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<document id="A68B2D5871B5480DA0B4E5858A56AACC" ID-DOI="10.5252/z2011n3a4" ID-ISSN="1638-9387" ID-Zenodo-Dep="4547017" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613606754513" checkinUser="felipe" docAuthor="Segniagbeto, Gabriel Hoinsoude, Trape, Jean François, David, Patrick, Ohler, Annemarie, Dubois, Alain &amp; Glitho, Isabelle Adolé" docDate="2011" docId="03B7FC4EC0346E78FF55FA8D6BEBE311" docLanguage="en" docName="Zoosystema.33.3.325-360.pdf" docOrigin="Zoosystema 33 (3)" docSource="http://dx.doi.org/10.5252/z2011n3a4" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Hapsidophrys lineatus Fischer 1856" docType="treatment" docVersion="3" lastPageNumber="334" masterDocId="FF8E8436C03D6E71FFEDFF816F6AE04F" masterDocTitle="The snake fauna of Togo: systematics, distribution and biogeography, with remarks on selected taxonomic problems" masterLastPageNumber="360" masterPageNumber="325" pageNumber="334" updateTime="1698873147243" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C66CED8F29C07859FBB040F9ADB87713">The snake fauna of Togo: systematics, distribution and biogeography, with remarks on selected taxonomic problems</mods:title>
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<mods:namePart id="A4FC22731ECF048A688FB4EA83986A42">Segniagbeto, Gabriel Hoinsoude</mods:namePart>
<mods:affiliation id="E68435F97700BDE6D1A87B66E3CAAF1F">Département de Zoologie et Biologie animale, Faculté des Sciences, Université de Lomé, BP 1515, Lomé (Togo) h _ segniagbeto @ yahoo. fr</mods:affiliation>
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<mods:namePart id="9CFE09E88061A38A8833264666E80E7D">Trape, Jean François</mods:namePart>
<mods:affiliation id="CAED7C13181EFB6D2E23895E3E75EFDC">Laboratoire de Paludologie et Zoologie médicale, UR 77, Institut de Recherche pour le Développement (IRD), Centre de Dakar, BP 1386, Dakar (Senegal) jean-françois. trape @ ird. fr</mods:affiliation>
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<mods:namePart id="73F76C94A492A64B18F74C3A77FB56AA">David, Patrick</mods:namePart>
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<mods:namePart id="09B83C422DA051A6DCEF9510C19ACC44">Ohler, Annemarie</mods:namePart>
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<mods:namePart id="323F22ABD6E139AC9C96ABDBBCA57EC4">Dubois, Alain</mods:namePart>
<mods:affiliation id="817BC0348A2C924CE5C68D92244F389A">UMR 7205 OSEB, Reptiles et Amphibiens, Département de Systématique et Évolution, Muséum national dHistoire naturelle, CP 30, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) pdavid @ mnhn. fr ohler @ mnhn. fr adubois @ mnhn. fr</mods:affiliation>
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<mods:namePart id="8B425C23D4DFF28415C1EB73A55CEA8B">Glitho, Isabelle Adolé</mods:namePart>
<mods:affiliation id="D08BFA2315676F3C51EB139CD275AFDD">Département de Zoologie et Biologie animale, Faculté des Sciences, Université de Lomé, BP 1515, Lomé (Togo) iglitho @ yahoo. fr</mods:affiliation>
<mods:nameIdentifier id="142AF24BAE7A1F0C7B2ABE8B3BBC2DDB" type="email">iglitho@yahoo.fr</mods:nameIdentifier>
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<mods:title id="AD0F8397DBAF4AAB2E6A3938F7C6CCAD">Zoosystema</mods:title>
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<mods:date id="737195DE5CF7AF7C942780530E116667">2011</mods:date>
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<treatment id="03B7FC4EC0346E78FF55FA8D6BEBE311" ID-DOI="http://doi.org/10.5281/zenodo.4547060" ID-Zenodo-Dep="4547060" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311" httpUri="http://treatment.plazi.org/id/03B7FC4EC0346E78FF55FA8D6BEBE311" lastPageNumber="334" pageId="9" pageNumber="334">
