980]" pageId="4" pageNumber="233">
|
||
<emphasis id="B962444AFFC49F0AFF68F9B15A6CF98F" box="[199,515,1631,1656]" italics="true" pageId="4" pageNumber="233">Forewing base articulations</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC49F0AFDBDF98E5A3AF98F" box="[530,597,1631,1656]" captionStart="FIGURE 3" captionStartId="5.[151,250,1842,1864]" captionTargetBox="[154,1424,195,1820]" captionTargetId="figure@5.[151,1436,193,1821]" captionTargetPageId="5" captionText="FIGURE 3. Illustrations of key wing structures in Mydiognathus eviohlhoffae (a – c) and other insect orders (d – f). (a) Forewing (composite drawing reconstructed from distal half of right forewing and basal half of left forewing). (b) Left forewing base articulation. (c) Right forewing base articulation. Forewing base articulation d – f: (d) Alloperla sp. (Plecoptera), (e) Longivalvus nubilus (Psocodea), (f) Petalolyma bicolor (Hemiptera)." httpUri="https://zenodo.org/record/1050651/files/figure.png" pageId="4" pageNumber="233">Fig. 3</figureCitation>
|
||
b,c). Tips of ANWP and MNWP invisible. PNWP narrow and long, tightly articulated with 3Ax. Tg well developed. HP and BSc fused. BSc and 2Ax separated by BR. First axillary sclerite hardly visible. Anterior region of 2Ax strongly swollen, divided into two sclerites; posterior region small and triangular, articulated with 3Ax. Proximal and distal arms of 3Ax short, anterior arm long. PMP deeply concave, distal margin strongly sclerotized, anteroproximal region broadly membranous, fused to BA posteriorly and distally. DMP small, located next to 2Ax.
|
||
</paragraph>
|
||
<paragraph id="8BA99858FFC49F0AFF68F8E95BF8F84B" blockId="4.[151,1437,1451,1980]" pageId="4" pageNumber="233">
|
||
<emphasis id="B962444AFFC49F0AFF68F8E95893F8B8" box="[199,252,1848,1871]" italics="true" pageId="4" pageNumber="233">Legs</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC49F0AFEBEF8E65939F8A7" box="[273,342,1847,1872]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="4" pageNumber="233">Fig. 1</figureCitation>
|
||
a,b,e). All legs analogous in structures; hind leg significantly longer than other legs but no specialization detected (e.g. for jumping). Coxae well developed, no specialization on internal surface of hind coxae. Tibiae with two well-developed tibial spurs distally. Tarsi four segmented; pretarsal claws symmetrical, without preapical tooth, with well-developed arolium between them.
|
||
</paragraph>
|
||
<caption id="DF69C8D0FFC59F0BFF38F8E35ABEF853" httpUri="https://zenodo.org/record/1050651/files/figure.png" pageId="5" pageNumber="234" targetBox="[154,1424,195,1820]" targetPageId="5">
|
||
<paragraph id="8BA99858FFC59F0BFF38F8E35ABEF853" blockId="5.[151,1436,1842,1956]" pageId="5" pageNumber="234">
|
||
<emphasis id="B962444AFFC59F0BFF38F8E3597DF8B0" bold="true" box="[151,274,1842,1864]" pageId="5" pageNumber="234">FIGURE 3.</emphasis>
|
||
Illustrations of key wing structures in
|
||
<taxonomicName id="4C16E3DBFFC59F0BFD36F8E35BC2F8BF" box="[665,941,1842,1864]" class="Insecta" family="Archipsyllidae" genus="Mydiognathus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="234" phylum="Arthropoda" rank="species" species="eviohlhoffae">
|
||
<emphasis id="B962444AFFC59F0BFD36F8E35BC2F8BF" box="[665,941,1842,1864]" italics="true" pageId="5" pageNumber="234">Mydiognathus eviohlhoffae</emphasis>
|
||
</taxonomicName>
|
||
(a–c) and other insect orders (d–f). (a) Forewing (composite drawing reconstructed from distal half of right forewing and basal half of left forewing). (b) Left forewing base articulation. (c) Right forewing base articulation. Forewing base articulation d–f: (d)
|
||