<subSubSection id="C3041ED3C0346E78FF55FA8D6D55E567" box="[184,575,1292,1320]" pageId="9" pageNumber="334" type="nomenclature">
<paragraph id="8BA14D58C0346E78FF55FA8D6D55E567" blockId="9.[184,575,1292,1320]" box="[184,575,1292,1320]" pageId="9" pageNumber="334">
<heading id="D0E9FA34C0346E78FF55FA8D6D55E567" box="[184,575,1292,1320]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="334" reason="2">
<taxonomicName id="4C1E36DBC0346E78FF55FA8D6D55E567" authority="Fischer, 1856" authorityName="Fischer" authorityYear="1856" box="[184,575,1292,1320]" class="Reptilia" family="Colubridae" genus="Hapsidophrys" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="lineatus">
<emphasis id="B96A914AC0346E78FF55FA8D6EC2E568" bold="true" box="[184,424,1292,1319]" italics="true" pageId="9" pageNumber="334">Hapsidophrys lineatus</emphasis>
Fischer, 1856
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3041ED3C0346E78FF9CFACD6E3DE651" pageId="9" pageNumber="334" type="materials_examined">
<paragraph id="8BA14D58C0346E78FF9CFACD6D78E530" blockId="9.[113,646,1356,1407]" pageId="9" pageNumber="334">
<materialsCitation id="3B764705C0346E78FF9CFACD6D64E530" collectionCode="MRAC" pageId="9" pageNumber="334" specimenCode="MRAC 73" specimenCount="2">
MATERIAL EXAMINED. —
<specimenCount id="9D1886D1C0346E78FE8AFACD6EB7E52B" box="[359,477,1356,1380]" pageId="9" pageNumber="334" type="generic">2 specimens</specimenCount>
(
<specimenCode id="DBB8E523C0346E78FE04FACD6D3BE52B" box="[489,593,1356,1380]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC 73</specimenCode>
14 R 32, Agou; GHS-W 1245, Yo [Agome Yo])
</materialsCitation>
.
</paragraph>
<paragraph id="8BA14D58C0346E78FF9CFA1D6E3DE651" blockId="9.[113,649,1436,1566]" pageId="9" pageNumber="334">
MORPHOLOGY. — TL 970 and
<quantity id="4CE6E0BDC0346E78FE57FA1D6D48E5FB" box="[442,546,1436,1460]" metricMagnitude="0" metricUnit="m" metricValue="1.095" pageId="9" pageNumber="334" unit="mm" value="1095.0">1095 mm</quantity>
; 15 MSR keeled and oblique; 159 and 165 Ven, keeled; 115 Sc, paired and smooth; anal single; 8-9 supralabials; 9 infralabials; 1 or 2 preoculars; 2 postoculars; temporal scale formula 2 + 2 + 3.
</paragraph>
</subSubSection>
<subSubSection id="C3041ED3C0346E78FF9CF9BD6CBDE15C" pageId="9" pageNumber="334" type="distribution">
<paragraph id="8BA14D58C0346E78FF9CF9BD6D78E6EC" blockId="9.[111,646,1596,1699]" pageId="9" pageNumber="334">
DISTRIBUTION. — This species is largely a forest-dweller as can be deduced from the collection localities. It is known only from ER IV. Previously it had been mentioned from
<collectingCountry id="F3090DC8C0346E78FF9DF90A6FC8E6EC" box="[112,162,1675,1699]" name="Togo" pageId="9" pageNumber="334">Togo</collectingCountry>
by
<bibRefCitation id="EF8F30A9C0346E78FF2BF90A6ECAE6EC" author="STERNFELD L." box="[198,416,1675,1699]" pageId="9" pageNumber="334" pagination="207 - 236" refId="ref29004" refString="STERNFELD L. 1908 b. - Die Schlangenfauna Togos. Mittheilungen Zoologischen Museum in Berlin 4: 207 - 236." type="journal article" year="1908">Sternfeld (1908b: 215</bibRefCitation>
,
<bibRefCitation id="EF8F30A9C0346E78FE47F90A6D6CE6EC" author="STERNFELD L." box="[426,518,1675,1699]" pageId="9" pageNumber="334" refId="ref29026" refString="STERNFELD L. 1909. - Die Schlangen Togos, in Die Fauna der deutschen Kolonien 1 (1): iv + 29 p., map." type="book" year="1909">1909: 15</bibRefCitation>
).