<taxonomicName id="4C16E3DBFFC59F0BFC41F8BE5C21F872" box="[1006,1102,1903,1925]" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B962444AFFC59F0BFC41F8BE5C21F872" box="[1006,1102,1903,1925]" italics="true" pageId="5" pageNumber="234">Alloperla</emphasis>
|
||
</taxonomicName>
|
||
sp. (
|
||
<taxonomicName id="4C16E3DBFFC59F0BFBD2F8BE5C82F872" box="[1149,1261,1903,1925]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="234" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
|
||
), (e)
|
||
<taxonomicName id="4C16E3DBFFC59F0BFA8FF8BE5934F853" authority="(Psocodea)" baseAuthorityName="Psocodea" class="Insecta" family="Psocidae" genus="Longivalvus" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="234" phylum="Arthropoda" rank="species" species="nubilus">
|
||
<emphasis id="B962444AFFC59F0BFA8FF8BE588DF853" italics="true" pageId="5" pageNumber="234">Longivalvus nubilus</emphasis>
|
||
(Psocodea)
|
||
</taxonomicName>
|
||
, (f)
|
||
<taxonomicName id="4C16E3DBFFC59F0BFE2BF85F5AA2F853" authority="(Hemiptera)" baseAuthorityName="Hemiptera" box="[388,717,1934,1956]" class="Insecta" family="Triozidae" genus="Petalolyma" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="234" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B962444AFFC59F0BFE2BF85F5A28F853" box="[388,583,1934,1956]" italics="true" pageId="5" pageNumber="234">Petalolyma bicolor</emphasis>
|
||
(Hemiptera)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF69C8D0FFC69F08FF38FBB05921FB61" httpUri="https://zenodo.org/record/1050653/files/figure.png" pageId="6" pageNumber="235" targetBox="[404,1174,197,1095]" targetPageId="6">
|
||
<paragraph id="8BA99858FFC69F08FF38FBB05921FB61" blockId="6.[151,1436,1121,1174]" pageId="6" pageNumber="235">
|
||
<emphasis id="B962444AFFC69F08FF38FBB0597CFB81" bold="true" box="[151,275,1121,1143]" pageId="6" pageNumber="235">FIGURE 4.</emphasis>
|
||
Male terminalia of
|
||
<taxonomicName id="4C16E3DBFFC69F08FE70FBB05A9BFB80" box="[479,756,1121,1143]" class="Insecta" family="Archipsyllidae" genus="Mydiognathus" kingdom="Animalia" order="Psocodea" pageId="6" pageNumber="235" phylum="Arthropoda" rank="species" species="eviohlhoffae">
|
||
<emphasis id="B962444AFFC69F08FE70FBB05A9BFB80" box="[479,756,1121,1143]" italics="true" pageId="6" pageNumber="235">Mydiognathus eviohlhoffae</emphasis>
|
||
</taxonomicName>
|
||
(St: sternite; T: tergite; Ad: aedeagus; Ep: epiproct; Pa: paraproct; Tr: trichobothria).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA99858FFC69F08FF68FB1259A9FA9B" blockId="6.[151,1437,1219,1639]" pageId="6" pageNumber="235">
|
||
<emphasis id="B962444AFFC69F08FF68FB12595EFB2B" box="[199,305,1219,1244]" italics="true" pageId="6" pageNumber="235">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="132D84DDFFC69F08FEEEFB1259E8FB2B" box="[321,391,1219,1244]" captionStart="FIGURE 1" captionStartId="3.[151,250,1956,1978]" captionTargetBox="[243,1343,375,1931]" captionTargetId="figure@3.[243,1344,375,1931]" captionTargetPageId="3" captionText="FIGURE 1. Photographs of Mydiognathus eviohlhoffae. (a) Habitus, right view. (b) Habitus, left view. (c) Head. (d) Connection between first (top) and second antennal flagellomeres. (e) Right hind tarsus, ventral view." httpUri="https://zenodo.org/record/1050647/files/figure.png" pageId="6" pageNumber="235">Figs 1</figureCitation>
|
||
a,b, 4). Weakly sclerotized, tergum and sternum well differentiated. First abdominal tergite clearly differentiated from second segment; sternite 1 not clearly differentiated. Terminal segments well sclerotized. Epiproct convex dorsally, pointed posteriorly. Paraproct rounded posteriorly, with swollen trichobothrial field. Ninth sternum strongly protruding posteriorly, genital chamber broadly open dorsally. Aedeagus pointed apically.