</paragraph>
<paragraph id="8BA14D58C0346E78FD2FFF566CBDE15C" blockId="9.[706,1180,215,275]" pageId="9" pageNumber="334">
<taxonomicName id="4C1E36DBC0346E78FD2FFF566BF6E0BC" authority="(Schlegel, 1837)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[706,1180,215,243]" class="Reptilia" family="Colubridae" genus="Hapsidophrys" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="334" phylum="Chordata" rank="species" species="smaragdinus">
<emphasis id="B96A914AC0346E78FD2FFF566C8FE0BD" bold="true" box="[706,997,215,242]" italics="true" pageId="9" pageNumber="334">Hapsidophrys smaragdinus</emphasis>
(Schlegel, 1837)
</taxonomicName>
(
<figureCitation id="132551DDC0346E78FC62FF796CA5E15C" box="[911,975,248,275]" captionStart="FIG" captionStartId="10.[677,688,626,643]" captionTargetBox="[677,1209,215,584]" captionTargetId="figure-614@10.[676,1209,214,584]" captionTargetPageId="10" captionText="FIG. 4. — Hapsidophrys smaragdinus (Schlegel, 1837), Agave. Photo by G. Segniagbeto." figureDoi="http://doi.org/10.5281/zenodo.4547027" httpUri="https://zenodo.org/record/4547027/files/figure.png" pageId="9" pageNumber="334">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3041ED3C0346E78FD48FEB46CE0E2EA" pageId="9" pageNumber="334" type="materials_examined">
<paragraph id="8BA14D58C0346E78FD48FEB46CECE248" blockId="9.[677,1212,309,519]" pageId="9" pageNumber="334">
<materialsCitation id="3B764705C0346E78FD48FEB46CE8E248" collectionCode="MRAC" country="Togo" county="Sodo" location="Kolokope" municipality="Agome Yo" pageId="9" pageNumber="334" specimenCode="MRAC 29585, MRAC 29626, MRAC 29715, MRAC 73017.0042, A7036.0016, A7036.0020, A7036.0004, 0005" specimenCount="17">
MATERIAL EXAMINED. —
<specimenCount id="9D1886D1C0346E78FC2FFEB46B3DE102" box="[962,1111,309,333]" pageId="9" pageNumber="334" type="generic">17 specimens</specimenCount>
(
<specimenCode id="DBB8E523C0346E78FB87FEB46D82E127" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC 29585</specimenCode>
,
<location id="8EC11B83C0346E78FD19FED16C32E127" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FD19FED16C32E127" box="[756,856,336,360]" country="Togo" county="Sodo" municipality="Agome Yo" name="Kolokope" pageId="9" pageNumber="334">Kolokope</location>
;
<specimenCode id="DBB8E523C0346E78FC89FED16C90E127" box="[868,1018,336,360]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC 29626</specimenCode>
,
<location id="8EC11B83C0346E78FBE8FED16B0EE127" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FBE8FED16B0EE127" box="[1029,1124,336,360]" country="Togo" county="Sodo" municipality="Agome Yo" name="Togoville" pageId="9" pageNumber="334">Togoville</location>
;
<specimenCode id="DBB8E523C0346E78FB9DFED16D82E1CD" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC 29715</specimenCode>
,
<location id="8EC11B83C0346E78FD18FEEB6C21E1CD" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FD18FEEB6C21E1CD" box="[757,843,362,386]" country="Togo" county="Sodo" municipality="Agome Yo" name="Kpalime" pageId="9" pageNumber="334">Kpalime</location>
;
<specimenCode id="DBB8E523C0346E78FCBBFEEB6B49E1CD" box="[854,1059,362,386]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC 73017.0042</specimenCode>
,
<location id="8EC11B83C0346E78FBC2FEEB6B0CE1CD" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FBC2FEEB6B0CE1CD" box="[1071,1126,362,386]" country="Togo" county="Sodo" municipality="Agome Yo" name="Agou" pageId="9" pageNumber="334">Agou</location>
;
<collectionCode id="ED0FD59DC0346E78FB9FFEEB6BD1E1CD" box="[1138,1211,362,386]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC</collectionCode>
<specimenCode id="DBB8E523C0346E78FD4BFE046C40E1D2" box="[678,810,389,413]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">A7036.0016</specimenCode>
,
<collectionCode id="ED0FD59DC0346E78FCDBFE046CEBE1D2" box="[822,897,389,413]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC</collectionCode>
<specimenCode id="DBB8E523C0346E78FC64FE046B67E1D2" box="[905,1037,389,413]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">A7036.0020</specimenCode>
, GHS-
<collectingCountry id="F3090DC8C0346E78FBB3FE046BFBE1D2" box="[1118,1169,389,413]" name="Togo" pageId="9" pageNumber="334">Togo</collectingCountry>
40,
<location id="8EC11B83C0346E78FD4BFE1E6C7BE1F8" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FD4BFE1E6C7BE1F8" box="[678,785,415,439]" country="Togo" county="Sodo" municipality="Agome Yo" name="Sodo Zion" pageId="9" pageNumber="334">Sodo Zion</location>
;
<collectionCode id="ED0FD59DC0346E78FCF7FE1E6C09E1F8" box="[794,867,415,439]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">MRAC</collectionCode>
<specimenCode id="DBB8E523C0346E78FC85FE1E6C82E1F8" box="[872,1000,415,439]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">A7036.0004</specimenCode>
-
<specimenCode id="DBB8E523C0346E78FC02FE1E6B4BE1F8" box="[1007,1057,415,439]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="9" pageNumber="334" type="Museum">0005</specimenCode>
, GHS-W 0016, GHS-W 0019, GHS-
<collectingCountry id="F3090DC8C0346E78FC6DFE3B6CD8E19D" box="[896,946,442,466]" name="Togo" pageId="9" pageNumber="334">Togo</collectingCountry>
63,
<location id="8EC11B83C0346E78FC32FE3B6B76E19D" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FC32FE3B6B76E19D" box="[991,1052,442,466]" country="Togo" county="Sodo" municipality="Agome Yo" name="Agave" pageId="9" pageNumber="334">Agave</location>
; GHS-W 0061, GHS-W 1238, GHS-W 1253-1254,
<location id="8EC11B83C0346E78FBF6FE556B5EE1A3" LSID="urn:lsid:plazi:treatment:03B7FC4EC0346E78FF55FA8D6BEBE311:8EC11B83C0346E78FBF6FE556B5EE1A3" box="[1051,1076,468,492]" country="Togo" county="Sodo" municipality="Agome Yo" name="Yo" pageId="9" pageNumber="334">Yo</location>
(
<collectingMunicipality id="6BC5D722C0346E78FBAEFE556BDAE1A3" box="[1091,1200,468,492]" pageId="9" pageNumber="334">Agome Yo</collectingMunicipality>
); GHS-
<collectingCountry id="F3090DC8C0346E78FD0AFE6E6C73E248" box="[743,793,495,519]" name="Togo" pageId="9" pageNumber="334">Togo</collectingCountry>
23,
<collectingCounty id="62C035D4C0346E78FCA8FE6E6C11E248" box="[837,891,495,519]" pageId="9" pageNumber="334">Sodo</collectingCounty>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA14D58C0346E78FD48FDA26CE0E2EA" blockId="9.[677,1211,547,677]" pageId="9" pageNumber="334">
MORPHOLOGY. — TL from
<quantity id="4CE6E0BDC0346E78FC58FDA26B37E274" box="[949,1117,547,571]" metricMagnitude="-1" metricUnit="m" metricValue="8.575" metricValueMax="11.5" metricValueMin="5.65" pageId="9" pageNumber="334" unit="mm" value="857.5" valueMax="1150.0" valueMin="565.0">565 to 1150 mm</quantity>
; 15 MSR keeled and oblique; 154-172 Ven, keeled; 139-160 Sc, paired and keeled; anal divided; loreal subrectangular, longer than high; 1 preocular; 2 postoculars; temporal scale formula 1 + 2 + 3.
</paragraph>
</subSubSection>
<subSubSection id="C3041ED3C0346E78FD48FD406BEBE311" pageId="9" pageNumber="334" type="distribution">
<paragraph id="8BA14D58C0346E78FD48FD406BEBE311" blockId="9.[675,1210,705,862]" pageId="9" pageNumber="334">
DISTRIBUTION. — This species is common in the forested area. It is present in the South of the country. Its northern limit is located in Mounts Aledjo. Previously, this species had been cited from
<collectingCountry id="F3090DC8C0346E78FC86FC906CF7E366" box="[875,925,785,809]" name="Togo" pageId="9" pageNumber="334">Togo</collectingCountry>
by
<bibRefCitation id="EF8F30A9C0346E78FC52FC906B0FE366" author="MATSCHIE P." box="[959,1125,785,809]" pageId="9" pageNumber="334" pagination="207 - 215" refId="ref28100" refString="MATSCHIE P. 1893. - Die Reptilien und Amphibien Togogebietes. Mittheilungen von Forschungsreisenden und Gelehrten aus den deutschen Schutzgebieten, mit Benutzung amtlicher Quellen 6: 207 - 215." type="journal article" year="1893">Matschie (1893)</bibRefCitation>
,
<bibRefCitation id="EF8F30A9C0346E78FB83FC906D81E30C" author="WERNER F." pageId="9" pageNumber="334" pagination="332 - 348" refId="ref29695" refString="WERNER F. 1902. - Uber westafrikanische Reptilien. Verhandlungen der kaiserlich-koniglichen Zoologischbotanischen Gesellschaft in Wien 52: 332 - 348." type="journal article" year="1902">Werner (1902)</bibRefCitation>
,
<bibRefCitation id="EF8F30A9C0346E78FD1AFCAA6CCFE30C" author="STERNFELD L." box="[759,933,811,835]" pageId="9" pageNumber="334" pagination="207 - 236" refId="ref29004" refString="STERNFELD L. 1908 b. - Die Schlangenfauna Togos. Mittheilungen Zoologischen Museum in Berlin 4: 207 - 236." type="journal article" year="1908">Sternfeld (1908b</bibRefCitation>
,
<bibRefCitation id="EF8F30A9C0346E78FC5CFCAA6B7AE30C" author="STERNFELD L." box="[945,1040,811,835]" pageId="9" pageNumber="334" refId="ref29026" refString="STERNFELD L. 1909. - Die Schlangen Togos, in Die Fauna der deutschen Kolonien 1 (1): iv + 29 p., map." type="book" year="1909">1909: 15</bibRefCitation>
),
<bibRefCitation id="EF8F30A9C0346E78FBC8FCAA6C26E311" author="HULSELMANS J. L. J. &amp; VERHEYEN W. N." pageId="9" pageNumber="334" pagination="200 - 204" refId="ref27452" refString="HULSELMANS J. L. J. &amp; VERHEYEN W. N. 1970. - Contribution a l'herpetologie de la Republique du Togo: Liste preliminaire des serpents recoltes par la deuxieme mission zoologique belge au Togo. Revue de Zoologie et de Botanique africaines 82 (1 - 2): 200 - 204." type="journal article" year="1970">Hulselmans &amp; Verheyen (1970)</bibRefCitation>
and Hulselmans
<emphasis id="B96A914AC0346E78FC10FCC76B47E312" box="[1021,1069,837,861]" italics="true" pageId="9" pageNumber="334">et al.</emphasis>
(1970).
</paragraph>
</subSubSection>
</treatment>
</document>