|
||
</paragraph>
|
||
<paragraph id="8BA99858FFC69F08FF68FAA75C8DFA43" blockId="6.[151,1437,1219,1639]" pageId="6" pageNumber="235">
|
||
<emphasis id="B962444AFFC69F08FF68FAA7594FFA78" box="[199,288,1398,1423]" italics="true" pageId="6" pageNumber="235">Lengths</emphasis>
|
||
(seen from right side, in mm). Body 2.12; antennal flagellomere f1 0.60, f2 0.56, f3 0.61; forewing 2.38, hindwing 2.20; hind femur 0.68, hind tibia 1.26, hind tarsus t1 0.38, t2 0.16, t3 0.05, t4 0.06.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CCBD3FFC69F08FF68FA6F5A45F9E8" pageId="6" pageNumber="235" type="materials_examined">
|
||
<paragraph id="8BA99858FFC69F08FF68FA6F5A45F9E8" blockId="6.[151,1437,1219,1639]" pageId="6" pageNumber="235">
|
||
<typeStatus id="54AD26FAFFC69F08FF68FA6F595BFA20" box="[199,308,1470,1495]" pageId="6" pageNumber="235" type="holotype">
|
||
<emphasis id="B962444AFFC69F08FF68FA6F595BFA20" bold="true" box="[199,308,1470,1495]" pageId="6" pageNumber="235">Holotype</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<collectingCountry id="F301D8C8FFC69F08FEEBFA6F59EEFA20" box="[324,385,1470,1495]" name="Malta" pageId="6" pageNumber="235">Male</collectingCountry>
|
||
. Embedded in Burmite, early Cenomanian (Upper Cretaceous, ca. 99 Mya:
|
||
<bibRefCitation id="EF87E5A9FFC69F08FB4CFA6F5DFFFA20" author="Shi" box="[1251,1424,1470,1495]" pageId="6" pageNumber="235" refString="Shi, G., Grimaldi, D. A., Harlow, G. E., Wang, J., Wang, J., Yang, M., Lei, W., Li, Q. & Li, X. (2012) Age constraint on Burmese amber based on U - Pb dating of zircons. Cretaceous Research, 37, 155 - 163. http: // dx. doi. org / 10.1016 / j. cretres. 2012.03.014" type="journal article" year="2012">
|
||
Shi
|
||
<emphasis id="B962444AFFC69F08FABEFA115D2CFA20" box="[1297,1347,1470,1495]" italics="true" pageId="6" pageNumber="235">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
), Hukawng Valley, northern
|
||
<collectingCountry id="F301D8C8FFC69F08FE66FA355A59FA0B" box="[457,566,1508,1532]" name="Myanmar" pageId="6" pageNumber="235">Myanmar</collectingCountry>
|
||
(to be deposited in State Museum of Natural History in Stuttgart,
|
||
<collectingCountry id="F301D8C8FFC69F08FA80FA325DF7FA0C" box="[1327,1432,1507,1531]" name="Germany" pageId="6" pageNumber="235">Germany</collectingCountry>
|
||
, under specimen
|
||
<collectionCode id="ED07009DFFC69F08FEE1F9D95904F9E8" box="[334,363,1544,1567]" pageId="6" pageNumber="235">ID</collectionCode>
|
||
<collectionCode id="ED07009DFFC69F08FEDCF9D659D0F9E8" LSID="urn:lsid:biocol.org:col:34840" box="[371,447,1543,1567]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="6" pageNumber="235">SMNS</collectionCode>
|
||
Bu-174).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30CCBD3FFC69F08FF68F9FA5A99F990" pageId="6" pageNumber="235" type="etymology">
|
||
<paragraph id="8BA99858FFC69F08FF68F9FA5A99F990" blockId="6.[151,1437,1219,1639]" pageId="6" pageNumber="235">
|
||
<emphasis id="B962444AFFC69F08FF68F9FA5929F9B3" bold="true" box="[199,326,1579,1604]" pageId="6" pageNumber="235">Etymology</emphasis>
|
||
. The specific epithet honours Evi Ohlhoff, wife of Rainer Ohlhoff (who made the specimen available for study) and codiscoverer of the specimen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |